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Shrub encroachment in alpine and subalpine environments has raised concerns in recent decades, particularly due to climate change and shifts in land use. This study investigates the varying impacts of shrub invasion on different alpine and subalpine herbaceous communities and analyzes the spatial characteristics of shrub invasion. Three sites in the central Italian Apennines were examined by comparing vegetation maps from 1954 and 2016, alongside environmental variables such as slope, ruggedness, elevation, and soil fertility. The findings indicate that alpine shrub vegetation more successfully invades open herbaceous communities compared to closed grasslands with nutrient-rich soils, which are more resistant to shrub invasion. The invasion process predominantly involves the expansion of existing shrub patches, gradually overtaking grasslands. This selective pattern of shrub invasion highlights the significance of varying competition dynamics in changing high-elevation ecosystems globally. [ABSTRACT FROM AUTHOR]

Invasive alien species represent a major threat to global biodiversity and the sustenance of ecosystems. Globally, mountain ecosystems have shown a degree of resistance to invasive species due to their distinctive ecological features. However, in recent times, the construction of linear infrastructure, such as roads, might weaken this resistance, especially in the Mediterranean basin region. Roads, by acting as efficient corridors, facilitate the dispersal of alien species along elevation gradients in mountains. Here, we investigated how the ecological features and road-associated disturbance in native plant communities affected both the occurrence and cover of alien plant species in Central Apennines (Italy). We implemented the MIREN road survey in three mountain transects conducting vegetation sampling in plots located both adjacent to and distant from the roads at intervals of ~ 100 m in elevation. We then used community-weighted means of Ecological Indicator Values for Europe together with Disturbance Indicator Values applied to plant species of native communities as predictors of alien species occurrence and cover in a machine-learning classification and regression framework. Our analyses showed that alien species' occurrence was greater in proximity to the road where high soil disturbance occurred and in warm- and light-adapted native communities. On the other hand, alien species cover was more strongly related to moderate grazing pressure and the occurrence of nitrophilic plant communities. These findings provide a baseline for the current status of alien plant species in this Mediterranean mountain region, offering an ecological perspective to address the challenges associated with their management under global change. [ABSTRACT FROM AUTHOR]

Abstract Background Long-term farmland abandonment has increased fuel build-up in many Euro-Mediterranean mountainous regions. The high fuel hazard in these landscapes, combined with ongoing climate change, is increasing the frequency of extreme wildfires, thus altering contemporary fire regimes. Mitigating the loss of the landscape’s capacity to regulate large and intense fires is crucial to prevent future harmful effects of fires. As such, effective strategies to manage these fire-prone landscapes are needed. Yet, further understanding of their performance under global change scenarios is required. This study assessed the effects of fire-smart management strategies on future landscape dynamics, fire regulation capacity (FRC), and fire regime in a Mediterranean fire-prone mountainous landscape in Portugal (30,650 ha) undergoing long-term land abandonment and climate change scenarios. For that, we applied the LANDIS-II model under climate change scenarios (RCP 4.5 and 8.5) and long-term farmland abandonment (2020–2050) according to three fire-smart management strategies focused on fire prevention compared with a business-as-usual (BAU) strategy based on fire suppression. Results Future fire activity and land dynamics resulted in changes that fostered landscape heterogeneity and fragmentation and favoured fire-adapted forests and agroforestry systems while decreasing the dominance of shrublands and croplands. FRC decreased over time, particularly under RCP 8.5 and the BAU strategy. In turn, fire-smart strategies better prevented large and intense fires than the BAU strategy, but their effectiveness decreased under RCP 8.5. The loss of FRC resulted in increased burned area and fire frequency, which predicts a shift from contemporary fire regimes but more markedly under RCP 8.5 and in the BAU strategy. Conclusions Fire-smart strategies outperformed BAU in averting current fire regime intensification. Merging forest- and silvopasture-based management is the most promising approach in taming the effects of climate and farmland abandonment on future fire activity. Our study underlines that planning and management policies in fire-prone Mediterranean mountain landscapes must integrate fire-smart strategies to decrease landscape fuel hazard and buffer the impact of global change on future fire regimes.

Background: Long-term farmland abandonment has increased fuel build-up in many Euro-Mediterranean mountainous regions. The high fuel hazard in these landscapes, combined with ongoing climate change, is increasing the frequency of extreme wildfires, thus altering contemporary fire regimes. Mitigating the loss of the landscape's capacity to regulate large and intense fires is crucial to prevent future harmful effects of fires. As such, effective strategies to manage these fire-prone landscapes are needed. Yet, further understanding of their performance under global change scenarios is required. This study assessed the effects of fire-smart management strategies on future landscape dynamics, fire regulation capacity (FRC), and fire regime in a Mediterranean fire-prone mountainous landscape in Portugal (30,650 ha) undergoing long-term land abandonment and climate change scenarios. For that, we applied the LANDIS-II model under climate change scenarios (RCP 4.5 and 8.5) and long-term farmland abandonment (2020–2050) according to three fire-smart management strategies focused on fire prevention compared with a business-as-usual (BAU) strategy based on fire suppression. Results: Future fire activity and land dynamics resulted in changes that fostered landscape heterogeneity and fragmentation and favoured fire-adapted forests and agroforestry systems while decreasing the dominance of shrublands and croplands. FRC decreased over time, particularly under RCP 8.5 and the BAU strategy. In turn, fire-smart strategies better prevented large and intense fires than the BAU strategy, but their effectiveness decreased under RCP 8.5. The loss of FRC resulted in increased burned area and fire frequency, which predicts a shift from contemporary fire regimes but more markedly under RCP 8.5 and in the BAU strategy. Conclusions: Fire-smart strategies outperformed BAU in averting current fire regime intensification. Merging forest- and silvopasture-based management is the most promising approach in taming the effects of climate and farmland abandonment on future fire activity. Our study underlines that planning and management policies in fire-prone Mediterranean mountain landscapes must integrate fire-smart strategies to decrease landscape fuel hazard and buffer the impact of global change on future fire regimes. [ABSTRACT FROM AUTHOR]

