Search

FAMILY PRODOTIIDAE FAM. NOV. (FIGS 19, 20C–F) Type genus: Prodotia Dall, 1924.

FAMILY PROSIPHONIDAE POWELL, 1951 (NEW RANK) (FIGS 8, 9) Type genus: Prosipho Thiele, 1912., Published as part of Kantor, Yuri I., Fedosov, Alexander E., Kosyan, Alisa R., Puillandre, Nicolas, Sorokin, Pavel A., Kano, Yasunori, Clark, Roger & Bouchet, Philippe, 2022, Molecular phylogeny and revised classification of the Buccinoidea (Neogastropoda), pp. 789-857 in Zoological Journal of the Linnean Society 194 on page 811, {"references":["Powell AWB. 1951. Antarctic. and Subantarctic Mollusca: Pelecypoda and Gastropoda. Discovery Reports 26: 47 - 196, pls 5 - 10.","Thiele J. 1912. Die antarktischen Schnecken und Muscheln. Deutschen Sudpolar-Expedition 1901 - 1903 13: 183 - 285, pls 11 - 19."]}

FAMILY BELOMITRIDAE KANTOR ET AL., 2012 (FIGS 3F, G, 4D���F) Type genus: Belomitra P.Fischer, 1883., Published as part of Kantor, Yuri I., Fedosov, Alexander E., Kosyan, Alisa R., Puillandre, Nicolas, Sorokin, Pavel A., Kano, Yasunori, Clark, Roger & Bouchet, Philippe, 2022, Molecular phylogeny and revised classification of the Buccinoidea (Neogastropoda), pp. 789-857 in Zoological Journal of the Linnean Society 194 on page 803

FAMILY EOSIPHONIDAE FAM. NOV. (FIGS 6E–L, 7) Type genus: Eosipho Thiele, 1929.

SUBFAMILY TOMLINIINAE SUBFAM. NOV. Type genus: Tomlinia Peile, 1937., Published as part of Kantor, Yuri I., Fedosov, Alexander E., Kosyan, Alisa R., Puillandre, Nicolas, Sorokin, Pavel A., Kano, Yasunori, Clark, Roger & Bouchet, Philippe, 2022, Molecular phylogeny and revised classification of the Buccinoidea (Neogastropoda), pp. 789-857 in Zoological Journal of the Linnean Society 194 on page 846

FAMILY COLIDAE GRAY, 1857 (NEW RANK) (FIGS 5F���L, 6D) Type genus: Colus R��ding, 1798., Published as part of Kantor, Yuri I., Fedosov, Alexander E., Kosyan, Alisa R., Puillandre, Nicolas, Sorokin, Pavel A., Kano, Yasunori, Clark, Roger & Bouchet, Philippe, 2022, Molecular phylogeny and revised classification of the Buccinoidea (Neogastropoda), pp. 789-857 in Zoological Journal of the Linnean Society 194 on page 806, {"references":["Gray JE. 1857. Guide to the systematic distribution of Mollusca in the British Museum. Part I. London: Taylor & Francis.","Roding PF. 1798. Museum Boltenianum ..., Pars secunda continens conchylia sive testacea univalvia, bivalvia & multivalvia. Hamburg: Christ."]}

