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Fossil discoveries of early Australopithecus species from Woranso-Mille have played a significant role in improving our understanding of mid-Pliocene hominin evolution and diversity. Here, we describe two mandibles with dentitions, recovered from sediments immediately above a tuff radiometrically dated to 3.76 ± 0.02 Ma, and assess their taxonomic affinity. The two mandibles (MSD-VP-5/16 and MSD-VP-5/50) show morphological similarities with both Australopithecus anamensis and Australopithecus afarensis. Some of the unique features that distinguish Au. anamensis from Au. afarensis are present in the mandibles, which also share a few derived features with Au. afarensis. Their retention of more Kanapoi Au. anamensis-like traits, compared to the fewer derived features they share with Au. afarensis, and the presence of Au. anamensis at Woranso-Mille in 3.8-million-year-old deposits, lends support to their assignment to Au. anamensis. However, it is equally arguable that the few derived dentognathic features they share with Au. afarensis could be taxonomically more significant, making it difficult to conclusively assign these specimens to either species. Regardless of which species they are assigned to, the mosaic nature of the dentognathic morphology and geological age of the two mandibles lends further support to the hypothesized ancestor-descendant relationship between Au. anamensis and Au. afarensis. However, there is now limited fossil evidence indicating that these two species may have overlapped in time. Hence, the last appearance of Au. anamensis and first appearance of Au. afarensis are currently unknown. Recovery of Australopithecus fossils from 4.1 to 3.8 Ma is critical to further address the timing of these events.

Author(s): Park, Y; Swanson-Hysell, NL; MacLennan, SA; Maloof, AC; Gebreslassie, M; Tremblay, MM; Schoene, B; Alene, M; Anttila, ESC; Tesema, T; Haileab, B | Abstract: The Tonian-Cryogenian Tambien Group of northern Ethiopia is a mixed carbonate-siliciclastic sequence that culminates in glacial deposits associated with the first of the Cryogenian glaciations-the Sturtian "Snowball Earth. " Tambien Group deposition occurred atop arc volcanics and volcaniclastics of the Tsaliet Group. New U-Pb isotope dilution-thermal ionization mass spectrometry (ID-TIMS) dates demonstrate that the transition between the Tsaliet and Tambien Groups occurred at ca. 820 Ma in western exposures and ca. 795 Ma in eastern exposures, which is consistent with west to east arc migration and deposition in an evolving back-arc basin. The presence of intercalated tuffs suitable for high-precision geochronology within the Tambien Group enable temporal constraints on stratigraphic data sets of the interval preceding, and leading into, the Sturtian glaciation. Recently discovered exposures of Sturtian glacial deposits and underlying Tambien Group strata in the Samre Fold-Thrust Belt present the opportunity to further utilize this unique association of tuffs and carbonate lithofacies. U-Pb ID-TIMS ages from zircons indicate that Tambien Group carbonates were deposited from ca. 820 Ma until 0-2 m.y. before the onset of the Sturtian glaciation, making the group host to a relatively complete carbonate stratigraphy leading into this glaciation. New δ13C and 87Sr/86Sr data and U-Pb ID-TIMS ages from the Tambien Group are used in conjunction with previously published isotopic and geochronologic data to construct newly time-calibrated composite Tonian carbon and strontium isotope curves. Tambien Group δ13C data and U-Pb ID-TIMS ages reveal that a pre-Sturtian sharp negative δ13C excursion (referred to as the Islay anomaly in the literature) precedes the Sturtian glaciation by-18 m.y., is synchronous in at least two separate basins, and is followed by a prolonged interval of positive δ13C values. The composite Tonian 87Sr/86Sr curve shows that, following an extended interval of low and relatively invariant values, inferred seawater 87Sr/86Sr rose ca. 880-770 Ma, then subsequently decreased leading up to the ca. 717 Ma initiation of the Sturtian glaciation. These data, when combined with a simple global weathering model and analyses of the timing and paleolatitude of large igneous province eruptions and arc accretion events, suggest that the 87Sr/86Sr increase was influenced by increased subaerial weathering of radiogenic lithologies as Rodinia rifted apart at low latitudes. The following 87Sr/86Sr decrease is consistent with enhanced subaerial weathering of arc lithologies accreting in the tropics over tens of millions of years, lowering pCO2 and contributing to the initiation of the Sturtian glaciation.