Context: The Mediterranean mountains have witnessed the revegetation of former pasture and cultivated fields since the mid-20th century. The process started when the mountains joined a dynamic and competitive market, which led to depopulation, abandoning cultivated fields, and a reduction in extensive livestock farming. Objectives: We investigate changes in land use and land cover (LULC) and landscape structure in a valley of the Central Pyrenees in 1956, 1981 and 2017, and we discuss the implications for land management within a multidisciplinary context. Methods: Aerial photographs are used to map the LULC in 1956, 1981 and 2017, followed by landscape ecology metrics and indices to analyse the structure of the landscape. Results: The results show: (i) a reduction in cultivated area (-42%) and pastures (-63%) in the montane belt between 1956 and 2017, while forests doubled in area; (ii) changes in LULC are more significant in the montane belt than in subalpine and alpine areas; (iii) based on the Kappa index, it was found that the 29% of the area had undergone changes in LULC between 1956 and 2017, with the changes being greater in the first period (1956–1981) than in the second (1981–2017); (iv) the structure and heterogeneity of the landscape shows an increase between 1956 and 1981, and a change in trend from 1981 to 2017, so that the landscape is more homogeneous in 2017 than in 1956. Conclusion: The discussion suggests the need to maintain and recover mosaic landscapes, since their structure is akin to traditional cultural ones, to increase biodiversity and sustainability, and provide ecosystem services to the population. [ABSTRACT FROM AUTHOR]

Monitoring snowmelt dynamics in mountains is crucial to understand water releases downstream. Sentinel-1 (S-1) synthetic-aperture radar (SAR) has become one of the most widely used techniques to achieve this aim due to its high frequency of acquisitions and all-weather capability. This work aims to understand the possibilities of S-1 SAR imagery to capture snowmelt dynamics and related changes in streamflow response in semi-arid mountains. The results proved that S-1 SAR imagery was able not only to capture the final spring melting but also all melting cycles that commonly appear throughout the year in these types of environments. The general change detection approach to identify wet snow was adapted for these regions using as reference the average S-1 SAR image from the previous summer, and a threshold of −3.00 dB, which has been assessed using Landsat images as reference dataset obtaining a general accuracy of 0.79. In addition, four different types of melting-runoff onsets depending on physical snow condition were identified. When translating that at the catchment scale, distributed melting-runoff onset maps were defined to better understand the spatiotemporal evolution of melting dynamics. Finally, a linear connection between melting dynamics and streamflow was found for long-lasting melting cycles, with a determination coefficient (R2) ranging from 0.62 to 0.83 and an average delay between the melting onset and streamflow peak of about 21 days. [ABSTRACT FROM AUTHOR]

The average annual temperature, precipitation, and evaporation patterns are some of the characteristics which indicate to a globe where climate is rapidly changing. It remains uncertain regarding how this rapid climate change will impact freshwater biota; however the consequences could be absolutely catastrophic. It is urgently needed to better adapt and maintain the fragile system that comprises the Mediterranean ecosystems. Water shortages and changes in the land use can impact water quality and the watershed's ecosystem. Changes in temperature and precipitation trends can increase the intensity of land use, the increase of water depletion, and the degradation of water quality. The effects of climate change on eco hydrology change runoff, evaporation, surface storage, and soil moisture, that all have immediate effects on the biota and biodiversity of region. It is supported by studies on eco hydrological impacts on local watersheds. The spatial pattern and surface water dynamics of watershed systems are utilized as a perspective using which to examine the hydrologic consequences. The impacts of climate change on rainfall, evaporation, soil humidity, lakes level, and quality of water, habitats, species migration, biodiversity, and economies have been investigated. The potential climatic impacts on watershed ecosystems are determined by a spatiotemporal assessment of variations in temperature and precipitation projections from a multimodal ensemble. The susceptibility of a watershed changed depending over its geographic region, terrain, and human influence. As such a manner of dealing with climate change, improving watershed systems could be beneficial. Various techniques have been proposed which can protect watersheds against by the consequences of climate and land use change. Strengthening ecosystem vulnerability requires knowledge of the Eco hydrological processes. This review anticipated effects of climate change on the Eco hydrology of watershed systems in the Mediterranean and implies strategies for adaptation to create the mechanisms more responsive. [ABSTRACT FROM AUTHOR]

Protected areas are no longer focused solely on conservation and protection needs but play a central role in promoting sustainable development in local socio-territorial systems. The Tepilora Natural Regional Park (TNRP) in Sardinia offers an example of this phenomenon. This paper highlights the significance of the TNRP in balancing conservation needs with the sustainable development of local populations - an effort that resulted in the establishment of the Tepilora, Rio Posada and Montalbo Biosphere Reserve. [ABSTRACT FROM AUTHOR]

[Display omitted] • Mediterranean seasonal range pattern for chamois: summer is the stress period? • Population density and fecundity rate: low for the Balkan chamois on Mt. Olympus. • Chamois' escape terrain is defined as steep (>45⁰) and extended (≥2,500 m2) slopes. • Road avoidance and preference for proximity to hiking trails on Mt. Olympus. • Balkan chamois action plan priorities: roadless areas and water availability. The Balkan chamois (Rupicapra rupicapra balcanica) has a bad conservation status in Greece, and a national action plan will be implemented by the Ministry of Environment. We explored the seasonal distribution pattern and ecological preferences of the species on Mt. Olympus by collecting 1,182 observations of chamois occurrences across four seasonal surveys (2022–2023), and we estimated its population size (2022). The annual range of the species reached 103 km2 (Kernel Density Estimator). We recorded the smallest seasonal range during autumn (rutting season), then in summer, and the largest in winter. The species followed the Mediterranean seasonal range use pattern, implying that summer is the stress period due to drought. The population size was c. 430 individuals, showing an increasing trend, but the fecundity rate was low in 2022. The Ecological Niche Factor Analysis (ENFA) showed that chamois preferred areas near hiking trails throughout the year, likely being habituated with visitors, while avoiding motorized roads, as adopted behaviors of poaching risk minimization. It also favored rocky areas and proximity to escape terrains during spring and winter. This work provides new methodological insights for habitat mapping and escape terrain definition (slope steepness and extent) and supports the suggested measures of the national action plan for implementing a roadless and road control policy, securing water availability in arid mountains, implementing a management plan for tourists, abating poaching and enlarging the hunting banning zone. [ABSTRACT FROM AUTHOR]

