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Monatractides persicus Pe��ić, 2004 Monatractides stadleri ��� Asadi et al. 2010: 46 Material examined ��� Iran, Mazandaran Province, IR13 area of Ramsar city, Elburs Mountain, small waterfall, 36.8897 N, 50.5681 E, 29.viii.2017 leg. Pe��ić, Zawal & Saboori, 2♀ (sequenced [CCDB38233 F07, -F08]). Remarks��� Monatractides persicus Pe��ić, 2004 was described from a stream in Elburs mountain of North Iran (Pe��ić & Saboori 2004). Later on, this species was placed in synonymy with M. stadleri (Walter, 1924) by Asadi et al. (2010). The latter authors felt that some differences such as the presence of knob-shaped protrusions at the margin of the gnathosomal bay are not suitable for separating the two species. The specimen from Northern Iran used in this study for molecular analysis perfectly matches the description of M. persicus. The latter species can be distinguished from M. stadleri on the basis of a relatively thicker chelicera (ratio L/H 5.6���5.9 vs. 6.2���6.4, data taken from Pe��ić & Saboori 2004). Based on our COI tree, Monatractides persicus is a sister taxon of M. stadleri (see Fig. 6). The average K2P genetic distance between M. persicus and M. stadleri was estimated to 9.35%. Thus, the morphological and molecular data suggest that M. persicus is a valid species distinct from M. stadleri. Distribution��� Iran., Published as part of Pe��i��, Vladimir, Zawal, Andrzej, Saboori, Alireza & Smit, Harry, 2021, New records of water mites (Acari, Hydrachnidia) from Iran with the description of one new species based on morphology and DNA barcodes, pp. 425-440 in Zootaxa 5082 (5) on pages 434-435, DOI: 10.11646/zootaxa.5082.5.2, http://zenodo.org/record/5794932, {"references":["Pesic, V. & Saboori, A. (2004) Water mite species of the genus Monatractides K. Viets (Acari, Hydrachnidia, Torrenticolidae) from Iran, with the description of two new species. Zootaxa, 673, 1 - 10.","Asadi, M., Pesic, V. & Etemadi, I. (2010) A revised survey of water mites (Acari: Hydrachnidia) from Iran: new synonyms and descriptions of three new species. Zootaxa, 2628, 43 - 55."]}

Hydrodroma golestanica sp. nov. Figs. 1-2 Material examined��� Holotype ♂ (RMNH), Iran, Golestan Province, IR3 Shirabad near Khanbebin, stream, downstream near bridge, 36.967499 N, 55.027779 E, 27.viii.2017 leg. Pe��ić, Zawal & Saboori, dissected and slidemounted. Paratypes: 2♂, 1♀, same place and data as holotype (RMNH); 1♂ sequenced [CCDB38361 F03]; 1♀ dissected and slide-mounted (RMNH). Diagnosis��� Integument papillae rounded. Genital plates with 40���51 pairs of Ac in at most four longitudinal rows. Numbers of leg swimming setae: II-L-posterior 1; III-L-4 posterior 6-7; III-L-5 posterior 5; IV-L-4 anterior 6-8, posterior 6-7; IV-L-5 anterior 2-3, posterior 4-5. Description. Both sexes: Integument papillae rounded (Fig. 1C). Cx-I+II medially separated by a fine membranous line, with a row of long fine setae at medial margins of Cx-I, and posterior margins of Cx-I, Cx-II, -III, and ���IV. Genital flaps with rounded lateral and concave medial margins (Figs. 1 A-B). Excretory pore sclerotized. Leg claws with a dorsal clawlet. Male (holotype; in parentheses some measurements of the paratype specimen [CCDB 38361 F03]): Idiosoma L 1130, W 1090. Coxal field L 527; Cx-III W 559, L Cx-I+II 266, Cx-III+IV 278; coxal setae numbers: Cx-I, 24 (23); Cx-II, 23 (22); Cx-III, 14 (18); Cx-IV 11 (19). Genital plate (Fig. 1A) L 259 (238), on each plate Ac number 49-51 (49), with 36-40 setae, all hollow. Ejaculatory complex L 198. Gnathosoma vL 228; chelicera (Fig. 2D) total L 297, L basal segment 238, claw 63, L ratio basal segment/claw 3.8. Palp (Figs. 2 A-B) total L 444, dL/H, dL/H ratio: P-1, 55/45, 1.23; P-2, 73/56, 1.3; P-3, 50/53, 0.94; P-4, 186/47, 4.0; P-5, 80/22, 3.6; L ratio P-2/P-4, 0.4. dL IV-L: 128, 150, 200, 309, 313, 291. Female (paratype, n = 1; juvenile specimen ([CCDB 38361 F05]): Idiosoma L 790. Coxal field L 486; Cx-III W 494, L Cx-I+II 256, Cx-III+IV 281; coxal setae numbers: Cx-I, 18; Cx-II, 18; Cx-III, 13; Cx-IV 12. Genital plate large, L 210-217, on each plate Ac number 40-42, with 30 setae. Gnathosoma vL 228; chelicera total L 297, L basal segment 233, claw 63, L ratio basal segment/claw 3.7. Palp total L 446, dL/H, dL/H ratio: P-1, 53/45, 1.19; P-2, 77/57, 1.34; P-3, 52/55, 0.94; P-4, 189/42, 4.5; P-5, 75/22, 3.4; L ratio P-2/P-4, 0.41. dL IV-L: 125, 150, 205, 309, 328, 272. Species delimitation using DNA-barcodes ��� The final alignment for species delimitation using COI sequence data of ten Hydrodroma specimens listed in Table 2 and one outgroup, Protzia cabardinica (BOLD: 8. IR8 2017 H2) from Iran to root the tree. The NJ tree is presented in Fig. 3. The COI tree sequence retrieved from a specimen of H. golestanica sp. nov. from Iran appeared as a sister taxa of H. torrenticola (Fig. 3). The average K2P genetic distance between the COI sequence of one specimen of H. golestanica sp. nov. from Iran and a specimen of H. torrenticola of from Montenegro was estimated to 10.1% (Table 3). Etymology��� Named after the province where the new species was collected. Discussion ���The new species from Golestan Province closely resembles Hydrodroma torrenticola, a species widely distributed in the Palaearctic (Di Sabatino et al. 2010). It shares the presence of only one short swimming seta on II-L-5 and 2-3 swimming setae on anterior IV-L-5. Only one specimen of the new species could be acquired for use in molecular analyses, so intraspecific differences in COI sequence could not be examined. The phylogenetic analysis based on COI data reveals that the new species is most similar to H. torrenticola. Level of COI differentiation between the new species from Iran and H. torrenticola from Montenegro was 10.1% K2P divergence. From a morphological point of view, H. torrenticola can be separated from the new species by the shape of the integument papillae (pointed vs. distally rounded in H. golestanica sp. nov.). Distribution��� Iran., Published as part of Pe��i��, Vladimir, Zawal, Andrzej, Saboori, Alireza & Smit, Harry, 2021, New records of water mites (Acari, Hydrachnidia) from Iran with the description of one new species based on morphology and DNA barcodes, pp. 425-440 in Zootaxa 5082 (5) on pages 427-430, DOI: 10.11646/zootaxa.5082.5.2, http://zenodo.org/record/5794932, {"references":["Di Sabatino, A., Gerecke, R., Gledhill, T. & Smit, H. (2010) Acari: Hydrachnidia II. In: Gerecke, R. (Ed.), Chelicerata: Acari II. Susswasserfauna von Mitteleuropa, Vol. 7, 2 - 2, Elsevier Spektrum Akademischer Verlag, Heidelberg, pp. 1 - 234."]}

Sperchon amuzgari Bader & Sepasgosarian, 1979 Material examined ��� Iran, Mazandaran Province, IR7 small stream along road to Kandelous village, 36.426388 N, 51.470001 E, 28.viii.2017 leg. Pe��ić, Zawal & Saboori, 3♀ (sequenced [CCDB38233 F09, -F11, -G01]). Remarks ��� Sperchon amuzgari was described by Bader & Sepasgosarian (1979) from a karstic spring in Lorestan Province, Iran. Later on, Asadi et al. (2010) re-examined the holotype of S. amuzgari and found that in view of the similar shape of the palp and dorsum, III/IV-L-3���5 with numerous pinnate dorsal setae and sclerotized excretory pore the later species matches the morphology of S. hispidus Koenike, 1895, a species widely distributed in the Western Palaearctic (Di Sabatino et al. 2010). As a result, Asadi et al. (2010) proposed to place S. amuzgari in synonymy with S. hispidus. During our 2017 survey in Northern Iran we collected and successfully barcoded three specimens of hispidus -like mites that match the original description of S. amuzgari. According to the original description, S. amuzgari can be separate from S. hispidus by having a parallelogram-shaped Cx-IV (Bader & Sepasgosarian 1979). The final alignment for species delimitation using COI sequence data comprised sequences of 34 Sperchon specimens listed in Table 4 and one outgroup, Sperchonopsis verrucosa (Protz, 1896) [BOLD 46. M19_16B_1_ G10] from Montenegro to root the tree. The neighbor-joining (NJ) tree is presented in Fig. 4. The COI tree sequence recovered S. amuzgari as a sister branch to the clade grouping COI sequences found in S. hispidus (Fig. 4). The average genetic distance between the COI sequence of the three specimens of S. amuzgari from Iran and two specimens of S. hispidus from Montenegro was estimated to 16.5% K2P indicating a long independent history of these two species. Therefore, we propose to resurrect Sperchon amuzgari Bader & Sepasgosarian, 1979 as a valid species distinct from S. hispidus. Distribution��� Iran., Published as part of Pe��i��, Vladimir, Zawal, Andrzej, Saboori, Alireza & Smit, Harry, 2021, New records of water mites (Acari, Hydrachnidia) from Iran with the description of one new species based on morphology and DNA barcodes, pp. 425-440 in Zootaxa 5082 (5) on page 431, DOI: 10.11646/zootaxa.5082.5.2, http://zenodo.org/record/5794932, {"references":["Bader, C. & Sepasgosarian, H. (1979) Wassermilben (Acari, Prostigmata, Hydrachnellae) aus dem Iran. 10. Mitteilung. Entomologische Mitteilungen aus dem Zoologischen Museum Hamburg, 6 (105), 243 - 252.","Asadi, M., Pesic, V. & Etemadi, I. (2010) A revised survey of water mites (Acari: Hydrachnidia) from Iran: new synonyms and descriptions of three new species. Zootaxa, 2628, 43 - 55.","Di Sabatino, A., Gerecke, R., Gledhill, T. & Smit, H. (2010) Acari: Hydrachnidia II. In: Gerecke, R. (Ed.), Chelicerata: Acari II. Susswasserfauna von Mitteleuropa, Vol. 7, 2 - 2, Elsevier Spektrum Akademischer Verlag, Heidelberg, pp. 1 - 234."]}

Genus Docleiana Delicado & Pe��ić gen. nov. urn:lsid:zoobank.org:act: 04639DB3-91D2-46B7-BFAB-FFC1EA3C27BD Type species Arganiella tabanensis Boeters, Gl��er & Pe��ić, 2014. Revised diagnosis Shell valvatiform; whorls ca 3; aperture complete, from rounded to ellipsoidal; outer lip narrow, straight in lateral profile; umbilicus wide. Operculum corneous, yellowish, thin, pliable, rounded, paucispiral with a central nucleus, without peg. Two pairs of basal cusps on each central radular tooth. Ctenidium well developed, with 10���11 gill filaments. Animal unpigmented. Osphradium positioned opposite approximately to the middle of the ctenidium. Stomach without gastric caecum; rectum forms a gentle V-shape in the mantle cavity. Bursa copulatrix small, globular, pedunculated and positioned beyond the posterior edge of the albumen gland; bursal duct longer than bursal length; unpigmented renal oviduct; one pyriform seminal receptacle arising at the insertion point with the bursal duct loop. Penis small and simple, gradually tapering. Etymology The new genus is named after Doclea, the name of the Roman city located on the site of modern Podgorica on whose municipal territory a new genus was found; gender feminine. Remarks Docleiana Delicado & Pe��ić gen. nov. can be distinguished from Arganiella as the former has a more ellipsoidal shell aperture, a narrower base of the penis, a smaller bursa copulatrix that is positioned beyond the posterior edge of the albumen gland, a larger seminal receptacle and more cusps on the lateral radular teeth (Fig. 2; for comparison, see Giusti & Pezzoli 1980; Boeters et al. 2014). The new genus differs from the closely related genus Kerkia by its smaller shell, absence of lobes on the inner edge of the penis, a smaller bursa copulatrix, a more globular seminal receptacle and V-shaped rectum (see Radoman 1978; Bodon et al. 2001). Mean COI sequence divergence for Docleiana Delicado & Pe��ić gen. nov. was 14.5% with Arganiella and 12.3% with Kerkia., Published as part of Delicado, Diana, Pe��i��, Vladimir & Ramos, Marian A., 2021, Arganiella Giusti & Pezzoli, 1980 (Caenogastropoda: Truncatelloidea: Hydrobiidae): a widespread genus or several narrow-range endemic genera?, pp. 140-155 in European Journal of Taxonomy 750 (1) on pages 146-147, DOI: 10.5852/ejt.2021.750.1369, http://zenodo.org/record/5451701, {"references":["Boeters H. D., Gloer P. & Pesic V. 2014. Arganiella tabanensis n. sp. from Montenegro (Mollusca: Gastropoda: Hydrobiidae). Ecologica Montenegrina 1 (3): 131 - 139. https: // doi. org / 10.37828 / em. 2014.1.20","Giusti F. & Pezzoli E. 1980. Hydrobioidea nuove o poco conosciute dell Italia appenninica (Gastropoda: Prosobranchia). Archiv fur Molluskenkunde 111: 207 - 222.","Radoman P. 1978. Neue Vertreter der Gruppe Hydrobioidea von der Balkanhalbinsel. Archiv fur Molluskenkunde 109 (1 - 3): 27 - 44.","Bodon M., Manganelli G. & Giusti F. 2001. A survey of the European valvatiform hydrobiid genera with special reference to Hauffenia Pollonera, 1898 (Gastropoda: Hydrobiidae). Malacologia 43: 103 - 215."]}