Several hypotheses posit a link between the origin of Homo and climatic and environmental shifts between 3 and 2.5 Ma. Here we report on new results that shed light on the interplay between tectonics, basin migration and faunal change on the one hand and the fate of Australopithecus afarensis and the evolution of Homo on the other. Fieldwork at the new Mille-Logya site in the Afar, Ethiopia, dated to between 2.914 and 2.443 Ma, provides geological evidence for the northeast migration of the Hadar Basin, extending the record of this lacustrine basin to Mille-Logya. We have identified three new fossiliferous units, suggesting in situ faunal change within this interval. While the fauna in the older unit is comparable to that at Hadar and Dikika, the younger units contain species that indicate more open conditions along with remains of Homo. This suggests that Homo either emerged from Australopithecus during this interval or dispersed into the region as part of a fauna adapted to more open habitats., Key events in human evolution are thought to have occurred between 3 and 2.5 Ma, but the fossil record of this period is sparse. Here, Alemseged et al. report a new fossil site from this period, Mille-Logya, Ethiopia, and characterize the geology, basin evolution and fauna, including specimens of Homo.

Author(s): MacLennan, S; Park, Y; Swanson-Hysell, N; Maloof, A; Schoene, B; Gebreslassie, M; Antilla, E; Tesema, T; Alene, M; Haileab, B | Abstract: In order to understand the onset of Snowball Earth events, precise geochronology and chemostratigraphy are needed on complete sections leading into the glaciations. While deposits associated with the Neoproterozoic Sturtian glaciation have been found on nearly every continent, time-calibrated stratigraphic sections that record paleoenvironmental conditions leading into the glaciation are exceedingly rare. Instead, the transition to glaciation is normally expressed as erosive contacts with overlying diamictites, and the best existing geochronological constraints come from volcanic successions with little paleoenvironmental information. We report new stratigraphic and geochronological data from the upper Tambien Group in northern Ethiopia, which indicates that the glacigenic diamictite at the top of the succession is Sturtian in age. U-Pb zircon dates obtained from two tuffaceous siltstones that are 74 and 84 m below the diamictite are 719.68 ± 0.46 Ma and 719.68 ± 0.56 Ma (2σ), respectively. We also report a U-Pb date of 735.25 ± 0.25 Ma from a crystal-rich tuff located 2 m above the nadir of a high-amplitude, basin-wide, negative δ13C excursion previously correlated with the Islay anomaly. This age for the anomaly agrees with Re-Os age constraints from Laurentia, suggesting that the δ13C signal is globally synchronous and preceded the Sturtian glaciation by ~18 m.y. The interval between the Islay anomaly and Sturtian glaciation is recorded in the Tambien Group as an ~600 m succession of predominantly shallow-water carbonates and siliciclastics with δ13C values recording a prolonged period at +5‰, followed by an interval of lower, but still positive, values leading up to the glaciation. Our data are consistent with synchronous global onset of the Sturtian glaciation at ca. 717 Ma. Shallow-water carbonates in strata directly below the first diamictite suggest that glacial onset was rapid in terranes of the Arabian-Nubian Shield.

The Woranso–Mille (WORMIL) area in the west-central Afar, Ethiopia, contains several Pliocene basalt flows, tuffs, and fossiliferous volcaniclastic beds. We present whole-rock major- and trace-element data including REE, and Sr–Nd–Pb isotope ratios from these basalts to characterize the geochemistry, constrain petrogenetic processes, and infer mantle sources. Six basalt groups are distinguished stratigraphically and geochemically within the interval from ~ 3.8 to ~ 3 Ma. The elemental and isotopic data show intra- and inter-group variations derived primarily from source heterogeneity and polybaric crystallization ± crustal inputs. The combined Sr–Nd–Pb isotope data indicate the involvement of three main reservoirs: the Afar plume, depleted mantle, and enriched continental lithosphere (mantle ± crust). Trace element patterns and ratios further indicate the basalts were generated from spinel-dominated shallow melting, consistent with significantly thinned Pliocene lithosphere in western Afar. The on-land continuation of the Aden rift into western Afar during the Pliocene is reexamined in the context of the new geochemistry and age constraints of the WORMIL basalts. The new data reinforce previous interpretations that progressive rifting and transformation of the continental lithosphere to oceanic lithosphere allows for increasing asthenospheric inputs through time as the continental lithosphere is thinned.