Mediterranean mountains have been and continue to be used by human populations along an interweaving of numerous uses: agro-sylvo-pastoralism, trade, industry and mining have all gone hand in hand for several millennia. Mines are however a so important source of wealth that, by putting in contact external powers and mountain locals, it creates an imbalance of powers inducing structural violence and tensions. The 1830–1962 colonial era did change the magnitude of these imbalances and this affect all Mediterranean mountainous ranges. The French expansion did affect as a result the Moroccan Atlas, the Tunisia Coastal Mounts but also the French Pyrenees. The article explores the available archives regarding the history of three mines in each of these emblematic mountains with a shared mining and agro-sylvo-pastoral past and where mining were actually well-known: Sem-Rancié and Puymorens in the French Pyrenees, Mibladen and Zeïda in the Moroccan Middle Atlas and Jebel Ressass in Tunisia. These reconstructions show that the initial social and political situations, as diverse as they are, are of little importance in the trajectory of these mines: all of them see a rapid appropriation by economic powers that are more and more powerful and more and more distant as far as Paris, the common capital in colonial times, despite several revolts and tensions. The initial expansion then gave way to a structural crisis due to the competition with other mining sites until abandonment. The following powers, post-colonial in Morocco or Tunisia or decentralized in France, did not endorse any responsibility of this the post-mining environmental, social and economic legacy. For each of these sites, the mine could be seen as an indicator of the power balance evolution among activities and actors, a canary in the mine on which we propose a methodology for further investigations.

Question: Vegetation around the alpine–treeline ecotone faces changes in both climate and land use (i.e. grazing abandonment). Broad‐scale shrub encroachment is considered an effect of these changes, but it remains unclear how this process is mediated by local‐scale environmental heterogeneity. Our goal is to determine which local‐scale environmental factors shape the spatial distribution and temporal trends of alpine dwarf shrub vegetation dominated by Juniperus communis in Mediterranean mountains. Location: Three sites in the Central Apennine Mountain Range (Italy): Mt. Terminillo, Mt. Duchessa and Mt. Ernici. Methods: Combining a set of environmental variables obtained at a fine scale (25‐m resolution) from multi‐year remote sensing imagery and field‐collected vegetation data, we modelled the occurrence and cover of Juniperus communis‐dominated dwarf shrubland, as well as its change over time, as a function of local climate, topography and land use, using linear and generalized mixed effects models. Results: Over a period of almost 60 years (1954–2012) shrubland occurrence increased by 12% and shrubland cover by 10% in our study sites. Its current distribution and change over time appear to be strongly shaped by the joint influences of fine‐scale topography, above‐ground biomass, land use and microclimate. Shrublands have been favoured locally in areas with harsher alpine environmental constraints and stronger resource limitations. Conclusions: Our study shows that in the Mediterranean high‐mountain environment J. communis dwarf shrubland acts as a stress‐tolerant pioneer vegetation, occurring in areas that are otherwise sparsely vegetated. Contrary to our expectations, at fine scales, warmer temperatures and the regional decline in grazing did not favour shrub encroachment. Despite increasing overall, J. communis shrubs have little competitive ability to successfully encroach grasslands, and remain restricted to less‐productive areas. Our results confirm that fine‐scale environmental heterogeneity can strongly influence shrub distribution and dynamics, thereby modulating future responses in evolving alpine ecosystems. [ABSTRACT FROM AUTHOR]

The meridional mountains of Europe are located in Andalusia, in the south of the Iberian Peninsula. In this region, orographic and climatic factors are the main circumstances that condition the landscape and cultural peculiarities. Nowadays, its ecosystems show a high vulnerability to climate change, its traditional ways of life are in crisis and its cultural heritage is endangered by abandonment and depopulation. This paper analyzes some of the main documents that include proposals of environmental education at European, national and regional levels from the point of view of its adaptation the territory of the Andalusian Mediterranean mountains. Some educational actions are also evaluated in order to detect the possible existence of dysfunctionalities or deficiencies among the different educational policies and projects. Finally, a series of actions are proposed to optimize environmental education programs. [ABSTRACT FROM AUTHOR]

Forests cover 30% of the Earth's landmass, host 80% of the biodiversity on land, and represent one of the main sinks of carbon. Studying forest ecosystems and dynamics is more crucial than ever now that the climate is changing. On the other hand, forest structural attributes and microhabitats data acquisition is challenging, and require huge efforts.Here we provide a georeferenced dataset of living trees, deadwood, and microhabitats referring to 199 plots (13 m radius), collected between 2012 and 2018, and located over six Apennine mountainous forest types across Italy. The dataset we provide promotes collaboration among researchers and improves the possibilities to analyze the evolution of forest ecosystems.

Climate change is predicted to dramatically affect mountain biodiversity and especially mountain pollination systems due to the mutual dependence between plants and pollinators. In this work, we investigate climate change effects on pollinator distribution and diversity along the altitudinal gradient of Mt. Olympus, a biodiversity hotspot.We used a species distribution modelling framework and predicted species richness hotspots, potential distribution and altitude change for 114 pollinator species, comprising bees, butterflies, and hoverflies along the altitudinal gradient (327–2596 m a.s.l.).We projected potential loss of suitable habitat and upward shift for most pollinator groups, with the exception of bumblebees and hoverflies which were predicted to descend. Pollinator extinctions were not forecasted; instead, we observed a pronounced species-specific response to climate change. Species richness hotspots will be relocated to higher altitudes and to the north-eastern mountain side.Projections for substantial but not detrimental climate change effects on pollinator fauna may be due to species differential resilience to climate change along with the existence of microrefugia on Mt. Olympus. Divergent response to global warming by bumblebees and hoverflies is probably due to species distribution modelling limitations, resulting in exclusion of the rarest species. We conclude that the predicted climate change impact stresses for the need of urgent conservation measures, including the expansion of the protection status over the whole mountain.

