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This article explores the importance of national governments and national-local relationships for scaling up local climate action to achieve global goals. From the Sustainable Development Goals to the New Urban Agenda to the Paris Climate Change Agreement, the achievement of global sustainability goals will depend on deep changes to national infrastructure and urban systems. Through an analysis of climate action planning and investments in exemplary cases in Africa, the paper highlights the opportunities and challenges that come with integrating national governments into urban-focused priorities and needs, especially for mobilising financial resources. The paper finds that scaling up city climate action in the selected African countries benefits from a constructive multi-level relationship between local and national institutions and stakeholders to shape and improve the legislative, financial and operating frameworks to enable systemic change. Large-scale urban climate action can be enabled by formal multi-level institutional arrangements, links to politically prioritised policy frameworks, and transformative aims that address both climate and socio-economic benefits for communities. [ABSTRACT FROM AUTHOR]