International audience; A fossil hominin cranium was discovered in mid-Pliocene deltaic strata in the Godaya Valley of the northwestern Woranso-Mille study area in Ethiopia. Here we show that analyses of chemically correlated volcanic layers and the palaeomagnetic stratigraphy, combined with Bayesian modelling of dated tuffs, yield an age range of 3.804 ± 0.013 to 3.777 ± 0.014 million years old (mean ± 1σ) for the deltaic strata and the fossils that they contain. We also document deposits of a perennial lake beneath the deltaic sequence. Mammalian fossils associated with the cranium represent taxa that were widespread at the time and data from botanical remains indicate that the vegetation in the lake and delta catchment was predominantly dry shrubland with varying proportions of grassland, wetland and riparian forest. In addition, we report high rates of sediment accumulation and depositional features that are typical of a steep topographic relief and differ from younger Woranso-Mille fossil localities, reflecting the influence of active rift processes on the palaeolandscape. Q1 Until recently, field work in the Woranso-Mille area (Afar, Ethiopia) has concentrated on exposures along the Mille River that include more than 120 m of fossiliferous sedimentary and volcanic strata that are between 3.8 and 3.2 million years (Myr) old. East of Korsi Dora (Fig. 1a), sedimentary sequences that are younger than the extra-regional approximately 3.57-Myr-old Kilaytoli tuff 1 have yielded specimens of at least two hominin species, Australopithecus afarensis and Australopithecus deyiremeda 2-5

A fossil hominin cranium was discovered in mid-Pliocene deltaic strata in the Godaya Valley of the northwestern Woranso-Mille study area in Ethiopia. Here we show that analyses of chemically correlated volcanic layers and the palaeomagnetic stratigraphy, combined with Bayesian modelling of dated tuffs, yield an age range of 3.804 ± 0.013 to 3.777 ± 0.014 million years old (mean ± 1σ) for the deltaic strata and the fossils that they contain. We also document deposits of a perennial lake beneath the deltaic sequence. Mammalian fossils associated with the cranium represent taxa that were widespread at the time and data from botanical remains indicate that the vegetation in the lake and delta catchment was predominantly dry shrubland with varying proportions of grassland, wetland and riparian forest. In addition, we report high rates of sediment accumulation and depositional features that are typical of a steep topographic relief and differ from younger Woranso-Mille fossil localities, reflecting the influence of active rift processes on the palaeolandscape.

Fossiliferous deposits at Woranso-Mille span the period when Australopithecus anamensis gave rise to Australopithecus afarensis (3.8–3.6 Ma) and encompass the core of the A. afarensis range (ca. 3.5–3.2 Ma). Within the latter period, fossils described to date include the intriguing but taxonomically unattributed Burtele foot, dentognathic fossils attributed to Australopithecus deyiremeda, and one specimen securely attributed to A. afarensis (the Nefuraytu mandible). These fossils suggest that at least one additional hominin lineage lived alongside A. afarensis in the Afar Depression. Here we describe a collection of hominin fossils from a new locality in the Leado Dido’a area of Woranso-Mille (LDD-VP-1). The strata in this area are correlated to the same chron as those in the Burtele area (C2An.3n; 3.59–3.33 Ma), and similar in age to the Maka Sands and the Basal through lower Sidi Hakoma Members of the Hadar Formation. We attribute all but one of the LDD hominin specimens to A. afarensis, based on diagnostic morphology of the mandible, maxilla, canines, and premolars. The LDD specimens generally fall within the range of variation previously documented for A. afarensis but increase the frequency of some rare morphological variants. However, one isolated M3 is extremely small, and its taxonomic affinity is currently unknown. The new observations support previous work on temporal trends in A. afarensis and demonstrate that the large range of variation accepted for this species is present even within a limited spatiotemporal range. The value added with this sample lies in its contribution to controlling for spatiotemporal differences among site samples in the A. afarensis hypodigm and its contemporaneity with non-A. afarensis specimens at Woranso-Mille.

Middle Pliocene hominin species diversity has been a subject of debate over the past two decades, particularly after the naming of Australopithecus bahrelghazali and Kenyanthropus platyops in addition to the well-known species Australopithecus afarensis. Further analyses continue to support the proposal that several hominin species co-existed during this time period. Here we recognize a new hominin species (Australopithecus deyiremeda sp. nov.) from 3.3-3.5-million-year-old deposits in the Woranso-Mille study area, central Afar, Ethiopia. The new species from Woranso-Mille shows that there were at least two contemporaneous hominin species living in the Afar region of Ethiopia between 3.3 and 3.5 million years ago, and further confirms early hominin taxonomic diversity in eastern Africa during the Middle Pliocene epoch. The morphology of Au. deyiremeda also reinforces concerns related to dentognathic (that is, jaws and teeth) homoplasy in Plio-Pleistocene hominins, and shows that some dentognathic features traditionally associated with Paranthropus and Homo appeared in the fossil record earlier than previously thought.