Mountain areas are sensitive to changes in precipitation and temperature, which significantly impact traditional pastoralist communities, their economy, and their lifestyle. Alarming climate change scenarios justify the investigation of the ecological and socioeconomic vulnerabilities that characterize Portugal's mountain regions. This work explores how the traditional production systems of small ruminants—sheep and goats—could adapt in the Montesinho mountain range as it changes over the next 2 decades. Land use–land cover maps from 1995 and 2018 show how the pastoral landscape has changed and indicate trends for a future scenario. Documented landscape grazing patterns are used to determine sheep and goat landscape preferences under different climatic conditions. Finally, we identify the near-future constraints on traditional sheep and goat systems, contrasting landscape changes with sheep and goat preferences. Over coming decades, the balance between rangelands and cultivated lands will persist in the Montesinho mountain landscape, despite some trade-offs between both. Woodlands could emerge from scrublands colonizing rangelands, and permanent crops could significantly replace arable lands in agricultural areas. Therefore, it is likely that the agricultural areas preferred for sheep, and rangelands preferred for goats, may not be affected by the forecast landscape changes, but rather be favored by the expansion of permanent crops. However, pasture areas must expand, as they are key to pastoral landscape function in a warming climate scenario. Landscape decision makers and managers should implement a landscape-monitoring system to inform policies and strategies aimed at protecting and safeguarding mountain pastoralism and its vital ecosystem services.

Mountain areas are sensitive to changes in precipitation and temperature, which significantly impact traditional pastoralist communities, their economy, and their lifestyle. Alarming climate change scenarios justify the investigation of the ecological and socioeconomic vulnerabilities that characterize Portugal's mountain regions. This work explores how the traditional production systems of small ruminants—sheep and goats—could adapt in the Montesinho mountain range as it changes over the next 2 decades. Land use–land cover maps from 1995 and 2018 show how the pastoral landscape has changed and indicate trends for a future scenario. Documented landscape grazing patterns are used to determine sheep and goat landscape preferences under different climatic conditions. Finally, we identify the near-future constraints on traditional sheep and goat systems, contrasting landscape changes with sheep and goat preferences. Over coming decades, the balance between rangelands and cultivated lands will persist in the Montesinho mountain landscape, despite some trade-offs between both. Woodlands could emerge from scrublands colonizing rangelands, and permanent crops could significantly replace arable lands in agricultural areas. Therefore, it is likely that the agricultural areas preferred for sheep, and rangelands preferred for goats, may not be affected by the forecast landscape changes, but rather be favored by the expansion of permanent crops. However, pasture areas must expand, as they are key to pastoral landscape function in a warming climate scenario. Landscape decision makers and managers should implement a landscape-monitoring system to inform policies and strategies aimed at protecting and safeguarding mountain pastoralism and its vital ecosystem services. [ABSTRACT FROM AUTHOR]

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Climate change impacts are of a particular concern in small mountain ranges, where cold-adapted plant species have their optimum zone in the upper bioclimatic belts. This is commonly the case in Mediterranean mountains, which often harbour high numbers of endemic species, enhancing the risk of biodiversity losses. This study deals with shifts in vascular plant diversity in the upper zones of the Sierra Nevada, Spain, in relation with climatic parameters during the past two decades. We used vegetation data from permanent plots of three surveys of two GLORIA study regions, spanning a period of 18 years (2001–2019); ERA5 temperature and precipitation data; and snow cover durations, derived from on-site soil temperature data. Relationships between diversity patterns and climate factors were analysed using GLMMs. Species richness showed a decline between 2001 and 2008, and increased thereafter. Species cover increased slightly but significantly, although not for endemic species. While endemics underwent cover losses proportional to non-endemics, more widespread shrub species increased. Precipitation tended to increase during the last decade, after a downward trend since 1960. Precipitation was positively related to species richness, colonisation events, and cover, and negatively to disappearance events. Longer snow cover duration and rising temperatures were also related to increasing species numbers, but not to cover changes. The rapid biotic responses of Mediterranean alpine plants indicate a tight synchronisation with climate fluctuations, especially with water availability. Thus, it rather confirms concerns about biodiversity losses, if projections of increasing temperature in combination with decreasing precipitation hold true. [ABSTRACT FROM AUTHOR]

Forest biomass is a renewable energy source, more climate-friendly than fossil fuels and widely available in Europe. The wood energy chain has been suggested as a means to re-activate forest management and improve the value of forest stands in marginalized rural areas. However, wall-to-wall estimates of forest biomass, needed to design the location and size of power and heat biomass plants in any given territory, are notoriously difficult to obtain. This paper tests an algorithm to predict forest biomass using publicly available Landsat satellite imagery in the Liguria region, northern Italy. We used regional forest inventory data to train and validate an artificial neural network (ANN) classifier that uses remotely-sensed information such as three principal components of Landsat-5 TM spectral bands, the Enhanced Vegetation Index (EVI), and topography, to retrieve aboveground live tree volume. Percent root mean square error was -9% and -23% for conifers and broadleaves respectively in the calibration dataset, and -27% and -24% in the validation dataset. The reconstructed volume map was updated to present day using current volume increment rates reported by the Italian National Forest Inventory. A wall-to-wall map of forest biomass from harvest residues was finally produced based on species-specific wood density, biomass expansion factors, volume logged for timber assortments, forest accessibility, and topography. Predicted aboveground forest volume ranged from 81 to 391 m3 ha-1. In forests available for wood supply (70% of the total), planned volume removals averaged 25.4 m3 ha-1, or 18.7% of the average standing stock across. Biomass available for bioenergy supply was 1.295.921 million Mg dry matter or 8.95 Mg ha-1. This analysis workflow can be replicated in all mountain regions with a predominant broadleaved coppice component.