Genus Aretiana Delicado & Ramos gen. nov. urn:lsid:zoobank.org:act: 7B207B6D-47E8-4211-8139-0CF98DA353D7 Type species Boetersiella wolfi Boeters & Gl��er, 2007. Revised diagnosis Shell trochiform; whorls 3.5���4.0; aperture complete, rounded; outer lip narrow, straight in lateral profile; umbilicus wide. Operculum corneous, yellowish, thin, pliable, oval to rounded, paucispiral with a central nucleus, without peg. Two pairs of basal cusps on each central radular tooth. Ctenidium well developed, with approximately 14 gill filaments. Osphradium positioned opposite to approximately the middle of the ctenidium. Stomach without gastric caecum; rectum forms a gentle U-shape in the mantle cavity. Bursa copulatrix medium-sized, pyriform, pedunculated and protruding beyond the posterior edge of the albumen gland; bursal duct shorter than bursal length; unpigmented renal oviduct that makes a complete loop over the pallial gland; one elongated or pyriform seminal receptacle arising from the renal oviduct, just above the insertion point with the bursal duct. Prostate gland bean-shaped, about twice as long as wide. Penis small and simple, gradually tapering. Nervous system unpigmented. Etymology The genus is named after Villa Aretiana, the Roman name of the town Aracena, which gives its name to the mountain range where the genus was found (i.e., Sierra de Aracena); gender feminine. Remarks Aretiana Delicado & Ramos gen. nov. can be distinguished from Arganiella as the former has a taller shell, a more oval operculum, pigmentation on the body and eyespots, fewer gill filaments, a narrower penis, a larger and pyriform bursa copulatrix located beyond the posterior edge of the albumen gland and fewer cusps on the lateral radular teeth (for comparison, see Giusti & Pezzoli 1980; Boeters & Gl��er 2007; Arconada & Ramos 2007a). The new genus differs from the closely related genus Iberhoratia by its taller shell with a narrower umbilicus, the absence of lobes on the inner edge of the penis and of a proximal seminal receptacle (SR2) and presence of two pairs of basal cusps on each central radular tooth (see Arconada et al. 2007). Mean COI divergence for Aretiana Delicado & Ramos gen. nov. was 15.9% with Arganiella and 11.2% with Iberhoratia., Published as part of Delicado, Diana, Pe��i��, Vladimir & Ramos, Marian A., 2021, Arganiella Giusti & Pezzoli, 1980 (Caenogastropoda: Truncatelloidea: Hydrobiidae): a widespread genus or several narrow-range endemic genera?, pp. 140-155 in European Journal of Taxonomy 750 (1) on pages 145-146, DOI: 10.5852/ejt.2021.750.1369, http://zenodo.org/record/5451701, {"references":["Boeters H. D. & Gloer P. 2007. A contribution to the genus Boetersiella Arconada & Ramos 2001 in Spain with the description of Boetersiella wolfi n. sp. Heldia 5: 85 - 88.","Giusti F. & Pezzoli E. 1980. Hydrobioidea nuove o poco conosciute dell Italia appenninica (Gastropoda: Prosobranchia). Archiv fur Molluskenkunde 111: 207 - 222.","Boeters H. D., Gloer P. & Pesic V. 2014. Arganiella tabanensis n. sp. from Montenegro (Mollusca: Gastropoda: Hydrobiidae). Ecologica Montenegrina 1 (3): 131 - 139. https: // doi. org / 10.37828 / em. 2014.1.20","Arconada B. & Ramos M. A. 2007 a. Description of a new species of the genus Arganiella Giusti & Pezzoli, 1980 (Mollusca, Gastropoda, Hydrobiidae) from the Iberian Peninsula. Graellsia 63 (1): 61 - 70. https: // doi. org / 10.3989 / graellsia. 2007. v 63. i 1.81","Bodon M., Manganelli G. & Giusti F. 2001. A survey of the European valvatiform hydrobiid genera with special reference to Hauffenia Pollonera, 1898 (Gastropoda: Hydrobiidae). Malacologia 43: 103 - 215."]}

Atractides (Atractides) khuzestanicus sp. nov. (Fig. 5) Type material. Holotype female (RMNH), dissected and slide mounted, Iran, Khuzestan Province, White river of Sardasht, 30��18'60"N, 50��12'60"E, 29 July 2017, coll. Isa Etemadi. Paratypes (SBUk): one female, river of Bebahan, 30��35'17"N, 50��14'27"E, 16 April 2016, coll. Isa Etemadi, dissected and slide mounted. Diagnosis. Dorsum with one pair of small sclerotized muscle attachment platelets, Vgl-1 fused to Vgl-2; I-L-5 with setae S-1/-2 without interspace; I-L-6 stout, with straight ventral margin; gnathosoma with protruding rostrum; chelicera with a long, weakly curved claw, L ratio basal segment/claw 1.9; P-3 with two strong setae, inserting on medial and lateral surface, respectively. Description. Female. Integument striated; postoc and D-1 fused to a small and longish platelet (Fig. 5A). Cx-I with a relatively long median suture line, apodemes of Cx-II directed caudolaterally. Pregen strong and curved, gonopore relatively long, genital plates on the level of postgen, with the acetabula in triangular arrangement. Excretory pore sclerotized; Vgl-1 fused to Vgl-2 (Fig. 5B). I-L-5 with setae S-1/-2 inserting side by side, without interspace; I-L-6 stout, not curved, dorsal and ventral margin diverging from base to tip, ventral margin straight. P- 2 ventral margin slightly convex, without projection, P-3 with one seta on medial and lateral surface, respectively, P-4 sword seta between the ventral setae (Fig. 5E). Gnathsoma with a protruding rostrum (Fig. 5D). I-L-5 with S-1 and S-2 short, inserted side by side; I-L-6 stout, nearly straight, distally thickened, with strong claws bearing a ventral clawlet (Fig. 5C). Measurements. Idiosoma L 684, W 500. Coxal shield L 314; Cx-III W 378; Cx-I+II mL 122, Cx-I+II lL 231. Genital field L/W 138/152, genital plate L 82���83, gonopore L 116, pregenital sclerite W 91, L Ac-1-3: 22���25, 22��� 27, 31���34. Palp: palp total L 272; dL/H, dL/H ratio: P-1, 31/27, 1.18; P-2, 63/50, 1.26; P-3, 58/36, 1.61; P-4, 83/28 [basal 23], 3.0 [3.56]; P-5, 37/16, 2.35; length ratio P-2/P-4 0.75. Gnathosoma vL 158; chelicera total L 278, L basal segment 184, claw 97, L ratio basal segment/claw 1.9. Legs: I-L-5 dL 113, vL 83, dL/vL ratio 1.36, maximum H 36, dL/maximum H 3.2, S-1 L 41, L/W ratio 8.8, S- 2 length 39, L/W ratio 8.3, distance S-1-2, 3.0, dL ratio S-1/2 1.06; I-L-6 dL 114, central H 30, dL/central H ratio 3.8; length ratio I-L-5/6 0.99. Male. Unknown. Etymology. Named after its occurrence in Khuzestan province of Iran. Remarks. The new species from South Iran belongs to the ��� subasper species group���, characterized by a pointed, protruding gnathosomal rostrum and the chelicera with a long, needle-like claw, L ratio basal segment/ claw et al. 2016). The new species can easily be distinguished from all other species of the ��� subasper species group���, A. octoporus Piersig, 1898, A. subasper Koenike, 1902, A. digitatus Lundblad, 1954, A. heversi K.O.Viets, 1982 and A. martini Pe��ić, Erman & Esen, 2010 (see Gerecke 2003 and Erman et al. 2010), in having two strong setae, inserting on medial and lateral surface of P-3, respectively, and in setae S-1/-2 on I-L-5 not distanced from each other., Published as part of Pe��i��, Vladimir, Asadi, Mahdieh, Etemadi, Isa & Smit, Harry, 2019, New records of water mites (Acari: Hydrachnidia) from the Khuzestan Province (South Iran) with description of three new species, pp. 550-558 in Zootaxa 4559 (3) on pages 552-555, DOI: 10.11646/zootaxa.4559.3.5, http://zenodo.org/record/2627233, {"references":["Gerecke, R., Gledhill, T., Pesic, V. & Smit, H. (2016) Chelicerata: Acari III. In: Gerecke, R. (Ed.), Susswasserfauna von Mitteleuropa, Bd. 7 / 2 - 3, Springer-Verlag Berlin, Heidelberg, pp. 1 - 429. https: // doi. org / 10.1007 / 978 - 3 - 8274 - 2689 - 5","Erman, O., Pesic, V., Esen, Y. & Ozkan, M. (2010) A checklist of the water mites of Turkey (Acari: Hydrachnidia) with description of two new species. Zootaxa, 2624, 1 - 48 https: // doi. org / 10.11646 / zootaxa. 2624.1.1","Gerecke, R. (2003) Water mites of the genus Atractides Koch, 1837 (Acari: Parasitengona: Hygrobatidae) in the western Palaearctic region: a revision. Zoological Journal of the Linnean Society, 138, 141 - 378. https: // doi. org / 10.1046 / j. 1096 - 3642.06 - 0.00051. x"]}

Torrenticola (Megapalpis) asadiae Pe��ić, Smit & Saboori, 2012 Material examined. Iran, Khuzestan Province, Andika-Taluk, 32��17'53''N, 49��19'51''E, 29 September 2016, coll. Isa Etemadi, 2/1/2 (mounted). Distribution. Hormozgan Province���Pe��ić et al. (2012), Khuzestan Province���this study., Published as part of Pe��i��, Vladimir, Asadi, Mahdieh, Etemadi, Isa & Smit, Harry, 2019, New records of water mites (Acari: Hydrachnidia) from the Khuzestan Province (South Iran) with description of three new species, pp. 550-558 in Zootaxa 4559 (3) on page 552, DOI: 10.11646/zootaxa.4559.3.5, http://zenodo.org/record/2627233, {"references":["Pesic, V., Smit, H. & Saboori A. (2012) Water mites delineating the Oriental and Palaearctic regions-the unique fauna of southern Iran, with descriptions of one new genus, one new subgenus and 14 new species (Acari: Hydrachnidia). Zootaxa, 3330, 1 - 67."]}