The Neoproterozoic Era was an interval characterized by profound environmental and biological transitions. Existing age models for Neoproterozoic nonglacial intervals largely have been based on correlation of carbonate carbon isotope values, but there are few tests of the assumed synchroneity of these records between basins. In contrast to the ash-poor successions typically targeted for Neoproterozoic chemostratigraphy, the Tonian to Cryogenian Tambien Group (Tigray region, Ethiopia) was deposited in an arc-proximal basin where volcanic tuffs suitable for U-Pb geochronology are preserved within the mixed carbonate-siliciclastic sedimentary succession. The Tambien Group culminates in a diamictite interpreted to correlate to the ca. 717–662 Ma Sturtian snowball Earth glaciation. New physical stratigraphic data and high-precision U-Pb dates from intercalated tuffs lead to a new stratigraphic framework for the Tambien Group that confirms identification of negative δ13C values from Assem Formation limestones with the ca. 800 Ma Bitter Springs carbon isotope stage. Integration with data from the Fifteenmile Group of northwestern Canada constitutes a positive test for the global synchroneity of the Bitter Spring Stage and constrains the stage to have started after 811.51 ± 0.25 Ma and to have ended before 788.72 ± 0.24 Ma. These new temporal constraints strengthen the case for interpreting Neoproterozoic carbon isotope variation as a record of large-scale changes to the carbon cycle and provide a framework for age models of paleogeographic change, geochemical cycling, and environmental evolution during the radiation of early eukaryotes.

A wide range of lithologic units and tectonic disturbances by cross-cutting faults and folds has resulted in the quite complex hydrogeological setting of the sedimentary outlier and its surroundings at Mekelle, northern Ethiopia. The environmental isotopes of oxygen and hydrogen and patterns of dissolved ion concentrations in the groundwater, coupled with understanding of the three-dimensional geological framework, are used to conceptualize the groundwater flow model and recharge–discharge mechanisms in the area. In agreement with the piezometric-surface map, recharge areas are determined to be the highlands (northwest, north, east and south of the study area), characterized by relatively more depleted isotopic compositions, higher d-excess, and lower concentrations of dissolved ions in the groundwater samples; the narrow major river valleys of Giba, Illala, Chelekot and Faucea Mariam are discharge areas. The groundwater divide between the Tekeze and the Denakil basins coincides with the surface-water divide line of these two basins. In most cases, groundwater feeds the semi-perennial streams and rivers in the area. However, isotopic signatures in some wells indicate that there are localities where river flow and seepage from micro-dams locally feed the adjacent aquifers. The lithostratigraphic, geomorphologic, isotopic and hydrochemical settings observed in this study indicate that three groundwater flow systems (shallow/local, intermediate and deep/semi-regional) can exist here. Tritium data indicate that the groundwater in the study area has generally short residence time and is dependent on modern precipitation.

