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The ponerine ant Anochetus miosumatrensis Ngô-Muller, Garrouste& Nel, n. sp. is described from a fossil alate female preserved in amber of Sumatra which is reputedly of Miocene age. On the basis of the general morphology, the fossil could be attributed to the extant Sumatran species group risii Brown, 1978. By comparing with the living environment of the extant species, this ant probably lived in a warm humid forest where it was trapped in dipterocarpacean resin during nuptial flight. Until now, the known Cenozoic distribution of the genus Anochetus was restricted to the Neotropical region. Thus A.miosumatrensis Ngô-Muller, Garrouste& Nel, n. sp. brings the first record of the genus from Indomalaya biogeographic region.

MEGALOPSENELLA GEN. NOV. urn:lsid: zoobank.org: act:081F52D1-52404FC0-B022-66BE5E7CFB74. Type species: Megalopsenella pouilloni. sp. nov. Etymology: The generic name is composed of the Greek prefix μεγάλο, large, and the ending of the name Holopsenella Engel et al., 2016, in turn derived from ψινέλλα, brushes. Gender feminine. Diagnosis: Female, antennae with 11 flagellomeres; mandibles long; propodeum nearly straight, without propodeal spines and sculpture; notauli slightly impressed; stout femora; forewing cell 1R1, broad and short, with vein Rs clearly shorter than Rs + M; cell 2R1 elongate, nearly reaching wing apex; distal part of Rs straight, making angle with straight 2r-rs; hindwing with 5 hamuli., Published as part of Jouault, Corentin, Ngô-Muller, Valérie, Pouillon, Jean-Marc & Nel, André, 2021, New Burmese amber fossils clarify the evolution of bethylid wasps (Hymenoptera: Chrysidoidea), pp. 1044-1058 in Zoological Journal of the Linnean Society 191 (4) on page 1049, DOI: 10.1093/zoolinnean/zlaa078, http://zenodo.org/record/4681287, {"references":["Engel MS, Ortega-Blanco J, Azevedo CO. 2016. A new bethylid wasp in Lebanese Early Cretaceous amber (Hymenoptera: Chrysidoidea), with comments on other Mesozoic taxa. American Museum Novitates 3855: 1 - 14."]}

CRETAPRISTOCERA LONGISCAPA SP. NOV. (FIGS 2–4) u r n: l s i d: z o o b a n k. o r g: a c t: 2 1 A 7 C A E C - 4 2 5 0 45A9-A3B3-28F099EAEC25. Holotype: Female, accession number IGR.BU-009, nearly complete and well-preserved female specimen in a rectangular piece of amber measuring 9 × 7 × 2.5 mm, together with a small Hymenoptera: Chalcidoidea; housed in the amber collection of the Geological Department and Museum (IGR) of the University of Rennes, France. Type locality and horizon: Noije Bum Hill, Hukawng Valley, Kachin State, Myanmar; Lower Cenomanian, Upper Cretaceous. Etymology: The specific epithet is derived from Latin longus, long, and scapus, shaft, for the elongated scape of this species. Description: Female; body 2.1 mm long (as preserved), not depressed, almost smooth and not strongly foveolate; LFW 1.5 mm; castaneous; wings setose. LH 0.43 mm; WH 0.5 mm; WF 0.26 mm; HE 0.11 mm; OOL 0.25 mm; WOT 0.16 mm; DAO 0.04 mm; VOL 0.1 mm. Head not pubescent, with small, dorsal, median depression, finely punctuated but not sculptured; mandible with one strong and sharp tooth, projecting forward; six maxillary palpomeres, three (?) visible labial palpomeres; clypeus spined; antenna with ten flagellomeres with cylindrical crosssection; scape 0.37 × 0.07 mm, pedicel 0.13 × 0.04 mm (all measurements refer to length × width), F1 0.07 × 0.05 mm, F2 0.05 × 0.05 mm, F3 0.05 × 0.06 mm, F 4 0.06 × 0.07 mm, F5 0.06 × 0.08 mm, F 6 0.08 × 0.08 mm, F7 0.08 × 0.08 mm, F8 0.08 × 0.07 mm, F9 0.09 × 0.06 mm, F10 0.17 × 0.06 mm; occipital carina strongly marked; compound eye without microsetae; ocellar triangle not equilateral, with two posterior ocelli more distant from each other than from anterior. Mesosoma with dorsum smooth, not