This paper aims to characterize small ruminant farming systems in the Moroccan mountain regions and to assess undergoing socio-economic and ecological changes. Livestock farming systems in these regions are extensive with large flock sizes, especially sheep and goats raised mainly for meat production. Sheep producers utilized the inherent genetic diversity of their stock and developed adaptations to cope with seasonal availability of the mountain pastures and the need to diversify their own activities. Nowadays, farming systems in mountainous regions are undergoing socio-economic and ecological transformations that impact directly or indirectly on natural resources. Reduction and/or disappearance of herd transhumance, where flocks were moved seasonally from one climatic zone/vegetation type to another, was an important feature of the Middle Atlas sixty years ago Along with this reduced contribution from natural rangelands, other external resources and agricultural by-products (straw, stubble) are offered which make the system heavily dependent on agriculture. Intensification of feeding systems is associated with a choice for more productive but less hardy breeds from lowland areas. As a consequence of the reduction in mobility and the sedentarization of herds, there is an increase in the grazing pressure causing new and significant degradation of rangeland resources. Other profound institutional modifications concern the management and the utilization of the mountainous pasture. Indeed, the power of the group and the institutions for managing the resources is being replaced by the individual initiatives.

Climatic changes associated with Pleistocene glacial cycles had profound effects on the genetic diversification and distribution of many taxa, especially from environmentally sensitive habitats such as mountain regions. The DNA sequence data (mtCOI and 28S RNA genes) derived from the Merodon rufus group (Diptera: Syrphidae) in central, eastern, and southern Europe was analyzed. Environmental niche modeling (ENM) and coalescent simulations also were performed to describe diversification events previously recognized via morphological analysis and to elucidate its evolutionary mechanism. Results revealed that M. rufus group is composed of four well-supported lineages, three of which belong to new species from Mediterranean mountains, namely M. kozufensis sp. nova, M. olympius sp. nova, and M. orjensis sp. nova. According to the divergence time estimates, lineage diversification most probably occurred during the Pleistocene by the colonization model rather than the fragmentation of a widely distributed common ancestor's range. ENM results showed potential overlap of climatic refugia on the Balkan Peninsula of the M. rufus group and the beech forests, suggesting a possible association with this habitat type. Merodon rufus Meigen, 1838 probably colonized further from these refugia and spread across other parts of Europe during the Holocene. [ABSTRACT FROM AUTHOR]

Climate change is increasingly favouring the occurrence of extreme weather events, affecting species distribution and tree growth. It has been proposed that mixed forests are more resistant to drought spells and heat waves than corresponding monocultures. We sampled tree cores in nine plots along a latitudinal gradient of the Italian Peninsula (Trentino, Molise and Calabria). The objective of the study was to identify radial growth responses in mixed-species and pure plots of European beech and silver fir to climate dynamics and extreme events. Pointer years, as well as correlation and response functions, were used to analyse the climate–growth relationships in mixed-species and pure plots. Climate–growth relationships showed differences between mixed-species and pure plots only at the regional level. In particular, Trentino differed from Molise and Calabria, when considering maximum temperatures. In Trentino, winter (for silver fir) and summer (for both species) temperatures had a less negative impact on the radial growth of trees, in comparison with Molise and Calabria. Nevertheless, correlations between the radial growth and drought indices, and the principal component analysis, showed that plots were relatively unresponsive to summer drought. These results can be important for the implementation of operational techniques that increase the adaptation of silver fir and European beech to climate change, as well as for assisted migration. [ABSTRACT FROM AUTHOR]

Many continental species distributed in the Eurasian steppe occur as relict populations in the mountains of Western Europe. Their biogeographical responses to Quaternary climate changes have been poorly studied; however, they could have responded as cold-adapted species. We investigated the biogeographic history of a steppe beetle, Hycleus polymorphus , using mitochondrial and nuclear DNA sequences (COI , CAD , ITS2), and species distribution modelling (SDM) under present and past bioclimatic envelopes. We first performed a phylogenetic assessment to define species boundaries within the H. polymorphus species group. Specimens previously treated as Hycleus humerosus on morphological grounds are assigned to H. polymorphus , and those identified as Hycleus zebraeus assigned to Hycleus atratus. ITS2 data analyses revealed a strong phylogeographical structure of H. polymorphus populations, with four haplogroups corresponding to the (i) Italian Alps, (ii) French Alps and Pyrenees, (iii) South Balkan and Pontic mountains, and (iv) North Dinaric Alps. Based on these analyses and the SDM, we propose that during a glacial period, following the spread of steppic habitat, H. polymorphus underwent a range expansion from Asia to South-West Europe. Within the Mediterranean area, during the last interglacial the climatic suitability for the species was limited to mountains that acted as refugia and prompted allopatric divergence into four main lineages. [ABSTRACT FROM AUTHOR]

Genetic diversity defines the evolutionary potential of a species, yet mounting evidence suggests that epigenetic diversity could also contribute to adaptation. Elucidating the complex interplay between genetic and epigenetic variation in wild populations remains a challenge for evolutionary biologists, and the intriguing possibility that epigenetic diversity could compensate for the loss of genetic diversity is one aspect that remains basically unexplored in wild plants. This hypothesis is addressed in this paper by comparing the extent and patterns of genetic and epigenetic diversity of phylogenetically closely related but ecologically disparate species. Seven pairs of congeneric species from Cazorla mountains in south-eastern Spain were studied, each pair consisting of one endemic, restricted-range species associated to stressful environments, and one widespread species occupying more favourable habitats. The prediction was tested that endemic species should have lower genetic diversity due to population fragmentation, and higher epigenetic diversity induced by environmental stress, than their widespread congeners. Genetic (DNA sequence variants) and epigenetic (DNA cytosine methylation variants) diversities and their possible co-variation were assessed in three populations of each focal species using amplified fragment length polymorphism (AFLP) and methylation-sensitive AFLP (MSAP). All species and populations exhibited moderate to high levels of genetic polymorphism irrespective of their ecological characteristics. Epigenetic diversity was greater than genetic diversity in all cases. Only in endemic species were the two variables positively related, but the difference between epigenetic and genetic diversity was greater at populations with low genetic polymorphism. Results revealed that the relationship between genetic and epigenetic diversity can be more complex than envisaged by the simple hypothesis addressed in this study, and highlight the need of additional research on the actual role of epigenetic variation as a source of phenotypic diversity before a realistic understanding of the evolutionary relevance of epigenetic phenomena in plant adaptation can be achieved. [ABSTRACT FROM AUTHOR]