Eighty-one paleomagnetic cores were collected from 10 locations across a black limestone unit within the core of Negash Synclinorium, northern Ethiopia in order to test the proposed Snowball Earth events for the diamictite unit of the Tambien Group. Cores were cut into two standard paleomagnetic specimens and were subjected to stepwise demagnetizations using both Thermal (TH) and alternating field (AF) techniques. Rock magnetic analyses on representative specimens were done and results revealed goethite, pyrrhotite, titano-magnetite, and titano-hematite to be the major magnetic materials carrying the magnetizations with PSD (pseudo single domain) grain size range. In most cases paleomagnetic directions are defined by a single component of magnetization, where a viscous component is present it is usually removed by heating to a temperature of ∼200 °C or an AF of ∼10 mT. The high stability component isolated above temperature of 200 °C or AF of 15 mT, defined straight line trajectories directed towards the origin and considered as the Characteristic Remanent Magnetization Direction (ChRM). The direction of magnetization of the ChRM is determined for samples with stable straight line segments by the best-fit line using the least square technique of Kirschvink (1980). In the cases of overlapping spectra and unblocking temperatures, direction of magnetization is determined by remagnetization circles of Halls (1976, 1978). When site mean ChRM directions are plotted on stereogram, their distribution is relatively clustered in geographic coordinates and the overall mean direction is Dec g = 358.5°, Inc g = 16.6° ( α 95 = 3.8°, K = 162.8, N = 10). After a structural restoration to the horizontal is made the directions disperse and fail the fold test of both McElhinny’s and McFadden’s tests and the mean direction is Dec s = 353.5°, Inc s = 8.8° ( α 95 = 18.9°, K = 7.5, N = 10). This is interpreted to result from a later remagnetization of the black limestone. All directions are normal in polarity and have mean unrestored paleomagnetic directions comparable to the Quaternary paleomagnetic directions. Virtual Geomagnetic poles (VGP) in the unrestored position is used to calculate overall mean VGP position resulting lon g = 235.7°E, lat g = 84.5°N ( A 95 = 3.0°, N = 10). Comparison of the obtained pole with the apparent polar wander path (APWP) curve for Africa of Besse and Courtillot (1991, 2003) and with the 2 Ma reference pole of stable Africa (Kidane et al., 2003) is found to be consistent with remagnetizations during the Quaternary period. Hence supporting evidence for the proposed Snowball Earth event of the Sturtian glaciation in the Negash rocks could not, unfortunately, be obtained from paleomagnetism.

Australopithecus afarensis is the best-known and most dimorphic species in the early hominin fossil record. Here, we present a comparative description of new fossil specimens of Au. afarensis from Nefuraytu, a 3.330-3.207 million-years-old fossil collection area in the Woranso-Mille study area, central Afar, Ethiopia. These specimens include NFR-VP-1/29, one of the most complete mandibles assigned to the species thus far and among the largest mandibles attributed to Au. afarensis, likely representing a male individual. NFR-VP-1/29 retains almost all of the distinctive archaic features documented for Au. afarensis. These features include a posteriorly sloping symphysis, a low and rounded basally set inferior transverse torus, anterosuperiorly opening mental foramen, a lateral corpus hollow bound anteriorly by the C/P3 jugae and posteriorly by the lateral prominence, and the ascending ramus arising high on the corpus. Dental morphology and metrics of the Nefuraytu specimens also falls within the range of Au. afarensis. The presence of this species at Woranso-Mille between 3.330 and 3.207 million years ago confirms the existence of this species in the area in close spatial and temporal proximity to other middle Pliocene hominin taxa such as the one represented by the Burtele foot (BRT-VP-2/73) and the recently named species Australopithecus deyiremeda. This has important implications for our understanding of middle Pliocene hominin diversity.

(40)Ar/(39)Ar dating of tuffs and mafic lavas, tephra geochemistry, and paleomagnetic reversal stratigraphy have been used to establish the chronostratigraphy of the Pliocene hominid-bearing fossiliferous succession at Woranso-Mille, a paleontological study area in the western part of the central Afar region of Ethiopia. The succession in the northwestern part of the study area ranges in (40)Ar/(39)Ar age from 3.82-3.570 Ma, encompassed by paleomagnetic subchron C2Ar (4.187-3.596 Ma). One of the major tuff units, locally named the Kilaytoli tuff, is correlative on the basis of age and geochemistry to the Lokochot Tuff of the Turkana Basin. A hominid partial skeleton (KSD-VP-1) was found in strata whose precise stratigraphic position and age is still under investigation, but is believed to correspond to the later part of this interval. Woranso-Mille fills a significant gap in the fossil record of northeastern Africa at the time of the lower to middle Pliocene transition, when many extant species lineages of African fauna were established.

Twenty obsidian artifacts from the terminal Pleistocene/early Holocene phase of the site of Beseka, central Ethiopia, have been instrumentally characterized using EDXRF and electron microprobe. Results show that most of the artifacts were derived from the silicic centers of Fentale and Kone. A nearby source of Abadir, although with better quality raw material, does not seem to have been utilized.