roughly sculptured; femora strongly thickened, tibia thin, tarsal claws simple; tibial spur formula 1-2-2; notauli well marked, posteriorly convergent; parapsidal furrows weak; metanotum developed medially and overlapping mesoscutellum posteriorly; metapectal–propodeal complex rectangular, not foveolate, with marginal carinae and no posterior spines; posterior margin of propodeum straight. Forewing covered with microsetae and bordered with small setae; anterior border not angularly incurved anterior to pterostigma; costal vein present; only veins C, Sc + R, Rs + M, 2r-rs&Rs, M + Cu, Cu and cells C (hardly visible), R and 2R1 present; vein cu-a absent and cell Cu opened; cell 2R1 distally opened; no basal stub of Rs + M; pterostigma long and narrow; poststigmal abscissa of R1 short, spectral veins not visible. Hindwing with only veins C and Sc + R and with three hamuli. Metasoma with short petiole; six tergites present (including three preserved detached from the body), T2 enlarged, convex, partly covering sternites; sting not preserved; length of clearly visible tergites: T1 0.23 mm, T2 0.29 mm, T3 at least 0.19 mm. Remarks: It seems that the internal musculature of the fossil was preserved during the fossilization process, especially those of the legs (Figs 2–3). It could be interesting to compare the evolution of the musculature and the insertion of the muscles between the Cretaceous and extant Bethylidae in a future study, using 3D X-ray tomography.

MEGALOPSENELLA POUILLONI SP. NOV. (FIGS 5, 6) urn:lsid:zoobank.org:act: 578342A1-78A5-4F5E-9A08E5D3E1B46A83. Holotype: Female, accession number Po 55 (complete and well-preserved female specimen in a rounded piece of amber measuring 20 × 20 × 5 mm, without syninclusions), housed in the Musée d’Histoire Naturelle de Colmar, France. Type locality and horizon: Noije Bum Hill, Hukawng Valley, Kachin State, Myanmar; Lower Cenomanian, Upper Cretaceous. Etymology: Named after Jean-Marc Pouillon, who donated the holotype. Description: Female; body about 7 mm long, not depressed, almost smooth and not strongly foveolate; LFW 4.31 mm; castaneous; wings setose. LH 1.13 mm; WH 0.29 mm; WF 1.3 mm; HE 0.33 mm; OOL 0.55 mm; WOT 0.32 mm; DAO 0.14 mm; VOL 0.18 mm. Head not pubescent, smooth; mandible elongate, teeth not visible; six maxillary palpomeres, three (?) visible labial palpomeres; gena slightly projecting forward; reduced clypeus medially emarginate; antennal sockets partially closed dorsally by a tooth-shaped projection; antenna with 11 cylindrical flagellomeres; scape 0.47 × 0.22 mm, pedicel 0.1 2 × 0.1 2 mm, F1 0.1 4 × 0.1 2 mm (all measurements refer to length × width), F2 0.15 × 0.13 mm, F3 0.15 × 0.13 mm, F4 0.15 × 0.13 mm, F5 0.15 × 0.11 mm, F6 0.18 × 0.11 mm, F7 0.15 × 0.11 mm, F8 0.16 × 0.10 mm, F9 0.17 × 0.10 mm, F10 0.17 × 0.09 mm; F11 0.3 × 0.09 mm; occipital carina strongly marked; compound eye without microsetae; ocellar triangle not visible. Mesosoma with dorsum smooth, not sculptured; - Antenna with 12 antennomeres, scape elongated; forewing venation always reduced (Cenomanian: Myanmar)................................................................................................................... Cretapristocera gen. nov. 2. Head elongated with small eyes.......................................................................................................................3- Head almost globose with large eyes...............................................................................................................4 3. Mandibles only with few teeth (five) along masticatory margin; legs with pro- and meta-femora clearly thickened; forewing cell 2 R 1 open (Cenomanian: Myanmar)........................... Bethylitella Cockerell, 1917a - Mandibles with small teeth along