Plant structural and biochemical traits are frequently used to characterise the life history of plants. Although some common patterns of trait covariation have been identified, recent studies suggest these patterns of covariation may differ with growing location and/or plant functional type (PFT). Mediterranean forest tree/shrub species are often divided into three PFTs based on their leaf habit and form, being classified as either needleleaf evergreen (Ne), broadleaf evergreen (Be), or broadleaf deciduous (Bd). Working across 61 mountainous Mediterranean forest sites of contrasting climate and soil type, we sampled and analysed 626 individuals in order to evaluate differences in key foliage trait covariation as modulated by growing conditions both within and between the Ne , Be , and Bd functional types. We found significant differences between PFTs for most traits. When considered across PFTs and by ignoring intraspecific variation, three independent functional dimensions supporting the Leaf-Height-Seed framework were identified. Some traits illustrated a common scaling relationship across and within PFTs, but others scaled differently when considered across PFTs or even within PFTs. For most traits much of the observed variation was attributable to PFT identity and not to growing location, although for some traits there was a strong environmental component and considerable intraspecific and residual variation. Nevertheless, environmental conditions as related to water availability during the dry season and to a smaller extend to soil nutrient status and soil texture, clearly influenced trait values. When compared across species, about half of the trait-environment relationships were species-specific. Our study highlights the importance of the ecological scale within which trait covariation is considered and suggests that at regional to local scales, common trait-by-trait scaling relationships should be treated with caution. PFT definitions by themselves can potentially be an important predictor variable when inferring one trait from another. These findings have important implications for local scale dynamic vegetation models. [ABSTRACT FROM AUTHOR]

Los bancales configuran uno de los paisajes agrarios más característico de las regiones de montaña mediterránea. Sin embargo, cambios demográficos y socioeconómicos, ocurridos en la segunda mitad del siglo pasado, provocaron el abandono de las terrazas, con la consecuente desaparición de las labores de mantenimiento que requerían estas estructuras agrarias. Esto favoreció la activación de diversos procesos de erosión entre los que destacan el colapso de los muros de piedra y los pequeños desprendimientos que se activan en los saltos de las terrazas. En este trabajo se analizan estos movimientos en masa cuantificando sus dimensiones y frecuencia, y definiendo las variables que intervienen en su desencadenamiento en los altos valles del Leza, Jubera y Cidacos (Camero Viejo, Sistema Ibérico, España), donde los bancales llegaron a ocupar 13.274 hectáreas, es decir el 63% del espacio agrícola. Un total de 240 desprendimientos fueron contabilizados en las 53 terrazas analizadas, lo que supone un número medio de 4,5 desprendimientos por parcela y 33,1 m3 de material movilizado por 100 m de muro. En el estudio se comprueba que las causas más decisivas para la iniciación de desprendimientos están vinculadas a la capacidad de infiltración de los suelos y a la forma en que el agua circula por la ladera.

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The present paper highlights the importance of hedgerows and enclosures in the mountains of Central Spain. Now, these landscapes have suffered profound variations in terms of agroforestry practices, especially in the Mediterranean mountains where the characteristic multifunctional has largely been lost. The article analyzes land uses changes, dynamics, and their morphological features between the first half of the 20th Century (1956) and the second decade of the present time (2019). The paper was divided into three sections. First, the identification of land uses using orthophotograph and aerial photograph; after that the info was checked with fieldwork. Eleven categories were identified according to the dominant use and land use changes and size of land parcels were taken into consideration. Second, the configuration and the information collected through the type and intensity of change in land uses made it possible to recognise and quantify their distribution and trend between these two dates. Also, the kernel density algorithm available in the Arcgis 10.5 software was used to obtain density and changes in land parcels. Finally, an overview is given of the main role that this agroforestry plays due to the social, ecological, and economic benefits that they provide for allowing sustainable development.

Highlights • We focused on tree-ring width (RWI) and potential evapotranspiration index (SPEI). • The relationship between RWI and SPEI was stronger under drier conditions. • SPEI of November and June were the most important predictors explaining RWI. • Beech tree growth was controlled by soil water supply and atmospheric evaporative demand. Abstract Due to the recent spread of forest die-off worldwide, concerns arise about the relative influence of specific climate parameters on tree growth decline in semi-arid environments, such as the Mediterranean mountain forests. As temperatures increase, drought may reduce tree productivity and survival across these forest ecosystems. Drought-induced tree growth decline can be interpreted as an early-warning signal of forest vulnerability. Here, we modeled the relationship between tree-ring width index (RWI) of beech populations in mountain forests of south-central Italy and the standardized precipitation evapotranspiration index (SPEI) derived from local weather stations. The aim of the research was to propose a procedure to determine the tipping point of drought severity triggering tree decline in beech forests. We focused on the cumulative water balance over the previous 1 month, which was particularly appropriate in relationship with the soil water conditions of these Mediterranean mountains. Under drier conditions, the correlation between RWI and SPEI was stronger, soil water supply (early fall of current year) and atmospheric evaporative demand (late spring of current year) being the dominant factors limiting tree growth of southern beech populations. [ABSTRACT FROM AUTHOR]

Abstract This paper explores the dynamics of temporal evolution of the high mountain Mediterranean grasslands, (Sierra Nevada, Spain SE). The indicator used is the emission of pollen (Pollen Index, PI) with respect to two important aspects: the incidence of the snow dynamic together with other hydro-meteorological parameters, and the changes in land use, which can Influence the evolution of the grasslands throughout time. The results reveal that pollen emissions in the last 25 years have shown a slight downward trend, with large interannual fluctuations, which are a consequence of diverse environmental factors, both general and specific to the area. One of the most influential parameters on pollen concentrations is snow cover, which reinforces the importance of the presence of snow-packs as water resource outside the winter season in the High Mediterranean Mountain environments. The changes in land use experienced in the area are a driver of change, especially due to the losses experienced in the last decades in the preferred habitats for many species of grasses. It can be concluded that the vulnerability of these ecosystems will be affected by an increase in winter temperatures and/or a decrease in rainfall (climate change) and an increase in the intensity of anthropogenic activities on land use. In this context, the PI is shown as a useful indicator of global change given its sensitivity to both anthropic and hydro-meteorological changes. In addition, it has a wide range of spatial detection and discrimination capacity by altitudinal dimensions. Graphical abstract Unlabelled Image Highlights • Temporal evolution of the high mountain Mediterranean grasslands was explored. • Grass pollen emissions were used as indicators of response to environmental changes. • Snow-packs outside the winter season is one of the most influential parameters on the pollen index. • Changes in land use in the preferred habitats of grasses are also driver of change. [ABSTRACT FROM AUTHOR]