Litarachna enigmatica Pe��ić, Durucan & Chatterjee sp. nov. (Figs. 1 A���D) Type series. Holotype female, dissected and mounted in Hoyer, Turkey, Antalya, Hamit Bey Plaji, 36.87555556��N, 30.70722222��E, 30.vii.2017 leg. Durucan. Diagnosis (Male unknown): Suture lines Cx-II/III and Cx-III/IV complete; medial posterior apodemes of Cx- IV absent; postgenital sclerite with a pair of wheel-like acetabula; P-2 with a anteroventral tubercle; P-4 ventral margin with a peg-like seta. Description. Female (holotype): Idiosoma elongated L/W 347/241; coxal field L/W 138/188; Cx-I separated medially; suture lines Cx-II/III and Cx-III/IV complete; posterior margin of Cx-IV without medial apodemes, lateral apodemes shortened and broad (Fig. 1A). Genital field L/W 67/34, gonopore elongated, pregenital sclerite small (W 27), postgenital sclerite bowed, with a pair of wheel-like acetabula (Fig. 1A). A pair of small platelets with coxoglandularia 4 and associated setae free in the integument posterior to postero-lateral apodemes of Cx-IV. Posterior to the genital field three pairs of probably wheel-like acetabula sensu Cook (1996); excretory pore unsclerotized. Gnathosoma with a short rostrum, vL 66; chelicera two-segmented, typically claw-like in form, total L 95; palp total L 222, dL/H ratio: P-1, 16/16, 1.0; P-2, 69/34, 2.0; P-3, 38/31, 1.2; P-4, 72/22, 3.3; P-5, 27/13, 2.0; dL P-2/P- 4 ratio 0.96; P-2 with an anteroventral tubercle; P-4 ventral margin with a well developed peg-like seta, dividing the segment into two parts, the proximal one longer than the distal one, both with a concave ventral margin, ventral hair-like setae at the level of peg-like seta insertion (Fig. 1D). Legs: dL of I-L (Fig. 1B): 33, 36, 38, 48, 59, 69; I-L-6 H 18, I-L-6 dL/H ratio 3.9; dL of IV-L (Fig. 1C): 52, 32, 38, 59, 76, 64; IV-L-5 with one short swimming seta. Male: Unknown. Etymology. enigmaticu s���an adjective from Latin origin meaning ���mysterious���; refers to the elusive features of the new species. Remarks. The new species resembles Litarachna gracilis Pe��ić, 2013 due to the absence of medial apodemes of Cx-IV, the presence of a ventral projection on P-2 and the peg-like seta on P-4. The latter species, described from South Korea (Pe��ić 2013a), is a characteristic inhabitant of marine interstitial (as indicated by the absence of eye pigment) and can be distinguished by a number of features: suture lines Cx-II/III and Cx-III/IV incomplete, gonopore more elongated, a different arrangement of wheel-like acetabula with the the anterior pair positioned on the level of postgenital sclerite, and two posterior pairs almost touching each other, and P-4 ventral setae approached to the distoventral edge of the segment (Pe��ić 2013a). Litarachna enigmatica also resembles to L. smiti Pe��ić, Chatterjee & Ahmed Abada, 2008 from the Red Sea (Pe��ić et al. 2008a, b), L. bartschae Smit, 2003 from Australia and Brunei (Smit 2003, Pe��ić et al. 2013b) and L. madagascariensis Pe��ić, 2013 from Madagascar (Pe��ić 2013b) in having a ventral projection on P-2 and the peg- like seta on P-4. However, the new species can be easily distinguished from them by the lack of a setal tubercle on P-4 and completely reduced medial apodemes of Cx-IV. Moreover, the new species is unique within the genus Litarachna in the postgenital sclerite having a pair of wheel-like acetabula, a character found only in the females of Pontarachna. As pointed out by Cook (1996), in males of Pontarachna and both sexes of Litarachna no wheel-like acetabula are present associated with the genital sclerites. Anyhow it is worth to mention that the structure of the so-called wheel-like acetabula associated with the postgenital sclerite should be determined under scanning electron microscope (SEM). Unfortunately, with the single specimen of our study this was not possible. Distribution. Turkey; known only from the type locality., Published as part of Durucan, Furkan, Chatterjee, Tapas & Pe��i��, Vladimir, 2018, Two new species of the marine water mite family Pontarachnidae (Acari: Hydrachnidia) from the Gulf of Antalya, Turkey, pp. 271-278 in Zootaxa 4531 (2) on pages 272-273, DOI: 10.11646/zootaxa.4531.2.8, http://zenodo.org/record/2614636, {"references":["Cook, D. R. (1996) A freshwater species of Pontarachna (Acari, Pontarachnidae) from South Africa, with a discussion of genital acetabula in the family. Anales Instituto de Biologia, Universidad Nacional Autonoma de Mexico, Seria Zoologia, 67, 259 - 264.","Pesic, V., Chatterjee, T. & Ahmed Abada, A. E. (2008 a) A new species of marine water mite (Acari: Hydrachnidia: Pontarachnidae) from the Red Sea. Systematic and Applied Acarology, 13, 133 - 136. https: // doi. org / 10.11158 / saa. 13.2.6","Pesic, V, Chatterjee, T. & Ahmed Abada, A. E. (2008 b) Marine water mites (Acari: Hydrachnidia: Pontarachnidae) from the Red Sea, with description of one new species. Cahiers de Biologie Marine, 49, 375 - 379.","Smit, H. (2003) Five new species of the water mite family Pontarachnidae from Western Australia (Acari: Hydrachnidia). In: Wells, F. E., Walker, D. I. & Jones, D. S. (Eds.), The Marine Flora and Fauna of Dampier, Western Australia. Western Australian Museum, Perth, pp. 547 - 562."]}

Sterkspruitia comorosensis Pe��ić & Smit, 2015 (Fig. 5) Material examined. Comoros, Mayotte: River Banano, upstream, 12��54'1,2"S, 45��11'9, 8"E, alt. 50 m asl., 21-vii- 2015 0/1/0 (dissected and slide mounted, MNHN). Morphology. Female���Idiosoma L/W 619/467; dorsal shield L/W 469/366, consisting of four pairs of platelets, each with one glandularium (Fig. 5A); suture lines of coxal plates visible, dividing coxae in 3 groups, separated by strips of porose sclerotization, tips of Cx-I extending beyond frontal idiosoma margin; gnathosoma broadly fused with Cx-I (Fig. 5B), L gnathosoma + Cx-I mL 328, Cx-III W 412; genital field with three pairs of Ac, altogether fused to medial margin of a plate bearing a pair of glandularia at posterior margin, anterior to gonopore a large, transverse plate separated from ventral shield and other surrounding sclerites by narrow strips of soft integument, flanked by a pair of small plates, each with a glandularium; excretory pore on a small, subtriangular platelet. Palp: total L 348, dL/H, dL/H ratio: P-1, 25/34, 0.73; P-2, 81/40, 2.0; P-3, 111/39, 2.8; P-4, 92/31, 3.0; P-5, 39/ 16, 2.5; P-2/P-4 ratio 0.87; P-2 with a spatula-shaped ventrodistal protrusion, P-3 with a central protrusion; P-4 with ventral setae and a short, fine sword seta, inserted far proximally (Fig. 5D). Legs: I-L-5 strongly thickened, with ventral and dorsal margins diverging from base to S-1 insertion, distal part abruptly narrowed, I-L-5 dL 204, vL 86, dL/vL ratio 2.37, maximum H 86, dL/maximum H ratio 2.37, S-1 and -2 heteromorphic and distanced, S-1 ending in a fine pointed tip, S-2 with well developed dorsal projection, apically rounded, S-1 L 92, L/W ratio 4.9, S-2 L 50, L/W ratio 2.67, distance, L ratio S-1/2 1.84; I-L-6 long and slender, only slightly narrowed from the base to the tip, I-L-6 L 186, central H 18, L/central H ratio 10.3; L I-L-5/6 ratio 1.1; dL of IV-L: 84, 95, 141, 167, 200, 182; as in male IV-L-3-6 with strongly serrated setae (see Pe��ić et al. 2015a, fig. 9D). Remarks. This species was described by Pe��ić et al. (2015a) based on a single male specimen from Koual�� River. No further records were published, this is the first description of the female. Distribution. Comoros: Mayotte., Published as part of Pe��i��, Vladimir, Smit, Harry & Mary, Nathalie, 2018, Fifth contribution to the knowledge of water mites (Acari: Hydrachnidia) from the Comoros: a checklist and description of one new genus and four new species, pp. 331-348 in Zootaxa 4483 (2) on page 339, DOI: 10.11646/zootaxa.4483.2.6, http://zenodo.org/record/1437784

Platymamersopsis mayottensis Pe��ić & Smit sp. nov. (Fig. 2, 11A���B) Type series. Holotype male (MNHN), dissected and slide mounted, Comoros, Mayotte, Koual�� River, intermediate, 12��47'50,2375"S, 45��11'08,8249"E, alt. 50 m asl., 28-vii-2016, leg. Mary. Diagnosis (Female unknown): Dorsal shield elongated, ridge on dorsal shield apically tapering; P-4 with well developed ventral projection, curved, directed anteriorly; I-III-L claws with comb-like ventral clawlets; IV-L-6 slender, with two small terminal setae and a row of three stronger sub-terminal setae. Description��� Idiosoma oval; dorsal shield elongated, L/W 503/350, ratio 1.44, anteriorly rounded, ridge on dorsal shield apically bluntly pointed, caudally dissolved, glandularia far laterally (Fig. 2A); ventral shield (Fig. 2B) oval, L/W 541/438; coxal field L/W 297/356, anteriorly extending beyond frontal margin, medial margin of Cx-I long (mL 70), medial margin of Cx-III mid-sized (mL 36), mL Cx-I/Cx-III ratio 1.9; Cxgl-2 close to posterior margin of Cx-II, Cxgl-4 at anterior margin of Cx-III; Cx-IV posteriorly and laterally rounded, dorso-laterally extended to the level of Cx-II. Genital field L/W 103/100, rounded, laterally smoothly convex, posteriorly slightly extended beyond genital bay; acetabula mid-sized, L Ac-1-3: 28, 28, 31; a few small setae present on genital flaps; setae of Vgl-1 posterior to caudal corner of Cx-IV, Vgl-3 close to caudo-lateral margin of Cx-IV; excretory pore and Vgl-2 close together, distanced from the posterior margin of ventral shield. Palp: total L 214, dL/H, dL/H ratio: P-1, 19/43, 0.44; P-2, 83/57, 1.45; P-3, 34/42, 0.82; P-4, 55/33, 1.67; P-5, 23/14, 1.66; P-2/P-4 ratio 1.5; P-2 compact, dorsal margin curved, ventral margin slightly concave, ventral seta mid-sized, pointed; P-4 ventral projection well developed, curved, flanked by a fine subterminal seta (Fig. 2E); P-5 with a strong distal claw flanked by two shorter claws. Legs: Due to the fixation all claws retracted in claw sheath, but comb-like clawlets can be observed; dL of I-L: 52, 63, 56, 81, 86, 106; dL of IV-L-4-6: 83, 98, 105; IV-L-6 slender, with two small terminal setae and a row of three stronger sub-terminal setae (Fig. 2C). Female: Unknown. Etymology. Named for its presence on Mayotte. Discussion. The new species belongs to the group of species bearing comb-like ventral clawlets (mysorensis - like species, see Goldschmidt 2008 for discussion). The new species from Mayotte differs from all other species of this group by the characteristic ventral projection at P-4. The only species with a similarly shaped P-4, P. nordenskioeldi K. Viets, 1914 from Cameroon and Liberia (see Cook 1966), differs in simple dorsal and ventral clawlets on I���III-L, a large, rounded dorsal shield, a very short coxal field, not protruding over the frontal margin, and a more concave ventral margin of P-2 (Cook 1966). Distribution. Comoros; only known from the type locality., Published as part of Pe��i��, Vladimir, Smit, Harry & Mary, Nathalie, 2018, Fifth contribution to the knowledge of water mites (Acari: Hydrachnidia) from the Comoros: a checklist and description of one new genus and four new species, pp. 331-348 in Zootaxa 4483 (2) on page 334, DOI: 10.11646/zootaxa.4483.2.6, http://zenodo.org/record/1437784, {"references":["Goldschmidt, T. (2008) Taxonomical, ecological and zoogeographical studies on anisitsiellid water mites (Acari: Hydrachnidia: Anisitsiellidae Koenike, 1910) from Madagascar. Zootaxa, 1954, 1 - 120.","Cook, D. R. (1966) The water mites of Liberia. Memoirs of the American Entomological Institute, 6, 1 - 418."]}

Lagoenaxonopsalbia Pe��ić & Smit gen. nov. Diagnosis. Characters of Axonopsalbia -like mites as given by Cook (1974): Dorsal and ventral shields fused posteriorly; lateroglandularia fused with dorsal shield; two pairs of glandular platelets located near the anterior margin of ventral shield, no glandularia in the dorsal furrow; tips of Cx-I not extending to frontal idiosoma margin; Cx-IV with two pairs of glandularia, one located immediately medial, the other posterior to IV-L-insertions; projections associated with openings of IV-L-insertions relatively long and bluntly pointed; three pairs of genital acetabula. Male: Posterior end of dorsal shield modified as illustrated in Fig. 6A, forming a well developed axeshaped cauda; excretory pore on a separate plate lying on the caudal portion of the dorsal shield, excretory pore plate more elevated in the centre, almost circular in lateral view, with paired groups of long setae flanking the excretory pore and paired glandularia associated with humps. Female: Excretory pore lying at extreme end of dorsal shield (Fig. 7A); acetabula-bearing plates and postgenital sclerite fused with the dorsal shield; a pair of glandularia near the anterior margins of the acetabula-bearing genital plates fused to the ventral shield (Fig. 7B). Type species: Lagoenaxonopsalbia comorosensis sp. nov. Remarks. The combination of a well developed flask-shaped idiosoma with a large, widened cauda, the genital field with the gonopore located in terminal position on the cauda, flanked by three pairs of acetabula, and the excretory pore plate elevated in the centre and almost circular in lateral view, lying on the caudal portion of the dorsal shield, will easily distinguish the male of the new genus. The combination of relatively long projections associated with the openings of IV-L-insertions, the acetabulabearing plates fused with the postgenital sclerite and the presence of a pair of glandularia medially to the insertions of IV-L, makes the female of the new species most similar to Axonopsalbia K. Viets, 1913 and Pseudaxonopsalbia (Motasaxona) Cook, 1974. The females of the latter two taxa can be distinguished from the new species in: 1) dorsal furrow with three pairs of small glandularia platelets, all restricted to the posterior half, and not fused to dorsal shield, 2) two pairs of glandularia typically fused to anterior margin of ventral shield, 3) genital field with 4 pairs of acetabula, and 4) the glandularia located near the anterior margins of the acetabula-bearing genital plates free, not fused with the ventral shield. The latter character state is also found in females of Villaxonopsalbia sudafricana Cook, 1999, representative of a monotypic genus known from South Africa. Villaxonopsalbia females are unique among Axonopsalbia -like mites in the fusion of the excretory pore with the combined acetabular plates and postgenital sclerite (Cook 1999). Etymology. The new genus name is a combination of lagoena (Latin, = bottle, flask) and Axonopsalbia., Published as part of Pe��i��, Vladimir, Smit, Harry & Mary, Nathalie, 2018, Fifth contribution to the knowledge of water mites (Acari: Hydrachnidia) from the Comoros: a checklist and description of one new genus and four new species, pp. 331-348 in Zootaxa 4483 (2) on pages 339-340, DOI: 10.11646/zootaxa.4483.2.6, http://zenodo.org/record/1437784, {"references":["Cook, D. R. (1974) Water mite genera and subgenera. Memoirs of the American Entomological Institute, 21, 1 - 860.","Cook, D. R. (1999) Axonopsalbia - like water mites from South Africa (Acari, Hydrachnida, Aturidae). International Journal of Acarology, 25 (4), 317 - 327. https: // doi. org / 10.1080 / 01647959908684170"]}