masticatory margin and with basal one elongated: legs, except pro- femora, only slightly thickened; forewing cell 2 R 1 almost closed (Eocene: Baltic amber).............................................................................................................................................................. Protopristocera Brues, 1923 4. Notauli well developed, parapsidal furrow weak (Eocene: Oise amber).... Gynopteron Falières & Nel, 2019 - Notauli weak, parapsidal furrow well-visible (Eocene: Baltic amber)................. Bethylopteron Brues, 1933 Forewing covered with microtrichia and bordered with small setae; anterior border not angularly incurved anterior to pterostigma; seven closed cells present, and all veins C, Sc + R, Rs, M + Cu, Cu, A tubular; pterostigma rather broad; poststigmal abscissa of R1 long; hindwing 3.23 mm long with five hamuli. Metasoma c. 5.76 mm long, with short petiole; length of clearly visible tergites: T1 1.45 mm, T2 1 mm, T3 at least 0.8 mm; short part of sting visible. - Antenna with 12 antennomeres, scape elongated; forewing venation always reduced (Cenomanian: Myanmar)................................................................................................................... Cretapristocera gen. nov. 2. Head elongated with small eyes.......................................................................................................................3- Head almost globose with large eyes...............................................................................................................4 3. Mandibles only with few teeth (five) along masticatory margin; legs with pro- and meta-femora clearly thickened; forewing cell 2 R 1 open (Cenomanian: Myanmar)........................... Bethylitella Cockerell, 1917a - Mandibles with small teeth along masticatory margin and with basal one elongated: legs, except pro- femora, only slightly thickened; forewing cell 2 R 1 almost closed (Eocene: Baltic amber).............................................................................................................................................................. Protopristocera Brues, 1923 4. Notauli well developed, parapsidal furrow weak (Eocene: Oise amber).... Gynopteron Falières & Nel, 2019 - Notauli weak, parapsidal furrow well-visible (Eocene: Baltic amber)................. Bethylopteron Brues, 1933, Published as part of Jouault, Corentin, Ngô-Muller, Valérie, Pouillon, Jean-Marc & Nel, André, 2021, New Burmese amber fossils clarify the evolution of bethylid wasps (Hymenoptera: Chrysidoidea), pp. 1044-1058 in Zoological Journal of the Linnean Society 191 (4) on pages 1049-1051, DOI: 10.1093/zoolinnean/zlaa078, http://zenodo.org/record/4681287, {"references":["Cockerell TDA. 1917 a. Arthropods in Burmese amber. American Journal of Science ser. 4 44: 360 - 368.","Brues CT. 1923. Some new fossil parasitic Hymenoptera from Baltic amber. Proceedings of the American Academy of Arts and Sciences 58: 327 - 346.","Falieres E, Nel A. 2019. The first representative of the extinct flat wasp subfamily Protopristocerinae (Insecta: Hymenoptera, Bethylidae). Palaeoentomology 2: 322 - 326.","Brues CT. 1933. The parasitic Hymenoptera of the Baltic amber. Part 1. Bernstein-Forschungen (Amber Studies) 3: 4 - 179."]}