Background: High-mountain ecosystems are centres of plant diversity that are particularly sensitive to land-use and climate change. Aims: We investigated the ecological trends associated with land use and climate change since the 1950s in different vegetation types in high-mountain habitats in the central Apennines. Methods: We analysed temporal changes in: Pinus mugo scrub, calcareous subalpine grasslands and alpine scree vegetation, comparing historical and recent vegetation records from vegetation plots from two periods (1955-1980 and 1990-2014) for their ecological indicator values (Landolt temperature and nutrient indicators) and structural traits (growth forms) over time using generalised linear models (GLMs). Results: We observed significant temporal differences in the ecology and structure of the analysed habitats. In the Pinus mugo scrub we detected a reduction of subalpine and herbaceous species and in calcareous alpine screes we observed an increment of the lower montane, montane and subalpine species and of dwarf shrubs. Conversely, subalpine grasslands were stable over time. Conclusions: Ecological changes that have occurred in the Central Apennines, following changes in type and intensity of land use and recent warming are consistent with those observed in other European mountains, for which climate and land-use changes are claimed as the main driving forces. [ABSTRACT FROM AUTHOR]

Abstract: Land abandonment is a global issue with important implications in Mediterranean mountain areas. Abandoned Mediterranean croplands start a process of secondary succession that is initially colonized by grasslands, shrubs and forest. In Mediterranean mountain areas, the process is very slow, so the shrubs remain for decades, preventing livestock from accessing pastureland. Therefore, farmers have to burn or clear the shrubs in order to provide pasture, a practice that has recently been encouraged by several regional governments in Spain. Data from experimental plots of the Aísa Valley Experimental Station in the Spanish Pyrenees allow to evaluate the effects of burning and clearing shrubs on physical and chemical soil properties, runoff production and soil erosion rates. The results suggested that clearing shrubs returned more positive results than burning, as it improved soil quality and slowed soil erosion while producing slightly higher runoff coefficients, which is very important in Mediterranean environments where water is a scarce resource. Clearing shrubs improves soil characteristics by increasing organic matter and CN ratio, which promote the expansion of herbaceous species with a high pastoral value. The results suggested that the policy of clearing shrubs is suitable for managing abandoned lands in Mediterranean mountain areas. Further studies at catchment scale will be needed to confirm the impact of substituting shrubs for meadows and to understand the connectivity of the flows measured from pedon and slope to the watershed scale. Copyright © 2016 John Wiley & Sons, Ltd. [ABSTRACT FROM AUTHOR]

This paper aims to characterize small ruminant farming systems in the Moroccan mountain regions and to assess undergoing socio-economic and ecological changes. Livestock farming systems in these regions are extensive with large flock sizes, especially sheep and goats raised mainly for meat production. Sheep producers utilized the inherent genetic diversity of their stock and developed adaptations to cope with seasonal availability of the mountain pastures and the need to diversify their own activities. Nowadays, farming systems in mountainous regions are undergoing socio-economic and ecological transformations that impact directly or indirectly on natural resources. Reduction and/or disappearance of herd transhumance, where flocks were moved seasonally from one climatic zone/vegetation type to another, was an important feature of the Middle Atlas sixty years ago Along with this reduced contribution from natural rangelands, other external resources and agricultural by-products (straw, stubble) are offered which make the system heavily dependent on agriculture. Intensification of feeding systems is associated with a choice for more productive but less hardy breeds from lowland areas. As a consequence of the reduction in mobility and the sedentarization of herds, there is an increase in the grazing pressure causing new and significant degradation of rangeland resources. Other profound institutional modifications concern the management and the utilization of the mountainous pasture. Indeed, the power of the group and the institutions for managing the resources is being replaced by the individual initiatives. [ABSTRACT FROM AUTHOR]

The analysis of past and present patterns of agrarian mountain areas allows researchers to characterize the influence of landscape heterogeneity on biodiversity, cultural heritage, and forest fire hazard. This process was mapped, quantified, and described through the use of digital mapping (GIS) and landscape indexes in a protected area in Alta Garrotxa (Catalonia, Spain). These areas require urgent management and modelling to provide alternative management scenarios, in order to maintain and recover habitats. A set of different scenarios have been designed using a multi-criteria evaluation and geospatial information available for the study area to identify the key areas for management action and to predict the potential effects on agricultural lands by prioritizing one or another management objective: biodiversity, landscape structure and perception, cultural heritage, fire hazard, and management cost. The observed progressive land abandonment of open areas with a small size and greater isolation will have a large impact on biodiversity and cultural heritage, and increase fire risk. Sustainable development will require planning objectives compatible with the conservation of biodiversity and the preservation of Mediterranean features with support for agricultural activities. This methodology can contribute to and be easily implemented by land managers, which could help to strengthen the link between managers and stakeholders. [ABSTRACT FROM AUTHOR]

We assessed the effects of landscape change on the climate regulation ecosystem service in a mountain river basin of Portugal, through the quantification, valuation and mapping of carbon sequestration and storage. The analyses were based on land use and land cover (LULC) changes that took place between 1990 and 2006 and on expected changes defined by three LULC change scenarios for 2020. We used the Integrated Valuation of Ecosystem Services and Tradeoffs model for scenario building and carbon assessment and valuation, and several modelling tools to assess past, current and future carbon in four different pools. Soil organic carbon data was obtained through an extensive sampling scheme across the entire study area. Recent (1990–2006) and expected landscape changes (2006–2020) affected considerably carbon sequestration and storage. Observed landscape changes generally promoted carbon sequestration and storage, and had a positive effect on the climate regulation ecosystem service, both biophysically and economically. Expected LULC changes further extend the capability of the landscape to increase carbon sequestration and storage in the near future. The carbon sequestered and stored in vegetation and soil contributes to avoid socio-economic damages from climate change, while increasing the economic value of particular LULC classes and the whole landscape. These results are essential to inform land planning, especially on how, where and when changes in landscapes may affect the provision of the climate regulation ecosystem service. [ABSTRACT FROM PUBLISHER]