Djeboa davecooki Pe��ić & Smit sp. nov. (Figs. 9���10, 11E���F) Type series: Holotype male (MNHN), dissected and slide mounted, Comoros, Mayotte, Demb��ni River, upstream, 12��51'38,7"S, 43��09'30,8"E, alt. 156 m asl., 18-vii-2015, leg. Mary. Paratypes: same data as holotype, 31-vii-2014 1/0/0 (RMNH); Hajangoua stream, 12��52'1,0"S, 45��10'25,7"E, alt. 265 m asl., 18-vii-2015 0/1/0 (dissected and slide mounted, RMNH); River Djalimou, upstream, 12��56'58,4"S, 45��07'23,7"E, alt. 165 m asl., 16-vii-2014 1/0/0 (MNHN). Diagnosis: Lateral ridges of central depression not extending to frontal and caudal margin of dorsal shield; gnathosomal bay Y-shaped, in posterior part widened; P-4 L/H 3.1, ventral margin almost straight; I-L-6 L/H 2.8- 2.9. Description. General features ���Dorsal shield oval (L/W ratio 1.15���1.16), with medial depression; muscle scars anterior and posterior to the postocularia, with moderately pronounced thickenings, second pair located well within the concavity (Figs. 9A, 11E); gnathosomal bay Y-shaped, in posterior part widened (Fig. 11F); tips of Cx-I extending to frontal margin; median margin of Cx-IV comparatively well developed. Palp: P-1 with a dorsal seta; P-2 with straight ventral and convex dorsal margin; P-3 ventral margin with a basal kink, distally straight; P-4 relatively slender, ventral margin almost straight (Figs. 9D, 10B). Legs: I-L (Fig. 9E) with I-L-6 L/H ratio 2.8���2.9, ventral margin distally convex; IV-L: Fig. 10C; swimming setae numbers: III-L-4, 4; III-L-5, 4; IV-L-4, 2; IV-L-5, 4. Measurements: Male (holotype)���Idiosoma (ventral view: Fig. 9B) L/W 463/442. Dorsal shield (Figs. 9A) L/ W 392/340, ratio 1.15; gnathosomal bay L 91; distance between IV-L insertions 288. Gonopore long and narrow, L 69; ejaculatory complex L 109. Palp: total L, dL/H, dL/H ratio: P-1, 23/22, 1.04; P-2, 52/40, 1.29; P-3, 52/23, 2.21; P-4, 64/20, 3.14; P-5, 17/10, 1.69; P-2/P-4 ratio 0.81. Legs: dL of I-L: 38, 46, 45, 53, 72, 89; I-L-6 H 31, I-L-6 L/H ratio 2.85; dL of IV-L-4-6: 86, 92, 81. Female (paratype)���Idiosoma (ventral view: Fig. 10A) L/W 463/428. Dorsal shield L/W 406/350, ratio 1.16; gnathosomal bay L 91; distance between IV-L insertions 275. Gonopore L/W 81/72. Palp: total L, dL/H, dL/H ratio: P-1, 22/22, 1.0; P-2, 53/42, 1.26; P-3, 53/25, 2.1; P-4, 63/20, 3.1; P-5, 16/10, 1.64; P-2/P-4 ratio 0.85. Legs: dL of I-L: 42, 47, 45, 52, 69, 83; I-L-6 H 30, I-L-6 L/H ratio 2.8; dL of IV-L: 66, 66, 64, 80, 105, 118. Etymology: The new species is named after David Cook (Paradise Valley) in appreciation of his studies of water mites. Discussion: The combination of an Y-shaped gnathosomal bay, a relatively long and slender P-3 of nearly same height along the entire length (except for the proximoventral bulge), and the large dorsal shield concavity with the lateral ridges not extending to frontal and caudal margin, makes the new species most similar to D. granatica Cook, Pe��ić & Gerecke, 2013 from Madagascar (Pe��ić et al. 2013) and D. dinosaurophila Pe��ić, Cook & Gerecke, 2013 from South Africa (Pe��ić et al. 2013). Djeboa granatica, known only from a male, can be distinguished from the new species by a comparatively slender P-4, with a concave ventral margin (see fig. 24E in Pe��ić et al. 2013) and Cx-III/IV with longitudinal striation. Djeboa dinosaurophila differs in having a gnathosomal bay comparatively narrower proximally, a proportionally thicker distal segment of I-L and in the dorsal shield medial depression rounded in its posterior half (Pe��ić et al. 2013). Distribution: Comoros; known only from the loci typici., Published as part of Pe��i��, Vladimir, Smit, Harry & Mary, Nathalie, 2018, Fifth contribution to the knowledge of water mites (Acari: Hydrachnidia) from the Comoros: a checklist and description of one new genus and four new species, pp. 331-348 in Zootaxa 4483 (2) on page 345, DOI: 10.11646/zootaxa.4483.2.6, http://zenodo.org/record/1437784, {"references":["Pesic, V., Cook, D., Gerecke, R. & Smit H. (2013) The water mite family Mideopsidae (Acari: Hydrachnidia): a contribution to the diversity in the Afrotropical region and taxonomic changes above species level. Zootaxa, 3720 (1), 1 - 75. https: // doi. org / 10.11646 / zootaxa. 3720.1.1"]}

Torrenticola mayottensis Pe��ić & Smit n. sp. (Figs. 3���4, 11C���D) Synonymy. Torrenticola sp. Smit et al. (2010, p. 52, figs. 2A���D). Type series. Holotype male (MNHN), dissected and slide mounted, Comoros, Mayotte, Demb��ni River, downstream, 12��51'38,7"S, 43��09'30,8"E, alt. 10 m asl., 31-vii-2014, leg. Mary. Paratypes (MNHN): same site and data as holotype, 1/0/0; same site, 18-vii-2015 4[3 juvenile]/6[2 juvenile]/0 (0/2/0 dissected and slide mounted) (RMNH). Other material. Comoros, Mayotte: River Bouyouni, intermediate, 12��44'55,6"S, 45��08'36,4"E, alt. 40 m asl., 5-viii-2014 0/1/0 (mounted); Mapouhera upstream, 12��43'4,3"S, 45��05'56,6"E, alt. 110 m asl., 24-vii-2015 0/1/0; River Banano, upstream, 12��54'1,2"S, 45��11'9, 8"E, alt. 50 m asl., 21-vii-2015 3[2 juvenile]/0/0 (1/0/0 mounted); River Dapani, upstream, 12��57'42,4"S, 45��09'22,7"E, alt. 37 m asl., 20-vii-2015 0/1[juvenile]/0; River Djalimou, upstream, 12��56'58,4"S, 45��07'23,7"E, alt. 165 m asl., 16-vii-2015 0/1/0; River Djalimou, downstream, 12��57'14,0"S, 45��06'53,2"E, alt. 30 m asl., 30-vii-2015 1/0/0. Diagnosis. Both sexes: Idiosoma elongated-oval (dorsal shield L/W ratio 1.3���1.4); gnathosomal rostrum long; P-4 stout, L/H 2.4���3.0; female: genital field roundish, L/W 0.95���1.05. Description. General features ���Idiosoma elongated-oval; gnathosomal bay U-shaped, proximally rounded; Cxgl-4 subapical, only slightly posterior to Cx-I tips; suture line of Cx-IV distinct, extending posteriorly beyond posterior margin of genital field; excretory pore in a medioposterior indentation of primary sclerotization, Vgl-2 posterior to excretory pore; gnathosomal rostrum long, ventral margin strongly curved (Fig. 3E); P-2 ventral margin slightly curved, ventrodistal protrusion slender, spatula-shaped, distally finely serrate; ventrodistal projection at P-3 irregularly cone-shaped, anterior margin frayed; P-4 relatively short, dorsal margin weakly curved, with ventral tubercles pointed and separated, bearing one long and three short setae (Figs. 3F���G). Male: medial suture line of Cx-II+III short; genital field subrectangular in shape; posterior suture lines of Cx-IV starting at right angle from genital field (Fig. 3C); ejaculatory complex apically long, cella proximalis small (Fig. 3D). Female: genital field roundish in shape Measurements. Male (holotype; in parentheses specimen from River Banano, n = 1)���Idiosoma (ventral view: Fig. 3C) L 650 (691), W 414 (450); dorsal shield (Figs. 3A, 11C) L 515 (534), W 363 (394), L/W ratio 1.4 (1.36); dorsal plate L 491 (511); shoulder plate L 159 (156���159), W 53 (53), L/W ratio 3.0 (2.8���2.9); frontal plate L 116��� 118 (114���116), W 42���44 (44), L/W ratio 2.6���2.8 (2.6); shoulder/frontal plate L 1.35���1.38 (1.35���1.4). Gnathosomal bay L 169 (173), Cx-I total L 292 (314), Cx-I mL 125 (141), Cx-II+III mL 78 (84); ratio Cx-I L/Cx-II+III mL 2.3 (2.2); Cx-I mL/Cx-II+III mL 3.7 (3.7). Genital field L/W 138 (141)/119 (124), ratio 1.16 (1.13); ejaculatory complex L 200 (222); distance genital field-excretory pore 89 (92), genital field-caudal idiosoma margin 141 (149). Gnathosoma vL 291 (325), chelicera L 345 (381); palp total L 274 (275), dL/H, dL/H ratio: P-1, 30/31, 0.95 (30/27, 1.1); P-2, 92/44, 2.1 (98/50, 2.0); P-3, 58/40, 1.45 (58/42, 1.37); P-4, 75/25, 3.0 (72/28, 2.6); P-5, 19/13, 1.5 (17/12, 1.5); P-2/P-4 ratio 1.23 (1.37). dL of I-L: 50, 56, 66, 83 (88), 95 (100), 86 (87); I-L-6 H 33 (33); dL/H I-L- 6 ratio 2.6 (2.6). Female (paratype, n = 2; in parentheses specimen from River Bouyouni, n = 1)���Idiosoma (ventral view: Fig. 4B) L 716���744 (740), W 497���519 (516); dorsal shield (Figs. 4A, 11D) L 572���575 (563), W 406���431 (422), L/W ratio 1.33���1.41 (1.33); dorsal plate L 541���550 (541); shoulder plate L 169���172 (159), W 56���59 (59), L/W ratio 2.8���3.0 (2.7); frontal plate L 119���131 (120���122), W 47���50 (49���50), L/W ratio 2.4���2.8 (2.4���2.5); shoulder/frontal plate L 1.29���1.42 (1.31���1.33). Gnathosomal bay L 186���195 (184), Cx-I total L 325���341 (325), Cx-I mL 140���145 (141), Cx-II+III mL 27���28 (34); ratio Cx-I L/Cx-I mL 2.3 (2.3); Cx-I mL/Cx-II+III mL 12.6 (9.6). Genital field L/ W 142���150 (147)/144���150 (141), ratio 0.95���1.04 (1.05); distance genital field-excretory pore 122���162 (138), genital field-caudal idiosoma margin 213���231 (231). Gnathosoma vL 331���344 (331); chelicera total L 413���419 (397); palp total L 302���311 (282), dL/H, dL/H ratio: P-1, 36���40/36���39, 1.0���1.12 (31/31, 1.0); P-2, 108/50���52, 2.09���2.16 (100/47, 2.13); P-3, 63���64/42���45, 1.42���1.48 (63/43, 1.44); P-4, 78���81/27���28, 2.8���3.0 (71/29, 2.45); P- 5, 17���18/14���16, 1.15���1.22 (17/14, 1.25); P-2/P-4 ratio 1.33���1.38 (1.41). dL of I-L: 52, 61, 69, 89 (92), 100 (100), 84 (80); I-L-6 H 34 (31); dL/H I-L-6 ratio 2.45 (2.6). Etymology. Named for its occurrence on Mayotte. Remarks. Smit et al. (2010) reported a single female of Torrenticola from Demb��ni River. Later on, the same authors assigned this specimen to T. comorosensis Pe��ić & Smit, 2015 described originally on the basis of a male and female from a tributary of Koual�� River (Pe��ić et al. 2015a). Additional material of this study reveals that females from Demb��ni River fit perfectly the description of the unnamed specimen described by Smit et al. (2010), but they have to be assigned to the new species T. mayottensis sp. nov. This new species differs from T. comorosensis (data taken from Pe��ić et al. 2015a) in a number of morphological features: 1) dorsal shield with a colour pattern (without colour in T. comorosensis, compare Figs. 11C���D with Figs. 4A, D in Pe��ić et al. 2015a), 2) gnathosomal rostrum comparatively longer (compare Fig. 3E with Fig. 2D in Pe��ić et al. 2015a), and 3) P-4 stouter in the both sexes, L/H 2.4���3.0 (3.6���3.7 in T. comorosensis). The female of the new species can be separated from T. comorosensis on the basis of more roundish genital field, L/ W 0.95���1.05 vs. more elongated in T. comorosensis, L/W ratio 1.12 (compare Fig. 4B with Fig. 3C in Pe��ić et al. 2015). Distribution. Comoros: Mayotte., Published as part of Pe��i��, Vladimir, Smit, Harry & Mary, Nathalie, 2018, Fifth contribution to the knowledge of water mites (Acari: Hydrachnidia) from the Comoros: a checklist and description of one new genus and four new species, pp. 331-348 in Zootaxa 4483 (2) on pages 334-338, DOI: 10.11646/zootaxa.4483.2.6, http://zenodo.org/record/1437784, {"references":["Smit, H., Pesic, V. & Mary-Sasal, N. (2010) Second contribution to the knowledge of water mites from the Comoros, with the description of one new species (Acari: Hydrachnidia). Zootaxa, 2413, 51 - 60."]}