KSD-VP-1/1, a partial skeleton of Australopithecus afarensis , was excavated from Pliocene strata at Korsi Dora, 3.3 km southeast of the confluence of the Waki and Mille rivers in the northwestern part of the Woranso-Mille paleoanthropological research site. A tuff collected from ~2.7 m below the fossil horizon, at the bottom of a trench dug 25 m to the east of the fossil excavation, yielded an 40Ar/39Ar age of 3.60 ± 0.03 Ma for anorthoclase feldspar. Strata in the trench and the fossil excavation site comprise a single normal magnetozone interpreted as part of the normal subchron C2An.3n, immediately above the Gauss/Gilbert paleomagnetic transition. Geologic mapping and tephrochemical analyses combined with paleomagnetic data place the fossil horizon and the trench section into local and regional stratigraphic context by constraining the partial skeleton to be younger than the Kilaytoli tuff (KT), a ~4 m thick vitric ash with an anorthoclase feldspar age of 3.570 ± 0.014 Ma. This unit is widely recognized at Korsi Dora, in collection areas north of the Waki-Mille confluence and outside the field area. The KT correlates with the Lokochot Tuff of the Omo-Turkana Basin in Kenya. Sedimentological features of the mudstone and sandstone in and near the excavation site are consistent with deposition in a floodplain or floodplain lake proximal to a stream channel.

Tephra geochemistry and (40)Ar/(39)Ar geochronology are reported for the Waki-Mille area in the northwestern part of the Woranso-Mille paleoanthropological project area in the west central Afar region of Ethiopia. Previous studies documented dentognathic fossils that are morphologically intermediate between Australopithecus anamensis and Australopithecus afarensis and some that are attributed to Australopithecus afarensis. Additional dentognathic remains from the study area were assigned to the newly identified species Australopithecus deyiremeda. These fossil hominin taxa were recovered from volcanic and sedimentary strata containing tuffs ranging in age from more than 3.77 million years ago (Ma) to less than 3.469 Ma. One of the tuffs was correlated based on geochemistry, feldspar mineralogy, and age to the Lokochot Tuff of the Omo-Turkana Basin of southern Ethiopia and Kenya. Variations in major and minor element abundances in volcanic glass demarcate ten geochemically distinct tuffs and tuff sequences, including three that are geochemically similar to widespread regional tuffs, specifically the Lomogol, Lokochot, and β- Tulu Bor/Sidi Hakoma tuffs. A new (40)Ar/(39)Ar age for the Waki Tuff, which is geochemically similar to the Lomogol Tuff, is 3.664 ± 0.016 Ma. Other tuffs in the Waki-Mille area are geochemically dissimilar to regional tuffs documented to date. Identification of tuffs based on character, stratigraphic position, and geochemistry refines local stratigraphic correlations and delineates the geographic distributions of precisely dated fossiliferous levels within the Waki-Mille area.

The source provenance of 10 Early Stone Age artifacts from the localities in Melka Konture has been determined by EDXRF. Results show that the early to mid-Pleistocene makers of the artifacts derived the raw material from a source located in their proximity, supporting the previously proposed short distance transport of raw material for the time period.

The Tambien Group in Tigrai, Ethiopia, comprises a number of inliers each containing 2–3 km thickness of interbedded carbonate and clastic sediments, capped in one inlier by a glaciogenic diamictite. A range of geochemical indices suggest near-pristine C- and Sr-isotope values are preserved and these, together with lithological variations, allow local correlation between these inliers and correlation with the global Neoproterozoic isotope stratigraphy. A composite section of the Tambien Group shows 13 C in carbonate of ∼+6‰ at its base, decreasing upwards to twin lows of ∼–4‰ separated by a brief excursion back to positive values, then rises again to a plateau of ∼+6‰ before finally decreasing sharply to −2‰ beneath the Negash diamictite at its top. No glaciogenic sediments are observed associated with the lower twinned negative anomalies. The 87 Sr/ 86 Sr values in carbonates change from ∼0.7063 in the lower units to ∼0.7067 in the upper units. The 13 C of organic matter changes little through the sequence (−24.2 ± 1.3‰), with the result that the C-isotope fractionation between carbonate and organic matter decreases from +26 to +21‰ up through the second low in carbonate 13 C before increasing to ∼+29.5‰ in the rest of the sequence. Together with existing radiometric age constraints, the Sr-isotope data indicate that the Negash diamictite is Sturtian in age, and the lower negative C-isotope anomalies appear to correlate with the non-glaciogenic Bitter Springs Stage recognised in Australia and Svalbard. According to this correlation the Tambien Group was deposited in the interval ca. 800–735 Ma. Our data reinforce the emerging view that not all Neoproterozoic negative C-isotope anomalies are associated with glaciation. The variations in C-isotope fractionations within our sequence contrast with those from Australia and are attributed to differences in local environmental variables, probably temperature. These differences are consistent with the recent proposal that the Bitter Springs Stage anomaly is the result of a pair of Inertial Interchange True Polar Wander events [Halverson, G.P., Maloof, A., Schrag, D., Dudas, F., Hurtgen, M., in press. Stratigraphy and geochemistry of a ca 800 Ma negative carbon isotope stage in northeastern Svalbard. Chem. Geol.] [Maloof, A.C., Halverson, G.P., Kirschvink, J.L., Schrag, D.P., Weiss, B.P., Hoffman, P.F., in press. Combined paleomagnetic, isotopic and stratigraphic evidence for true polar wander from the Neoproterozoic Akademikerbreen Group, Svalbard. GSA Bull.], and further high-resolution dating and palaeomagnetic studies of the Tambien Group should allow testing of this hypothesis.