Human-shaped treelines are a common feature in mountain landscapes across Europe, and particularly in secularly managed Mediterranean high-elevation areas. The abandonment of traditional land use, and especially the reduction in grazing pressure at high elevations, triggered secondary succession in treeless grassland areas, and favored the upward shift of anthropogenic treelines in some cases. We investigated this process in four anthropogenic treeline ecotone sites in the Central Apennines, Italy, populated by European black pine ( Pinus nigra Arn.). The upward treeline shift was controlled by microsite topography and the proximity to plantations acting as seed source. We found a 50% probability of producing cones in trees with basal diameters of 15–25 cm, heights of 2–5 m and ages of 20–25 years. The role played by climate on growth and the recruitment processes seems to be secondary, or could be masked by human-shaped processes. The presence of reproductive age trees at the treeline, mainly growing on debris-rich and steep slope sites, could indicate that the recruitment process will increase in future, leading to patchy tree patterns at different elevations. The high growth and encroachment rates observed at these human-shaped treelines would indicate that general growth dynamics are speeding up, including the tree colonization of treeless areas. These succession processes could cause a significant long-term decline in plant diversity in species-rich grasslands. Nonetheless, tree encroachment could increase forest protection against landslides and avalanches in the context of global change. [ABSTRACT FROM AUTHOR]

Forests of the Apennines are characterised by high canopy cover and high tree species diversity (being at the interface between two major climatic zones of Europe), and provide important ecosystem functions to millions of people. They exemplify cutting-edge themes such as forest ecology in warmer climates, consequences of heavy land use, and resilience at the trailing edge of the distribution of many European forest species (Silver fir, Norway spruce, Beech, Black pine, Birch). We introduce the setting under the geological and climatological point of view and review the literature on the interactions between these long-term drivers and the specific, structural, and genetic diversity of these forest communities (e.g., effects of glacial refugia or tectonic/volcanic activity), followed by a brief outline of what little is known about natural disturbance regimes and their range of variability. Anthropogenic disturbances (fire, grazing) and land use changes (abandonment of cropland and pasture) have been by far the main drivers of forest dynamics at least for the last two millennia, determining for examples overageing of coppices, treeline advances, forest encroachment on former agricultural land. We suggest considerations about the interplay between these land use changes and disturbance drivers (e.g. fuel continuity), summarize comparisons between managed and unmanaged forests (e.g., increase in tree size, deadwood, biodiversity indicators), and elaborate on current proposals for climate-adapted management, highlighting specific and genetic diversity as an important source of resilience and adaptive potential. [ABSTRACT FROM AUTHOR]

• Cropland abandonment significantly affects soil physico-chemical properties. • Differences between revegetated practices after farmland abandonment occurred. • Afforestation triggered higher soil organic carbon than natural revegetation. • Grassland sites enhance the accumulation of soil organic carbon. • Labile fractions are higher in both vegetated sites than in grasslands. Mediterranean mountains are sensitive agroecosystems that have suffered intense land use and land cover changes (LULCC) during the last century. From the middle of the twentieth century, most of the cultivated lands in Mediterranean mountains were abandoned, allowing the recovery of vegetation (through natural revegetation and afforestation programmes). To examine the effects of farmland abandonment, secondary succession (natural revegetation) and afforestation, an intensive soil sampling was carried out in the Araguás catchment (Central Spanish Pyrenees) including sparsely vegetated areas (badlands), grasslands, shrublands and afforested sites. LULCC were mapped, and soil physico-chemical properties were analysed in reference sites (unaltered areas during the last centuries) and in the different land uses. Likewise, the soil organic carbon (SOC) content in the bulk soils and in the fractions separated by density fractionation have been studied. This study evidenced that farmland abandonment led to a mosaic landscape with different land use and land covers. Results show that LULCC significantly affect soil physico-chemical properties (soil texture, stoniness, pH, SOC, total carbon, CorgN ratio, bulk density and field capacity). Significant differences were observed between secondary and afforested sites following farmland abandonment. Afforestation triggered higher SOC than shrubland sites (natural revegetation) (1.4 and 1.1% respectively), suggesting a slower process of organic matter accumulation after farmland abandonment in the natural revegetation compared to afforestation. The significant role of grassland sites for enhancing the accumulation of SOC has been also confirmed. The results showed also significant differences in the relative contribution of each organic fraction to the bulk SOC: the amount of labile fraction (free and occluded labile fractions) is significantly higher in afforested and shrubland sites (58.1 and 51.2% respectively) than in grassland sites (36.8%). Understanding the effects of LULCC on soil properties and SOC dynamics is essential when planning post-land management practices after farmland abandonment. [ABSTRACT FROM AUTHOR]

Time compression was studied at the Aísa Valley Experimental Station to assess the effect of large rainfall events on suspended sediment load. The study was focused on the response of land under traditional and current land uses/land covers in Mediterranean mountain areas, including dense shrub cover, grazing meadow, abandoned field, cereal cultivation, fallow land and both active and abandoned shifting agriculture. The results confirm the importance of the largest rainfall events in explaining the geomorphological effects that occur during a small number of such events. Three groups of land uses were distinguished: (i) dense shrub cover and grazing meadow, which showed relatively little time compression; (ii) cereal cultivation, fallow land, and both active and abandoned shifting agriculture, which were characterized by a high degree of time compression and a major impact of the largest events; and (iii) the abandoned field, which was intermediate in response between the other two groups. The greater geomorpho-logical effects of the largest events in the plots at the Aísa Valley Experimental Station relative to those reported in studies in the USA, is a consequence of the Mediterranean climate charac-teristics, and in part because of scale effects. The contribution of the three largest events to total accumulated suspended sediment load ranged from 25.4 to 71.2% whereas for the five largest events it ranged from 35.4 to 76.9%. For each of the land uses the 25 largest events represented more than 80% (in some cases more than 90%) of the total suspended sediment load recorded since 1991.