Atractides comorosensis Smit & Pesic, 2009 Material examined. Comoros, Mayotte: River Bouyouni, 12 �� 44 ' 55,6 "S, 45 ��08' 36,4 "E, 16 -iv- 2009, 4/ 19 / 1 (1 /0/0 mounted); River Bouyouni, upstream, 12 �� 45 ' 38,4 "S, 45 ��08' 46,7 "E, alt. 105 m asl., 19 -iv- 2009, 0/ 5 [1 juvenile]/0; ibid., 30 -vii- 2012, 0/ 1 /0; River Bouyouni, 12 �� 44 ' 25,35 "S, 45 ��08' 33,3 "E, 16 -iv- 2009, 2[1 juvenile]/ 10 [1 juvenile]/ 0; River Demb��ni, 20 -iv- 2009, 0/ 1 /0; Demb��ni River, 12 �� 50 ' 24,5 "S, 45 �� 10 ' 33,2 "E, 19 -iv- 2009, 0/0/ 1; River Goulou��, 12 �� 47 ' 27,2 "S, 45 �� 11 ' 22,9 "E, 15 -iv- 2009, 3/ 7 /0; River Koual��, tributary, 12 �� 47 ' 57,35 "S, 45 ��09' 51,2 "E, 21 -iv- 2009, 0/ 4 /0; River Koual��, 12 �� 47 ' 49,7 "S, 45 �� 11 ' 8,6 "E, 21 -vi- 2009, 0/ 2 /0; River Koual��, 12 �� 48 ' 22,1 "S, 45 �� 11 ' 55,1 "E, 13 -iv- 2009, 0/ 1 /0; River Koual��, 12 �� 48 ' 19,5 "S, 45 ��09' 49,7 "E, 21 -iv- 2009, 3/ 4 /0; River Longoni, 12 �� 44 ' 20,05 "S, 45 ��09' 53,6 "E, 16 -iv- 2009, 3/ 27 /0; River Mroni Mouala, 12 �� 46 ' 17,2 "S, 45 ��08' 51,1 "E, 20 -iv- 2009, 1/ 4 / 1; River Mtsangachehi, 12 �� 53 '04,15"S, 45 ��07' 55,4 "E, 14 -iv- 2009, 1/ 1 /0; River Ourov��ni, 12 �� 48 ' 29,9 "S, 45 ��07' 39,4 "E, 17 -iv- 2009, 1/ 1 /0. Distribution. Comoros: Mayotte., Published as part of Pe��i��, Vladimir, Smit, Harry & Mary, Nathalie, 2015, Third contribution to the knowledge of water mites from the Comoros, with the description of two new species (Acari: Hydrachnidia), pp. 445-459 in Zootaxa 3964 (4) on page 456, DOI: 10.11646/zootaxa.3964.4.4, http://zenodo.org/record/237029, {"references":["Smit, H., Pesic, V. & Mary-Sasal, N. (2009) New species of water mites from the Comoros (Acari: Hydrachnidia). Zootaxa, 2213, 47 - 56."]}

Lagoenaxonopsalbia comorosensis Pe��ić & Smit sp. nov. (Figs. 6���8) Type material. Holotype male (MNHN), dissected and slide mounted, Comoros, Mayotte, Hajangoua stream, upstream, 12��52'1,0"S, 45��10'25,7"E, alt. 265 m asl., 18-vii-2015, leg. Mary. Paratypes (RMNH, MNHN): same data as holotype, 1/3/0; River Antanana, upstream, 12��57'55,7"S, 45��08'6,5"E, alt. 192 m asl., 30-vii-2014 0/1/0 (dissected and slide mounted); River Djalimou, upstream, 12��56'58,4"S, 45��07'23,7"E, alt. 165 m asl., 30-viii-2014 0/1/0 (dissected and slide mounted, MNHN); ibid., 16-vii-2015, 0/2/0 (0/1/0 dissected and slide mounted, RMNH). Diagnosis. As given for genus. Description. Male (holotype)���Dorsal and ventral shields fused posteriorly; dorsal shield L/W 450/328; excretory pore plate L/W 91/120, excretory pore flanked by numerous thin setae; ventral shield L/W 391/375; gnathosomal bay L 82; distance between projections associated with IV-L-insertions 198; width between the outer edges of the most lateral pair of acetabula 102. Ejaculatory complex with extremely elongated, straight posterior arms (Fig. 6H). Palp: total L, dL/H, dL/H ratio: P-1, 20/16, 1.24; P-2, 47/30, 1.58; P-3, 27/25, 1.06; P-4, 61/23, 2.7; P-5, 26/10, 2.6; P-2/P-4 ratio 0.77; ventral margin of P-2 straight, without extension, P-4 not expanded ventrally, without heavy setae (Figs. 6D���E); gnathosoma vL 59. Legs: dL of I-L: 34, 41, 47, 56, 70, 55; I-L-6 central H 22, I-L-6 dL/central H ratio 2.5; dL of IV-L: 69, 73, 92, 92, 91, 86; a few swimming setae, restricted to distal margins of penultimate leg segments; numbers: III-L-5, 1, IV- L-5, 2. Female (from River Antanana, n = 1; in parentheses specimen from River Djalimou, n = 1)���Idiosoma L/W; dorsal shield L/W 447 (441)/344 (338), ratio 1.3 (1.31), Dgl-3 and -4 located close to each other; ventral shield L/ W 391 (384)/394 (372); gnathosomal bay L 92 (91), distance between IV-L-insertions 222 (211); posterior margin of the ventral shield irregular in shape, forming a large membranous genital bay with a longish gonopore; genital plates with postgenital gland flanked by two setae (Figs. 8B���C arrows); width between outer edges of acetabular plates 152. Palp: total L 195 (197); dL/H, dL/H ratio: P-1, 24/18-19, 1.27 (25/19, 1.33); P-2, 51/32, 1.59 (52/31, 1.66); P- 3, 29/25, 1.19 (29/25, 1.19); P-4, 62/19, 3.2 (62/19, 3.2); P-5, 29/12, 2.5 (29/11, 2.7); L P-2/P-4 ratio 0.83 (0.84); palp morphology as in male; gnathosoma vL 82 (92). Legs: dL of I-L: 35 (30), 43 (37), 43 (44), 60 (59), 68 (69), 53 (54); I-L-6 central H (20), I-L-6 dL/central H ratio (2.7); dL of IV-L: 83, 66, 80, 88, 89, 84; swimming setae numbers: III-L-5, 2, IV-L-5, 2. Etymology. Named after the archipelago (Comoros) where the new species was found. Distribution. Comoros (Mayotte), known only from the loci typici., Published as part of Pe��i��, Vladimir, Smit, Harry & Mary, Nathalie, 2018, Fifth contribution to the knowledge of water mites (Acari: Hydrachnidia) from the Comoros: a checklist and description of one new genus and four new species, pp. 331-348 in Zootaxa 4483 (2) on page 340, DOI: 10.11646/zootaxa.4483.2.6, http://zenodo.org/record/1437784

Unionicola (Majumderatax) srilankana sp. nov. (Figs. 1���3) Type material. Holotype female (RMNH), dissected and slide mounted, Sri Lanka, Mahaweli River E of Gallella; slow water flow, width 15 m, depth 0.5 m, sandy bottom with organic matter, Eichhornia crassipes on water surface, 7��54'35.6"N, 81��06'15.2"E, 39 m asl., 2.i.2017, leg. A. Zawal. Paratypes: one male, one deutonymph, same data as holotype, dissected and slide mounted (RMNH). Diagnosis. Dorsum with a two pairs of muscle attachment platelets; P-4 with a serrated hyaline margin and two proximal projections on ventral margin; in the female anterior pair of genital plates medially with a cone-shaped extension bearing three peg-like setae. Cx-II and -III with hook-like extensions. Description. General features (Adults). Dorsum with two pairs of muscle attachment platelets, anterior platelet elongated and close to each other, medial to Dgl-1, posterior platelet further apart, posteromedial to Dgl-2, knob-shaped (Fig. 1A). Coxae with a reticulate pattern, Cx-II laterally with two hook-like extensions, Cx-III laterally with two hooklike extensions. Cx-IV each with a short posterior apodeme (Fig. 1B) and a laterally directed extension. I-L-5 distally with three strong setae, two short stout setae located at distal margin and one larger and blade-like (Fig. 1E). Palp: P-2 medially with two short serrate setae; P-4 with a hyaline serrated ventral margin and with a small seta and two proximal projections on ventral margin, one distal bearing a peg-like seta; P-5 claw bifurcated into two very long, slender clawlets (Fig. 1C). Legs: number of swimming setae: III-L-4, 3; III-L-5, 3; IV-L-4, 3; IV-L-5, 3. Female (holotype). Idiosoma L 590, W 465. Genital field with paired anterior wing-shaped plates, two acetabula and medially a cone-shaped extension bearing three peg like setae, posterior plates with three acetabula in triangular arrangement (Fig. 1G). Palp: total L 412, dL/H, dL/H ratio: P-1, 19/55, 0.34; P-2, 128/80, 1.6; P-3, 42/71, 0.59; P-4, 159/ 44, 3.6; P-5, 64/36, 1.79; L P-2/P-4 ratio 0.81. Gnathosoma L 188, W 103; L chelicera 152, cheliceral claw 89. Legs: dL of I-L-4-6: 100, 89, 109; L of I-L-5 blade-like distoventral seta 63; dL of IV-L: 73, 92, 108, 145, 172, 145. Male (paratype). Idiosoma L 460. Genital field (L/W 80/131) with acetabula grouped, two anteriorly and three posteriorly and with a few small setae scattered over the genital plates (Fig. 2A): Palp (Fig. 2B): total L 374, dL/H, dL/H ratio: P-1, 16/47, 0.33; P-2, 103/69, 1.49; P-3, 39/58, 0.68; P-4, 144/43, 3.3; P-5, 72/30, 2.4; L P-2/P-4 ratio 0.72. Legs: dL of I-L-4-6: 88, 86, 88; L of I-L-5 blade-like distoventral seta (Fig. 2C) 55; dL of IV-L-4-6: 109, 152, 161. Deutonymph. Idiosoma L 331. Dorsum with one pair of muscle attachment platelet. Provisional genital field with two pairs of acetabula (Fig. 3A). Palp: total L 268, dL/H, dL/H ratio: P-1, 12/36, 0.33; P-2, 73/39, 1.88; P-3, 28/38, 0.75; P-4, 102/36, 2.83; P-5, 53/22, 2.43; L P-2/P-4 ratio 0.72; P-2 with one medial serrate seta (Fig. 3B). Legs: dL of I-L: 34, 39, 56, 58, 57, 78; L of I-L-5 blade-like distoventral seta 50; dL of IV-L: 39, 47, 59, 84, 103, 116; number of swimming setae: III-L-4, 2; III-L-5, 2; IV-L-4, 2; IV-L-5, 2. Etymology. The new species is named after its occurrence on Sri Lanka. Remarks. The combination of P-4 with a hyaline serrated ventral margin bearing a small seta and the two medial serrate setae on P-2 makes the new species from Sri Lanka most similar to Unionicola grandiseta Savatenalinton & Smit, 2017 described on the basis of two females from Thailand (Savatenalinton & Smit 2017). The female of latter species can easily be distinguished from Unionicola srilankana sp. nov. in having one pair of platelet on the dorsum, one small ventral seta located behind the hyaline serrated margin of P-4, and with one peg-like seta only on the anterior genital plate. Distribution. Sri Lanka; known only from the type locality., Published as part of Pe��i��, Vladimir & Zawal, Andrzej, 2018, A new species in the water mite subgenus Majumderatax Vidrine, 1993 from Sri Lanka (Acari: Hydrachnidia), pp. 346-350 in Zootaxa 4457 (2) on pages 346-349, DOI: 10.11646/zootaxa.4457.2.11, http://zenodo.org/record/1457692, {"references":["Savatenalinton, S. & Smit, H. (2017) New records of water mites from standing waters in Thailand, with the description of nine new species (Acari: Hydrachnidia). Zootaxa, 4312, 69 - 91. https: // doi. org / 10.11646 / zootaxa. 4312.1.3"]}