Juvenile continental crust of the Arabian–Nubian Shield (ANS) formed within a Neoproterozoic supercontinent cycle. Subsequent late Neoproterozoic deposition overlapped a series of dramatic climatic events that are unparalleled in subsequent Phanerozoic time, as proposed by the “Snowball Earth” hypothesis. In particular, extreme negative δ 13 C excursions coincident with glacial diamictite and cap carbonate sequences imply that profound carbon flux changes accompanied widespread glacial transitions (Snowball Events). Such a succession appears to be partially preserved in metasediments of the diamictite-bearing Tambien Group of northern Ethiopia (Negash syncline). Here, a pronounced negative carbon excursion from values of +7 to −2 per mil accompanies the transition to diamictite deposition. New zircon evaporation dates from granites intruding the base of the sequence but post-dating early deformation of the entire sequence suggest that the Tambien Group is older than 613 Ma, and therefore diamictite genesis is incompatible with younger Marinoan, Varanger, or “Ediacaran” glacial intervals. Syn-tectonic granites elsewhere within the region date between 750 and 800 Ma and provide an older age constraint for the Tambien Group. The limestone sequence underlying the diamictite has an average 87 Sr / 86 Sr composition of 0.7066, which supports either a middle (ca. 750–740 Ma) or late (ca. 725–720 Ma) Sturtian age. Combined Sr and C isotopic compositions of the limestones and the upper age constraint from the granite ages provide a compelling argument that the Tambien Group sediments are ∼720–750 Ma or a little older and strongly support the prospect that the Negash diamictite is the product of Sturtian glaciation. If so, these are the first Sturtian Snowball Earth sequences identified in the ANS.

Whole-rock geochemical analysis of metavolcanic and plutonic rocks from Mai Kenetal-Negash area, Tigrai, northern Ethiopia was carried out to evaluate their magma type and original tectonic environment. Their major element content has been modified by secondary alteration, and trace and rare-earth elements have been used to investigate their petrogenesis. The low content of compatible elements of the metavolcanic rocks and their relatively high content of incompatible elements and light REE point to a depleted source region, whereas the plutonic rocks show a more pronounced REE fractionation (LaN/LuN >4). Metavolcanic and plutonic rocks seem to be cogenetic. Discriminant diagrams suggest that the majority of the metavolcanic and all the plutonic rocks are members of a calc-alkaline suite developed in a volcanic arc setting. The overall geochemical characteristics of both sets of rocks are consistent with the arc accretion models postulated in Sudan, Egypt and Saudi Arabia for the Neoproterozoic evolution of the Arabian-Nubian Shield.

Utilising geochemical data, various discriminant diagrams have been employed to establish the magma type and original tectonic environment for some Neoproterozoic amphibolites, ultrabasic rocks and gabbros of the Moyale area, southern Ethiopia. The gneissic amphibolites are found to have mixed geochemical characteristics indicative of island arc and/or ocean ridge basalts with tholeiitic composition whereas the porphyritic amphibolites show alkalic features with no clear tectonic setting. The ultrabasic and gabbroic units of the Moyale area are described in terms of their relation with mantle melts and parental material. The majority of ultrabasics relate to a cumulate origin and the gabbroic rocks appear as more differentiated magma from the same source. The mainly dunite bodies in the eastern sub-area at Moyale probably represent refractory residues left after variable degree of partial melting of a periodotite mantle. It is concluded that the gneissic amphibolites were probably part of an accreting arc associated with closure of a pre-existing oceanic basin. The ultrabasic and gabbroic rocks (together with the porphyritic amphibolite) are considered to be remnants of oceanic crust.