Antenna with 12 antennomeres, scape elongated; forewing venation always reduced (Cenomanian: Myanmar)................................................................................................................... Cretapristocera gen. nov. 2. Head elongated with small eyes.......................................................................................................................3- Head almost globose with large eyes...............................................................................................................4 3. Mandibles only with few teeth (five) along masticatory margin; legs with pro- and meta-femora clearly thickened; forewing cell 2 R 1 open (Cenomanian: Myanmar)........................... Bethylitella Cockerell, 1917a - Mandibles with small teeth along masticatory margin and with basal one elongated: legs, except pro- femora, only slightly thickened; forewing cell 2 R 1 almost closed (Eocene: Baltic amber).............................................................................................................................................................. Protopristocera Brues, 1923 4. Notauli well developed, parapsidal furrow weak (Eocene: Oise amber).... Gynopteron Falières & Nel, 2019 - Notauli weak, parapsidal furrow well-visible (Eocene: Baltic amber)................. Bethylopteron Brues, 1933, Published as part of Jouault, Corentin, Ngô-Muller, Valérie, Pouillon, Jean-Marc & Nel, André, 2021, New Burmese amber fossils clarify the evolution of bethylid wasps (Hymenoptera: Chrysidoidea), pp. 1044-1058 in Zoological Journal of the Linnean Society 191 (4) on page 1050, DOI: 10.1093/zoolinnean/zlaa078, http://zenodo.org/record/4681287, {"references":["Cockerell TDA. 1917 a. Arthropods in Burmese amber. American Journal of Science ser. 4 44: 360 - 368.","Brues CT. 1923. Some new fossil parasitic Hymenoptera from Baltic amber. Proceedings of the American Academy of Arts and Sciences 58: 327 - 346.","Falieres E, Nel A. 2019. The first representative of the extinct flat wasp subfamily Protopristocerinae (Insecta: Hymenoptera, Bethylidae). Palaeoentomology 2: 322 - 326.","Brues CT. 1933. The parasitic Hymenoptera of the Baltic amber. Part 1. Bernstein-Forschungen (Amber Studies) 3: 4 - 179."]}

Clypeus with apical margin modified and medially emarginate; first metasomal tergum with transverse carina sometimes present [Early Cretaceous (Barremian): Lebanon]...........................................................3 2. Anterior wing margin beyond pterostigma swollen (margin discontinuous); second cubital cell open; pretarsal claws simple (Cenomanian: Myanmar).............................................. Holopsenelliscus Engel, 2019 - Anterior wing margin beyond pterostigma not swollen (margin continuous); second cubital cell closed; pretarsal claws dentate (Santonian: Taimyr amber, Russia).................................. Cretabythus Evans, 1973 3. Antennae with ten flagellomeres; forewings with cell 2 R 1 short, ending far from wing apex; veins Rs and 2r-rs curved; propodeum with propodeal spines (Cenomanian: Myanmar)................... Holopsenella Engel et al., 2016 - Antennae with 11 flagellomeres; forewings with cell 2 R 1 elongate, nearly reaching wing apex; veins Rs and 2r-rs straight; propodeum without propodeal spines (Cenomanian: Myanmar)...... Megalopsenella gen. nov.

CRETAPRISTOCERA GEN. NOV. u r n: l s i d: z o o b a n k. o r g: a c t: C 8 9 E 2 D 1 0 - E 2 8 F 4298-AC06-B20BCA318394. Type species: Cretapristocera longiscapa. sp. nov. Etymology: The generic name is composed of the Latin creta, chalk, used here in reference to the Cretaceous period and the genus Pristocera Klug, 1 8 0 8, Pristocerinae, which it resembles. Gender feminine. Diagnosis: Female fully winged; only cells 2R1 and R present; pterostigma narrow, elongate; antennae with ten flagellomeres; scape long; palpal formula 6-3; propodeum rectangular and not spinose; with stout femora., Published as part of Jouault, Corentin, Ngô-Muller, Valérie, Pouillon, Jean-Marc & Nel, André, 2021, New Burmese amber fossils clarify the evolution of bethylid wasps (Hymenoptera: Chrysidoidea), pp. 1044-1058 in Zoological Journal of the Linnean Society 191 (4) on page 1047, DOI: 10.1093/zoolinnean/zlaa078, http://zenodo.org/record/4681287