Carinozetes mangrovi Pfingstl, Lienhard & Jagersbacher-Baumann, 2014 Records associated with mangroves: Great Bay, Concrete Beach and Vine Cave, Bermuda, from Bostrychia on roots of Avicennia germinans; Walsingham Pond and Hungry Bay, Bermuda, Bostrychia montagnei from roots of Rhizophora mangle (Pfingstl et al. 2014). R��o San Juan, GriGri Lagoon, Dominican Republic, intertidal algae growing on pneumatophores of Rhizophora mangle (Pfingstl et al. 2016). Records from other habitats: Boca Chica and Puerto Santa B��rbara, Dominican Republic, algae (Bostrychia) growing on rocks (Pfingstl et al. 2016). Distribution: Bermuda, Dominican Republic. Remarks: Although only reported from Bermuda and the Dominican Republic this species shows a wider distribution in the northern Caribbean area (Pfingstl, unpublished)., Published as part of Chatterjee, Tapas, Pfingstl, Tobias & Pe��i��, Vladimir, 2018, A checklist of marine littoral mites (Acari) associated with mangroves, pp. 221-240 in Zootaxa 4442 (2) on page 232, DOI: 10.11646/zootaxa.4442.2.2, http://zenodo.org/record/1303078, {"references":["Pfingstl, T., Lienhard, A. & Jagersbacher-Baumann, J. (2014) Hidden in the mangrove forest: the cryptic intertidal mite Carinozetes mangrovi sp. nov. (Acari, Oribatida, Selenoribatidae). Experimental and Applied Acarology, 63, 481 - 495. https: // doi. org / 10.1007 / s 10493 - 014 - 9802 - 2"]}

Arotrobates lanceolatus Luxton, 1992 Records associated with mangroves: Sha Tan, Hong Kong, small stones with green algae in mangrove; Wu Kai Sha, Hong Kong, Chaetomorpha in mangrove (Luxton 1992). Distribution: Hong Kong., Published as part of Chatterjee, Tapas, Pfingstl, Tobias & Pe��i��, Vladimir, 2018, A checklist of marine littoral mites (Acari) associated with mangroves, pp. 221-240 in Zootaxa 4442 (2) on page 232, DOI: 10.11646/zootaxa.4442.2.2, http://zenodo.org/record/1303078, {"references":["Luxton, M. (1992) Oribatid mites from the marine littoral of Hong Kong (Acari: Cryptostigmata) In: Morton, B. (Ed.), The marine flora and fauna of Hong Kong and southern China III. Proceedings of the Fourth International marine Biological Workshop: The marine Flora and Fauna of Hong Kong and Southern China. Hong Kong University Press, Hong Kong, pp. 211 - 227."]}

Copidognathus lutarius Bartsch, 2003 Record associated with mangroves: East coast of the Burrup Peninsula, Western Australia, algal turf on pneumatophores of Avicennia mangroves (Bartsch 2003b). Distribution: Northwestern Australia., Published as part of Chatterjee, Tapas, Pfingstl, Tobias & Pe��i��, Vladimir, 2018, A checklist of marine littoral mites (Acari) associated with mangroves, pp. 221-240 in Zootaxa 4442 (2) on page 225, DOI: 10.11646/zootaxa.4442.2.2, http://zenodo.org/record/1303078, {"references":["Bartsch, I. (2003 b) Mangrove halacarid fauna (Halacaridae, Acari) of the Dampier region, Western Australia, with description of five new species. Journal of Natural History, 37, 1855 - 1877. https: // doi. org / 10.1080 / 00222930110089184"]}

Eutrachytes flagellates Moraza, Kontsch��n, Sahoo & Ansari, 2016 Records associated with mangroves: Divar Island, Mandovi estuary, Goa, India, pneumatophores of Avicennia officinalis having algal (Bostryschia sp.) growth (Moraza et al. 2016). Distribution: India., Published as part of Chatterjee, Tapas, Pfingstl, Tobias & Pe��i��, Vladimir, 2018, A checklist of marine littoral mites (Acari) associated with mangroves, pp. 221-240 in Zootaxa 4442 (2) on page 228, DOI: 10.11646/zootaxa.4442.2.2, http://zenodo.org/record/1303078, {"references":["Moraza, M. L., Kontschan, J., Sahoo, G. & Ansari, Z. A. (2016) A new species of Eutrachytes (Acari: Uropodina: Eutrachytidae) associated with the Indian mangrove (Avicennia officinalis). Acarologia, 56, 73 - 89. https: // doi. org / 10.1051 / acarologia / 20162189"]}

Litoribates bonairensis Pfingstl, Baumann, Lienhard & Schatz, 2017 Records associated with mangroves: Lac Bay, Lesser Antilles, Bonaire, mangrove leaves and leaf litter under Rhizophora mangle (Pfingstl et al. 2017). Records from other habitats: Punta Ausente and Santa Ana, Isla Margarita, Venezuela, soil and leaf litter (Ermilov 2016). Distribution: Bonaire, Venezuela Remarks: The record in Venezuela (Ermilov 2016) is listed under Alismobates sp. but refers to L. bonairensis., Published as part of Chatterjee, Tapas, Pfingstl, Tobias & Pe��i��, Vladimir, 2018, A checklist of marine littoral mites (Acari) associated with mangroves, pp. 221-240 in Zootaxa 4442 (2) on page 229, DOI: 10.11646/zootaxa.4442.2.2, http://zenodo.org/record/1303078, {"references":["Pfingstl, T., Baumann, J., Lienhard, A. & Schatz, H. (2017) New Fortuyniidae and Selenoribatidae (Acari, Oribatida) from Bonaire (Lesser Antilles) and morphometric comparison between Eastern Pacific and Caribbean populations of Fortuyniidae. Systematic and Applied Acarology, 22, 2190 - 2217. https: // doi. org / 10.11158 / saa. 22.12.11","Ermilov, S. (2016) Additions to the oribatid mite fauna of Venezuela, with description of two new species of the family Achipteriidae (Acari, Oribatida). Systematic and Applied Acarology, 21, 1591 - 1603. https: // doi. org / 10.11158 / saa. 21.12.1"]}

Alismobates inexpectatus Pfingstl & Schuster, 2012 Records associated with mangroves: Coney Island, Lover���s lake and the ���Lagoon���, Bermuda, Bostrychia on mangrove roots (Avicennia germinans) (Pfingstl 2013a). R��o San Juan, GriGri Lagoon, Dominican Republic, intertidal algae growing on pneumatophores of red mangrove (Rhizophora mangle) (Pfingstl et al. 2016). Records from other habitats: Tobacco Bay, Whalebone Bay, Devonshire Bay and numerous other locations on the island of Bermuda, Bostrychia tenella growing on intertidal rocks���(Pfingstl & Schuster 2012b). R��o San Juan, Playa Grande, Dominican Republic, intertidal algae growing on rock; Boca Chica and Puerto Santa B��rbara, Dominican Republic, Bostrychia sp. from littoral rocks (Pfingstl et al. 2016). Distribution: Bermuda, Dominican Republic. Remarks: Although this species can be found occasionally in mangrove habitats, it prefers rocky coasts where it grazes predominantly on Bostrychia tenella (Pfingstl 2013a)., Published as part of Chatterjee, Tapas, Pfingstl, Tobias & Pe��i��, Vladimir, 2018, A checklist of marine littoral mites (Acari) associated with mangroves, pp. 221-240 in Zootaxa 4442 (2) on page 229, DOI: 10.11646/zootaxa.4442.2.2, http://zenodo.org/record/1303078, {"references":["Pfingstl, T. (2013 a) Habitat use, feeding and reproductive traits of rocky-shore intertidal mites from Bermuda (Oribatida: Fortuyniidae and Selenoribatidae). Acarologia, 53, 369 - 382. https: // doi. org / 10.1051 / acarologia / 20132101","Pfingstl, T. & Schuster, R. (2012 b) First record of the littoral genus Alismobates (Acari: Oribatida) from the Atlantic ocean, with a redefinition of the family Fortuyniidae based on adult and juvenile morphology. Zootaxa, 3301, 1 - 33."]}

Amhyadesia pacifica Fain & Schuster 1984 Records associated with mangroves: Coconut Island, Camotes Sea, Philippines, from intertidal roots of mangrove (Fain & Schuster 1984). Distribution: Philippines., Published as part of Chatterjee, Tapas, Pfingstl, Tobias & Pe��i��, Vladimir, 2018, A checklist of marine littoral mites (Acari) associated with mangroves, pp. 221-240 in Zootaxa 4442 (2) on page 234, DOI: 10.11646/zootaxa.4442.2.2, http://zenodo.org/record/1303078, {"references":["Fain, A. & Schuster, R. (1984) Four new species of the genus Amhyadesia Fain and Ganning, 1979 from the intertidal zone of several Indo-pacific Islands (Acari: Hyadesidae). International Journal of Acarology, 10 (2), 103 - 111. https: // doi. org / 10.1080 / 01647958408683359"]}

Nesiacarus schusteri Schatz, 1994 Records associated with mangroves: Puerto Ayora, Isla Santa Cruz, Gal��pagos, decayed leaf litter under Rhizophora mangle; Lake Arcturus, Isla Genovesa, Gal��pagos, well-decayed leaf litter and black humus under Rhizophora mangle; Punta Espinosa, Isla Fernandina, Gal��pagos, moist and partially decayed mangrove leaf litter under Rhizophora mangle and Laguncularia racemosa trees (Schatz 1994). Distribution: Gal��pagos., Published as part of Chatterjee, Tapas, Pfingstl, Tobias & Pe��i��, Vladimir, 2018, A checklist of marine littoral mites (Acari) associated with mangroves, pp. 221-240 in Zootaxa 4442 (2) on page 231, DOI: 10.11646/zootaxa.4442.2.2, http://zenodo.org/record/1303078, {"references":["Schatz, H. (1994) Lohmanniidae (Acari: Oribatida) from the Galapagos Islands, the Cocos Islands, and Central America. Acarologia, 35, 267 - 287."]}

Agauopsis dasyderma Bartsch, 2003 Record associated with mangroves: East coast of the Burrup Peninsula, Western Australia, turf on trunk of Avicennia marina, also among epibionts (barnacles, mussels, turf) on the underside branches of A. marina mangroves (Bartsch 2003b). Distribution: Western Australia., Published as part of Chatterjee, Tapas, Pfingstl, Tobias & Pe��i��, Vladimir, 2018, A checklist of marine littoral mites (Acari) associated with mangroves, pp. 221-240 in Zootaxa 4442 (2) on page 223, DOI: 10.11646/zootaxa.4442.2.2, http://zenodo.org/record/1303078, {"references":["Bartsch, I. (2003 b) Mangrove halacarid fauna (Halacaridae, Acari) of the Dampier region, Western Australia, with description of five new species. Journal of Natural History, 37, 1855 - 1877. https: // doi. org / 10.1080 / 00222930110089184"]}

Copidognathus piger Bartsch, 2003 Record associated with mangroves: East coast of the Burrup Peninsula, Western Australia, turf on pneumatophores of Avicennia marina mangroves (Bartsch 2003b). Distribution: Western Australia., Published as part of Chatterjee, Tapas, Pfingstl, Tobias & Pe��i��, Vladimir, 2018, A checklist of marine littoral mites (Acari) associated with mangroves, pp. 221-240 in Zootaxa 4442 (2) on page 225, DOI: 10.11646/zootaxa.4442.2.2, http://zenodo.org/record/1303078, {"references":["Bartsch, I. (2003 b) Mangrove halacarid fauna (Halacaridae, Acari) of the Dampier region, Western Australia, with description of five new species. Journal of Natural History, 37, 1855 - 1877. https: // doi. org / 10.1080 / 00222930110089184"]}

Schusteria littorea Grandjean, 1968 Records associated with mangroves: Ara���� mangrove, S��o Sebasti��o, S��o Paulo, Brazil, green algae on mangroves (Pepato et al. 2010). Records from other habitats: S��o Sebasti��o, S��o Paulo, Brazil, algae from the rocky littoral ��� (Grandjean 1968). Distribution: Brazil., Published as part of Chatterjee, Tapas, Pfingstl, Tobias & Pe��i��, Vladimir, 2018, A checklist of marine littoral mites (Acari) associated with mangroves, pp. 221-240 in Zootaxa 4442 (2) on page 233, DOI: 10.11646/zootaxa.4442.2.2, http://zenodo.org/record/1303078, {"references":["Grandjean, F. (1968) Schusteria littorea n. g. n. sp. et les Selenoribatidae (Oribates). Acarologia, 10, 116 - 150.","Pepato, A. R, da Rocha, C. E. F. & Dunlop, J. A. (2010) Phylogenetic position of the acariform mites: sensitivity to homology assessment under total evidence. BMC Evolutionary Biology, 10, 235. https: // doi. org / 10.1186 / 1471 - 2148 - 10 - 235"]}