The Gademotta/Kulkuletti sites, located in the central part of the Main Ethiopian Rift Valley, represent the earliest Middle Stone Age (MSA) sequences in the country. Here we present the geochemical provenance of obsidian archaeological artefact recovered through excavation at the site. The artefacts and source materials were characterized by EDXRF and electron microprobe. Results show that the artefacts were procured from two sources, one local, and one presumably more distant, implying that despite the local availability of good quality raw material, not all obsidians were procured from a nearby source.

A newly discovered partial hominin foot skeleton from eastern Africa indicates the presence of more than one hominin locomotor adaptation at the beginning of the Late Pliocene epoch. Here we show that new pedal elements, dated to about 3.4 million years ago, belong to a species that does not match the contemporaneous Australopithecus afarensis in its morphology and inferred locomotor adaptations, but instead are more similar to the earlier Ardipithecus ramidus in possessing an opposable great toe. This not only indicates the presence of more than one hominin species at the beginning of the Late Pliocene of eastern Africa, but also indicates the persistence of a species with Ar. ramidus-like locomotor adaptation into the Late Pliocene.

The Precambrian tectonometamorphic evolution of the Moyale-Agal region, southernmost Ethiopia is described. This area forms part of the Pan-African, Mozambique Belt, forming the eastern margin of the Tanzanian Craton. It comprises polydeformed and metamorphosed mafic and ultramafic rocks, granodiorite, and subordinate amounts of metasedimentary rock. Four phases of deformation and two metamorphic events are recognised. The most pervasive phase, D1, is interpreted to have occurred during the Baragoian (c. 620 Ma) orogenic event. Peak metamorphism (M1) is synchronous with the development of the steep, N-S- or NW-SE-trending, S1 regional schistosity. In the Moyale sub-area amphibolite facies conditions prevailed, whereas in the Agal sub-area peak conditions did not exceed epidote-amphibolite facies. Plagioclase-hornblende thermometry gives estimates for peak metamorphic temperatures of 490°–650°C for the Moyale sub-area and

Only one partial skeleton that includes both forelimb and hindlimb elements has been reported for Australopithecus afarensis . The diminutive size of this specimen (A.L. 288-1 ["Lucy"]) has hampered our understanding of the paleobiology of this species absent the potential impact of allometry. Here we describe a large-bodied (i.e., well within the range of living Homo ) specimen that, at 3.58 Ma, also substantially antedates A.L. 288–1. It provides fundamental evidence of limb proportions, thoracic form, and locomotor heritage in Australopithecus afarensis . Together, these characteristics further establish that bipedality in Australopithecus was highly evolved and that thoracic form differed substantially from that of either extant African ape.

The phylogenetic relationship between Australopithecus anamensis and Australopithecus afar- ensis has been hypothesized as ancestor-descendant. However, the weakest part of this hypothesis has been the absence of fossil samples between 3.6 and 3.9 million years ago. Here we describe new fossil specimens from the Woranso-Mille site in Ethiopia that are directly rele- vant to this issue. They derive from sediments chrono- metrically dated to 3.57-3.8 million years ago. The new fossil specimens are largely isolated teeth, partial mandi- bles, and maxillae, and some postcranial fragments. However, they shed some light on the relationships between Au. anamensis and Au. afarensis. The dental morphology shows closer affinity with Au. anamensis from Allia Bay/Kanapoi (Kenya) and Asa Issie (Ethiopia) than with Au. afarensis from Hadar (Ethiopia). However, they are intermediate in dental and mandibular mor- phology between Au. anamensis and the older Au. afar- ensis material from Laetoli. The new fossils lend strong support to the hypothesized ancestor-descendant rela- tionship between these two early Australopithecus spe- cies. The Woranso-Mille hominids cannot be unequivo- cally assigned to either taxon due to their dental mor- phological intermediacy. This could be an indication that the Kanapoi, Allia Bay, and Asa Issie Au. anamensis is the primitive form of Au. afarensis at Hadar with the Laetoli and Woranso-Mille populations sampling a mosaic of morphological features from both ends. It is particularly difficult to draw a line between Au. anamen- sis and Au. afarensis in light of the new discoveries from Woranso-Mille. The morphology provides no evidence that Au. afarensis and Au. anamensis represent distinct taxa. Am J Phys Anthropol 141:406-417, 2010. V C 2009