Notophthiracarus schatzii Niedbała, 1989 Records associated with mangroves: Punta Espinoza, Isla Fernandina, Gal��pagos, moist well-decayed mangrove litter under Laguncularia racemosa and Rhizophora mangle mangroves (Niedbała1989). Distribution: Gal��pagos. Remarks: This species was only reported from mangrove habitats and is probably endemic to the Gal��pagos Islands (Niedbała & Schatz 1996)., Published as part of Chatterjee, Tapas, Pfingstl, Tobias & Pe��i��, Vladimir, 2018, A checklist of marine littoral mites (Acari) associated with mangroves, pp. 221-240 in Zootaxa 4442 (2) on page 231, DOI: 10.11646/zootaxa.4442.2.2, http://zenodo.org/record/1303078, {"references":["Niedbala, W. (1989) Notophthiracarus schatzii sp. nov. d´iles Galapagos (Acari, Oribatida, Phthiracaroides). Bulletin of the Polish Academy of Sciences, 37, 75 - 78.","Niedbala, W. & Schatz, H. (1996) Euptyctimous mites from the Galapagos Islands, Cocos Island, and Central America (Acari: Oribatida). Genus, 7, 239 - 317."]}

Fortuynia rotunda Marshall & Pugh, 2002 Records associated with mangroves: Barra reef near Inhambane, Mozambique, pneumatophores and stems of Avicennia mangroves (Marshall & Pugh 2002). Okukubi River, Kin Town, Okinawa Island, Japan, algae from mangrove forest floor (Karasawa & Hijii 2004). Distribution: Mozambique, Japan. Remarks: Fortuynia rotunda is the only species of this genus that inhabits exclusively mangrove habitats., Published as part of Chatterjee, Tapas, Pfingstl, Tobias & Pe��i��, Vladimir, 2018, A checklist of marine littoral mites (Acari) associated with mangroves, pp. 221-240 in Zootaxa 4442 (2) on page 230, DOI: 10.11646/zootaxa.4442.2.2, http://zenodo.org/record/1303078, {"references":["Marshall, D. J. & Pugh, P. J. A. (2002) Fortuynia (Acari: Oribatida: Ameronothroidea) from the marine littoral of southern Africa. Journal of Natural History, 36, 173 - 183. https: // doi. org / 10.1080 / 00222930010002775","Karasawa, S. & Hijii, N. (2004) Effects of microhabitat diversity and geographical isolation on oribatid mite (Acari: Oribatida) communities in mangrove forests. Pedobiologia, 48, 245 - 255. https: // doi. org / 10.1016 / j. pedobi. 2004.01.002"]}

Maculobates bruneiensis Ermilov, Chatterjee & Marshall, 2013 Records associated with mangroves: Brunei Bay, Pulau Bedukang, Brunei Darussalam, associated with mangrove tree bark and trunks���(Ermilov et al. 2013). Funaura Bay, Iriomote Island and Okukuibi River, Okinawa Island, Japan, bark of B. gymnorrhiza magrove (Iseki & Karasawa 2014). Distribution: Brunei Darussalam, Japan., Published as part of Chatterjee, Tapas, Pfingstl, Tobias & Pe��i��, Vladimir, 2018, A checklist of marine littoral mites (Acari) associated with mangroves, pp. 221-240 in Zootaxa 4442 (2) on page 230, DOI: 10.11646/zootaxa.4442.2.2, http://zenodo.org/record/1303078, {"references":["Ermilov, S. G., Chatterjee, T. & Marshall, D. J. (2013) Two new species of oribatid mites of Oripodoidea (Acari: Oribatida) from Brunei. Annales Zoologici, 63 (3), 393 - 400. https: // doi. org / 10.3161 / 000345413 X 672456","Iseki, A. & Karasawa, S. (2014) First record of Maculobates (Acari: Oribatida: Liebstadiidae) from Japan, with a redescription based on specimens from the Ryukyu Archipelago. Species Diversity, 19, 59 - 69. https: // doi. org / 10.12782 / sd. 19.1.059"]}

Lohmannia maya Iglesias & Palacios-Vargas, 2017 Records associated with mangroves: Isla Cozumel, Quintana Roo, Mexico, mangrove litter Rhyzophora mangle and Avicennia germinans on littoral sand (Iglesias & Palacios-Vargas 2017). Distribution: Mexico., Published as part of Chatterjee, Tapas, Pfingstl, Tobias & Pe��i��, Vladimir, 2018, A checklist of marine littoral mites (Acari) associated with mangroves, pp. 221-240 in Zootaxa 4442 (2) on page 230, DOI: 10.11646/zootaxa.4442.2.2, http://zenodo.org/record/1303078, {"references":["Iglesias, R. & Palacios-Vargas J. G. (2017) A new species of Lohmannia (Acari: Oribatei: Lohmanniidae) from mangroves at Quintana Roo (Mexico). Acarologia, 57, 655 - 663."]}

Litarachna minuta Pe��ić, Chatterjee & Marshall, 2013 Record associated with mangroves: Brunei Bay, Pulau Bedukang, Brunei Darussalam, among epiphytal algae from mangrove pneumatophores (Pe��ić et al. 2013). Distribution: Brunei Darussalam., Published as part of Chatterjee, Tapas, Pfingstl, Tobias & Pe��i��, Vladimir, 2018, A checklist of marine littoral mites (Acari) associated with mangroves, pp. 221-240 in Zootaxa 4442 (2) on page 228, DOI: 10.11646/zootaxa.4442.2.2, http://zenodo.org/record/1303078, {"references":["Pesic, V., Chatterjee, T. & Marshall, D. J. (2013) Marine water mites (Acari: Hydrachnidia, Pontarachnidae) from the Brunei Bay, with description of one new species. Cahiers de Biologie Marine, 54 (3), 405 - 410."]}

Arotrobates granulatus Luxton, 1992 Records associated with mangroves: Okukubi River, Kin Town, Okinawa Island, Japan, from bark of Bruguiera gymnorrhiza (black mangrove) and from algae on mangrove forest floor���(Karasawa & Aoki 2005). Records from other habitats: Starfish Bay, Hong Kong, rocky shore, from crevices with Saccostrea (oysters) at its mouth (Luxton 1992). Distribution: Hong Kong, Japan., Published as part of Chatterjee, Tapas, Pfingstl, Tobias & Pe��i��, Vladimir, 2018, A checklist of marine littoral mites (Acari) associated with mangroves, pp. 221-240 in Zootaxa 4442 (2) on page 232, DOI: 10.11646/zootaxa.4442.2.2, http://zenodo.org/record/1303078, {"references":["Luxton, M. (1992) Oribatid mites from the marine littoral of Hong Kong (Acari: Cryptostigmata) In: Morton, B. (Ed.), The marine flora and fauna of Hong Kong and southern China III. Proceedings of the Fourth International marine Biological Workshop: The marine Flora and Fauna of Hong Kong and Southern China. Hong Kong University Press, Hong Kong, pp. 211 - 227.","Karasawa, S. & Aoki, J. (2005) Oribatid Mites (Arachnida: Acari: Oribatida) from the Marine Littoral of the Ryukyu Archipelago, Southwestern Japan. Species Diversity, 10, 209 - 233. https: // doi. org / 10.12782 / specdiv. 10.209"]}

Schusteria melanomerus Marshall & Pugh 2000 Records associated with mangroves: Barra reef, near Inhambane, Mozambique, pneumatophores and stems of Avicennia mangroves (Marshall & Pugh 2000). Records from other habitats: Coast near Ukunda (south of Mombasa), Kenya, intertidal algae on rocks (Pfingstl 2016). Distribution: Mozambique, Kenya., Published as part of Chatterjee, Tapas, Pfingstl, Tobias & Pe��i��, Vladimir, 2018, A checklist of marine littoral mites (Acari) associated with mangroves, pp. 221-240 in Zootaxa 4442 (2) on page 233, DOI: 10.11646/zootaxa.4442.2.2, http://zenodo.org/record/1303078, {"references":["Marshall, D. J. & Pugh, P. J. A. (2000) Two new species of Schusteria (Acari: Oribatida: Ameronothroidea) from marine shores in southern Africa. African Zoology, 35 (2), 201 - 206. https: // doi. org / 10.1080 / 15627020.2000.11657091"]}

Litarachna caribica Pe��ić, Chatterjee & Schizas, 2008 Record associated with mangroves: Niewport, Netherland Antilles, Cura��ao, among epiphytal algae from red mangrove Rhizophora mangle roots (Pe��ić et al. 2008). Records from other habitats: Pe��ić et al. (2015) recorded this species in algae and rubble from Taboguilla Island, Panama (Pacific coast of Panama). Distribution: Caribbean Sea, Panama., Published as part of Chatterjee, Tapas, Pfingstl, Tobias & Pe��i��, Vladimir, 2018, A checklist of marine littoral mites (Acari) associated with mangroves, pp. 221-240 in Zootaxa 4442 (2) on page 228, DOI: 10.11646/zootaxa.4442.2.2, http://zenodo.org/record/1303078, {"references":["Pesic, V., Chatterjee, T. & Schizas, N. V. (2008) Marine water mites (Acari: Hydrachnidia: Pontarachnidae) from the Caribbean Sea, with description of one new species. Cahiers de Biologie Marine, 49 (3), 253 - 259.","Pesic, V., Chatterjee, T. & Schizas, N. V. (2015) First record of Litarachna caribica (Acari, Pontarachnidae) from the Pacific coast of Panama. Marine Biodiversity Records, 8 (e 85), 1 - 3. https: // doi. org / 10.1017 / S 1755267215000688"]}

Isobactrus australiensis Bartsch, 2003 Record associated with mangroves: East coast of the Burrup Peninsula, Western Australia, underside of Avicennia branches among barnacles (Bartsch 2003c). Distribution: Western Australia., Published as part of Chatterjee, Tapas, Pfingstl, Tobias & Pe��i��, Vladimir, 2018, A checklist of marine littoral mites (Acari) associated with mangroves, pp. 221-240 in Zootaxa 4442 (2) on page 226, DOI: 10.11646/zootaxa.4442.2.2, http://zenodo.org/record/1303078, {"references":["Bartsch, I. (2003 c) Rhombognathine mites (Halacaridae: Acari) from Dampier, Western Australia: taxonomy and biogeography. In: Wells, F. E., Walker, D. I. & Jones, D. S. (Eds.), The Marine Flora and Fauna of Dampier, Western Australia. Western Australian Museum, Perth, pp. 255 - 280."]}

Selenoribates quasimodo Pfingstl, 2013 Records associated with mangroves: Coney Island, Bermuda, algae growing on mangrove (Avicennia germinans) roots (Pfingstl 2013b). Distribution: Bermuda., Published as part of Chatterjee, Tapas, Pfingstl, Tobias & Pe��i��, Vladimir, 2018, A checklist of marine littoral mites (Acari) associated with mangroves, pp. 221-240 in Zootaxa 4442 (2) on page 233, DOI: 10.11646/zootaxa.4442.2.2, http://zenodo.org/record/1303078, {"references":["Pfingstl, T. (2013 b) Revealing the diversity of a once small taxon: the genus Selenoribates (Acari, Oribatida, Selenoribatidae). Zookeys, 312, 39 - 63. https: // doi. org / 10.3897 / zookeys. 312.5478"]}

Copidognathus bruneiensis Chatterjee, Marshall & Pe��ić, 2012 Record associated with mangroves: Kota Batu, Sungai Brunei Estuary and Bay, Brunei Darussalam, on mud flat and in algae covering the pneumatophores of Avicennia marina trees (Chatterjee et al. 2012b). Distribution: Brunei Darussalam., Published as part of Chatterjee, Tapas, Pfingstl, Tobias & Pe��i��, Vladimir, 2018, A checklist of marine littoral mites (Acari) associated with mangroves, pp. 221-240 in Zootaxa 4442 (2) on page 224, DOI: 10.11646/zootaxa.4442.2.2, http://zenodo.org/record/1303078, {"references":["Chatterjee, T., Marshall, D. J. & Pesic, V. (2012 b) New records of Copidognathus mites (Acari: Halacaridae) from mangroves in Brunei Darussalam with descriptions of two new species. Zootaxa, 3269, 18 - 30."]}

Galapagacarus schatzi Balogh, 1985 Records associated with mangroves: Bah��a Academy, Santa Cruz, Gal��pagos, in mangrove litter (Balogh 1985). Distribution: Gal��pagos., Published as part of Chatterjee, Tapas, Pfingstl, Tobias & Pe��i��, Vladimir, 2018, A checklist of marine littoral mites (Acari) associated with mangroves, pp. 221-240 in Zootaxa 4442 (2) on page 230, DOI: 10.11646/zootaxa.4442.2.2, http://zenodo.org/record/1303078, {"references":["Balogh, P. (1985) Galapagacarus schatzi gen. et sp. n. (Acari: Oribatei). Opuscula Zoologica Budapest, 21, 31 - 34."]}

Agauopsis longirostris Bartsch, 2015 Record associated with mangroves: End of Lim Chu Kang Road, Singapore, among green algae (Cladophorales) growing on mangrove (Bartsch 2015). Records from other habitats: Labrador Park, Singapore, among algae Catenella sp. (Rhodophyta) and other small algae on concrete piling (Bartsch 2015). Distribution: Singapore., Published as part of Chatterjee, Tapas, Pfingstl, Tobias & Pe��i��, Vladimir, 2018, A checklist of marine littoral mites (Acari) associated with mangroves, pp. 221-240 in Zootaxa 4442 (2) on page 224, DOI: 10.11646/zootaxa.4442.2.2, http://zenodo.org/record/1303078, {"references":["Bartsch, I. (2015) Halacaridae (Acari) amongst the epiflora and fauna on trunks, branches, roots, and pneumatophores on the coast of Singapore: A survey. Raffles Bulletin of Zoology, 31 (Supplement), 96 - 138."]}

Rhombognathus abirus Pepato & Da Silveira, 2015 Record associated with mangroves: Algal turfs on aerial roots, trunks and pneumatophores of mangroves Rhizophora mangle and Avicennia schaueriana from Ubatuba, S��o Paulo, Brazil, along the Fazenda River (Pepato & Da Silveira 2015). Distribution: Brazil., Published as part of Chatterjee, Tapas, Pfingstl, Tobias & Pe��i��, Vladimir, 2018, A checklist of marine littoral mites (Acari) associated with mangroves, pp. 221-240 in Zootaxa 4442 (2) on page 226, DOI: 10.11646/zootaxa.4442.2.2, http://zenodo.org/record/1303078, {"references":["Pepato, A. R. & Da Silveira, P. S. (2015) Two new species of Rhombognathus (Halacaridae, Trombidiformes) from a mangrove in the northern littoral zone of Sao Paulo state (Brazil). Zootaxa, 3905 (4), 500 - 510. https: // doi. org / 10.11646 / zootaxa. 3905.4.4"]}

Copidognathus riguus Bartsch, 2015 Record associated with mangroves: End of Lim Chu Kang Road, Singapore, mats of green algae amongst Avicennia mangrove trees (Bartsch 2015). Record from other habita ts: Pulau Ubin, OBS Camp NW, Singapore, sediment from beach (Bartsch 2015). Distribution: Singapore., Published as part of Chatterjee, Tapas, Pfingstl, Tobias & Pe��i��, Vladimir, 2018, A checklist of marine littoral mites (Acari) associated with mangroves, pp. 221-240 in Zootaxa 4442 (2) on page 226, DOI: 10.11646/zootaxa.4442.2.2, http://zenodo.org/record/1303078, {"references":["Bartsch, I. (2015) Halacaridae (Acari) amongst the epiflora and fauna on trunks, branches, roots, and pneumatophores on the coast of Singapore: A survey. Raffles Bulletin of Zoology, 31 (Supplement), 96 - 138."]}

Carinozetes bermudensis Pfingstl & Schuster, 2012 Records associated with mangroves: Concrete Beach and Lover���s Lake, Bermuda, algae on roots of Avicennia germinans (Pfingstl et al. 2014). Records from other habitats: Rocky Hill Park, Concrete Beach, Soldier Bay and numerous other locations on Bermuda, intertidal algae from rocks, boulder or soil (Pfingstl & Schuster 2012a, Pfingstl et al. 2014). Distribution: Bermuda. Remarks: Although this species is occasionally found in mangrove habitats it primarily prefers rocky or muddy shores without mangrove vegetation., Published as part of Chatterjee, Tapas, Pfingstl, Tobias & Pe��i��, Vladimir, 2018, A checklist of marine littoral mites (Acari) associated with mangroves, pp. 221-240 in Zootaxa 4442 (2) on page 232, DOI: 10.11646/zootaxa.4442.2.2, http://zenodo.org/record/1303078, {"references":["Pfingstl, T., Lienhard, A. & Jagersbacher-Baumann, J. (2014) Hidden in the mangrove forest: the cryptic intertidal mite Carinozetes mangrovi sp. nov. (Acari, Oribatida, Selenoribatidae). Experimental and Applied Acarology, 63, 481 - 495. https: // doi. org / 10.1007 / s 10493 - 014 - 9802 - 2","Pfingstl, T. & Schuster, R. (2012 a) Carinozetes nov. gen. (Acari: Oribatida) from Bermuda and remarks on the present status of the Family Selenoribatidae. Acarologia, 52, 377 - 409. https: // doi. org / 10.1051 / acarologia / 20122067"]}

Litoribates caelestis Pfingstl & Schatz, 2017 Records associated with mangroves: Pinnacle Rock, Bartolom�� Island, Gal��pagos, mangrove leaf litter; Puerto Ayora (near Charles Darwin Research Station), Santa Cruz Island, Gal��pagos, decaying mangrove leaf litter and soil under Rhizophora mangle and Avicennia germinans near shore (Pfingstl & Schatz 2017). Distribution: Gal��pagos., Published as part of Chatterjee, Tapas, Pfingstl, Tobias & Pe��i��, Vladimir, 2018, A checklist of marine littoral mites (Acari) associated with mangroves, pp. 221-240 in Zootaxa 4442 (2) on page 230, DOI: 10.11646/zootaxa.4442.2.2, http://zenodo.org/record/1303078

Fortuynia marina Hammen, 1960 Records associated with mangroves: Okukubi River, Kin Town, Okinawa Island, Japan, littoral algae from mangrove forest (Bruguiera gymnorrhiza and Rhizophora stylosa) floor (Karasawa & Hijii 2004). Records from other habitats: Near Base of the Royal Netherlands Navy, Biak Island, New Guinea, Cladophora socialis and red algae on stones in the intertidal zone (Hammen 1960) Distribution: Japan, Netherlands New Guinea., Published as part of Chatterjee, Tapas, Pfingstl, Tobias & Pe��i��, Vladimir, 2018, A checklist of marine littoral mites (Acari) associated with mangroves, pp. 221-240 in Zootaxa 4442 (2) on page 230, DOI: 10.11646/zootaxa.4442.2.2, http://zenodo.org/record/1303078, {"references":["Karasawa, S. & Hijii, N. (2004) Effects of microhabitat diversity and geographical isolation on oribatid mite (Acari: Oribatida) communities in mangrove forests. Pedobiologia, 48, 245 - 255. https: // doi. org / 10.1016 / j. pedobi. 2004.01.002"]}

Selenoribates satanicus Pfingstl, 2013 Records associated with mangroves: Lover���s Lake (marine pond), Bermuda, green algae growing on sandy substrate among mangrove (Avicennia germinans) roots (Pfingstl 2013b). Distribution: Bermuda., Published as part of Chatterjee, Tapas, Pfingstl, Tobias & Pe��i��, Vladimir, 2018, A checklist of marine littoral mites (Acari) associated with mangroves, pp. 221-240 in Zootaxa 4442 (2) on page 233, DOI: 10.11646/zootaxa.4442.2.2, http://zenodo.org/record/1303078, {"references":["Pfingstl, T. (2013 b) Revealing the diversity of a once small taxon: the genus Selenoribates (Acari, Oribatida, Selenoribatidae). Zookeys, 312, 39 - 63. https: // doi. org / 10.3897 / zookeys. 312.5478"]}

Litarachna bruneiensis Pe��ić, Chatterjee, Marshall & Pavićević, 2011 Record associated with mangroves: Kota Batu, Sangai Brunei Estuary and Bay, Brunei Darussalam, collected from dense Avicennia marina pneumatophore mats, depth about 2 m (Pe��ić et al. 2011). Pulau Bedukang, Brunei Bay, Brunei Darussalam, among epiphytal algae from mangrove pneumatophores (Pe��ić et al. 2013). Distribution: Brunei Darussalam., Published as part of Chatterjee, Tapas, Pfingstl, Tobias & Pe��i��, Vladimir, 2018, A checklist of marine littoral mites (Acari) associated with mangroves, pp. 221-240 in Zootaxa 4442 (2) on page 227, DOI: 10.11646/zootaxa.4442.2.2, http://zenodo.org/record/1303078, {"references":["Pesic, V., Chatterjee, T., Marshall, D. J. & Pavicevic, A. (2011) New records of water mites (Acari: Hydrachnidia) from Brunei Darussalam, Borneo, with descriptions of two new species. Zootaxa, 3018, 50 - 58.","Pesic, V., Chatterjee, T. & Marshall, D. J. (2013) Marine water mites (Acari: Hydrachnidia, Pontarachnidae) from the Brunei Bay, with description of one new species. Cahiers de Biologie Marine, 54 (3), 405 - 410."]}

Copidognathus glandulifer Bartsch, 1996 Record associated with mangroves: West Coast Park, Singapore, among Cladophorales (green algae) and also on algal tuft of red algae on Avicennia pneumatophores; end of Lim Chu Kang Road, Cladophorales (green algae) around Avicennia mangroves (Bartsch 2015). Records from other habitats: Macgillivray Reef, North east of Lizard Island, eastern Australia, 2m depth, coarse sand; Lizard Island, north end of Second beach, 1m depth, coarse sand and gravel; Heron reef, Heron island, 12 m depth fine to medium sand (Bartsch 1996i). St John���s Island, Singapore, sediment amongst colonies of polychaete tubes (Sabellariidae) (Bartsch 2015). Distribution: Eastern Australia, Singapore., Published as part of Chatterjee, Tapas, Pfingstl, Tobias & Pe��i��, Vladimir, 2018, A checklist of marine littoral mites (Acari) associated with mangroves, pp. 221-240 in Zootaxa 4442 (2) on page 224, DOI: 10.11646/zootaxa.4442.2.2, http://zenodo.org/record/1303078, {"references":["Bartsch, I. (1996) New records of Actacarus from the Caribbean area and notes on the subfamily Actacarinae and its species (Arachnida: Acari: Halacaridae). Senckenbergiana Biologica, 75, 229 - 241.","Bartsch, I. (2015) Halacaridae (Acari) amongst the epiflora and fauna on trunks, branches, roots, and pneumatophores on the coast of Singapore: A survey. Raffles Bulletin of Zoology, 31 (Supplement), 96 - 138."]}

Rhombognathus ocularis Bartsch, 2003 Record associated with mangroves: Western Australia, east coast of the Burrup Peninsula, in algal turf (mainly red algae Caloglossa and Bostrychia) present on branches, stem and pneumatophores of Avicennia marina mangroves (Bartsch 2003b, c). Distribution: Western Australia., Published as part of Chatterjee, Tapas, Pfingstl, Tobias & Pe��i��, Vladimir, 2018, A checklist of marine littoral mites (Acari) associated with mangroves, pp. 221-240 in Zootaxa 4442 (2) on page 227, DOI: 10.11646/zootaxa.4442.2.2, http://zenodo.org/record/1303078, {"references":["Bartsch, I. (2003 b) Mangrove halacarid fauna (Halacaridae, Acari) of the Dampier region, Western Australia, with description of five new species. Journal of Natural History, 37, 1855 - 1877. https: // doi. org / 10.1080 / 00222930110089184","Bartsch, I. (2003 c) Rhombognathine mites (Halacaridae: Acari) from Dampier, Western Australia: taxonomy and biogeography. In: Wells, F. E., Walker, D. I. & Jones, D. S. (Eds.), The Marine Flora and Fauna of Dampier, Western Australia. Western Australian Museum, Perth, pp. 255 - 280."]}

Isobactrus ponapensis Ab��, 1996 Record associated with mangroves: Pasir Ris Park, Singapore, from dead branch with shells of barnacles; Pulau Ubin, OBS Camp, Singapore, amongst root bale of uprooted palm (Bartsch 2015). Record from other habitats: Ab�� (1996) described this species among Phaeophyceae Turbinaria ornata and Monocotyledoneae Thalasia hemprichii, coarse, coral sandy sediment from Micronesia, Ponape Island (Central Pacific Ocean). Distribution: Ponape Island, Singapore., Published as part of Chatterjee, Tapas, Pfingstl, Tobias & Pe��i��, Vladimir, 2018, A checklist of marine littoral mites (Acari) associated with mangroves, pp. 221-240 in Zootaxa 4442 (2) on page 226, DOI: 10.11646/zootaxa.4442.2.2, http://zenodo.org/record/1303078, {"references":["Abe, H. (1996) A new species of the genus Isobactrus (Acari: Halacaridae) from Ponape, Micronesia, with special reference to intrageneric character state distribution. Species Diversity, 1, 17 - 29. https: // doi. org / 10.12782 / specdiv. 1.17","Bartsch, I. (2015) Halacaridae (Acari) amongst the epiflora and fauna on trunks, branches, roots, and pneumatophores on the coast of Singapore: A survey. Raffles Bulletin of Zoology, 31 (Supplement), 96 - 138."]}