International audience

Sibogasyrinx clausura sp. nov. (= PSH 10) urn:lsid:zoobank.org:act: 27ED0BF0-C78A-41E1-9064-E3EB0492864A Figs 10D–M, 11A–B Diagnosis Shell medium-sized, reaching 41.5 mm in length, fusiform, subsutural ramp concave on upper teleoconch whorls, weakly so on last whorl, nearly smooth (except for the axial riblets), shoulder bearing nodules on spire whorls, absent on last whorl. Spiral sculpture of weak, closely spaced cords on and below shoulder. Radula with central tooth and longitudinally folded marginal teeth with weakly thickened edges. Etymology Latin ‘ clausura ’ (noun in apposition), meaning ‘lockdown’, with reference to the sanitary restrictions associated with SARS-COV-2 that prevailed over much of the World when this manuscript was finalized. Material examined Holotype NEW CALEDONIA • Coral Sea, Coriolis Bank; 21°20′ S, 157°55′ E; depth 963–970 m; KANADEEP, stn CP4964; MNHN-IM-2013-48256. Other material (all sequenced) AUSTRALIA • 1 lv; New South Wales, off Byron Bay; 28°03′ S, 154°05′ E; depth 999–1013 m; R/V Investigator, cruise IN2017_ V03 _100; AMS C.519344. NEW CALEDONIA • 1 lv; Coral Sea, Coriolis Bank; 21°21′ S, 158°00′ E; depth 978–1000 m; KANADEEP, stn CP4963; MNHN-IM-2013-48244 • 3 lv; Coral Sea, Coriolis Bank; 21°20′ S, 157°50′ E; depth 961 m; KANADEEP, stn CP4965; MNHN-IM-2013-48144, MNHN-IM-2013-48169, MNHN-IM-2013-48258 • 2 lv; Coral Sea, Coriolis Bank; 21°10′ S, 157°46′ E; depth 975–980 m; KANADEEP, stn CP4966; MNHN-IM-2013-48167, MNHN-IM-2013-48235. SOLOMON ISLANDS • 1 lv; Sta Isabel I.; 08°47′ S, 159°38′ E; depth 762–1060 m; SALOMON 2, stn CP2182; MNHN-IM-2009-16763. Description MEASUREMENTS (holotype, largest specimen). SL 41.4 mm, AL (with canal) 23.0 mm, AL (without canal) 13.7 mm, SW 12.4 mm. SHELL (holotype). Moderately thick, slightly glossy, strong except for fragile and partially broken outer aperture lip, narrowly fusiform, with high spire and moderately long, straight siphonal canal. Protoconch small, globose, of about 1.5 strongly convex, eroded whorls. Protoconch/teleoconch transition marked by strongly arcuate axial rib, corresponding to shape of growth lines, followed by 10 thinner and weaker axial ribs and rather thickened growth lines. Protoconch diameter 1.0 mm, height 0.84 mm. Spire whorls distinctly angled at shoulder, last whorl with scarcely discernible shoulder. Total teleoconch whorls just over 8. Suture shallowly impressed, subsutural ramp broad, concave, on last whorl weakly concave. Subsutural region with a row of distinct, dense, narrow, short, prosocline axial wrinkles, corresponding to upper parts of thickened growth lines, extending from suture to upper ⅓ of ramp, increasing in number from 22 on first whorl to 42 on penultimate and 50 on last whorl. Shoulder of teleoconch whorls (except last one) with a row of distinct opisthoclinely elongated nodules, extending to abapical suture and intersected by spiral cords, increasing in number from 13 on first whorl to 21 on antepenultimate and penultimate whorls. Subsutural ramp smooth except for axial wrinkles mentioned above. Spiral cords intersecting shoulder nodules beginning on 2 rd teleoconch whorls (due to corrosion of shell their number is unclear) and becoming progressively stronger, 7 on antepenultimate and 8 on penultimate whorl. Cords closely spaced with intervals about half the width of cords. On last whorl cords covering entire shell surface below indistinct shoulder, about 40 cords in total, of which 20 on canal. Cords weak, slightly wavy, very closely spaced just below suture and with intervals 0.5–1.0 times cord width on shell base and canal. Shell base gradually narrowing towards narrow, nearly straight siphonal canal. Aperture narrow, constricted posteriorly, with narrow, moderately thick parietal callus, outer lip partially broken, evenly convex and weakly concave at transition to canal. Anal sinus shallow, subsutural, broadly arcuate, confluent with large forward extension of outer lip as deduced from growth lines. Shell off-white, with slightly darker subsutural ramp, protoconch light tan. Periostracum smooth, retained between cords and ribs. ANATOMY (n = 1; MNHN-IM-2009-16763). Male. Penis obliquely truncated at tip with short, large, conical papilla occupying entire anterior part of penis, surrounded by circular fold.Eyes present.Proboscis moderately long, conical, with expanded base. Proboscis retractors not defined, entire posterior part of proboscis base muscular. Salivary glands small, not fused, with very long ducts that run within walls of oesophagus. Buccal mass moderately large, about ⅓ of proboscis length, basal, protruding backwards beyond proboscis base, radular sac lying outside proboscis. Venom gland very large, thick and strongly convoluted, very constricted before opening into oesophagus in region of nerve ring. Muscular bulb moderately large. RADULA (n = 1; MNHN-IM-2009-16763, AL 10.6 mm) (Fig. 11A–B). Relatively short, comprising 38 rows of teeth, 16 nascent. Radula length 1.65 mm (15.5% of AL without canal), width up to 250 μm (2.35% of AL without canal). Central tooth with basal plate, having distinct anterior and lateral borders and long, narrow, sharply-pointed cusp. Anterior margin overlapped by preceding row, posterior margin almost evenly rounded except for narrow protrusion adjoining cusp. Marginal teeth flat when formed, becoming trough-shaped with weakly thickened edges during maturation, folded longitudinally when fully formed, with both margins overlapping at tooth tip. The resulting folded tooth is moderately broad with a sharp pointed tip, border between margins appears as a narrow groove along anterior edge. Tooth folding occurs within 15–16 th row of teeth (counting from rear). Remarks The other specimens are very similar to the holotype in shell shape and sculpture. In some specimens there is very indistinct spiral striation on the subsutural ramp and a single very weak spiral cord may even be present. This species is most similar to Sibogasyrinx maximei sp. nov., but differs in having more numerous and narrower subsutural axial wrinkles (50 vs 43 on last whorl and 42 vs 34 on penultimate one) and less numerous (40 vs 50) spiral cords on the last whorl, the holotypes of both species being of almost the same size. Nevertheless, S. maximei sp. nov. is only represented in our material by a single specimen and thus its intraspecific variability remains unknown. The two species are broadly sympatric in the Solomon Islands, but are clearly differentiated in our molecular analysis. Distribution This species occurs in the Solomon Islands, New South Wales and on the Coriolis Bank in the Coral Sea, at 762–1060 m. This bathymetric range corresponds to a single haul in the Solomon Islands, while all other specimens were collected at 960–1000 m, indicating a narrower bathymetric range for the species.

Sibogasyrinx cf. pyramidalis (Schepman, 1913) 1 (= PSH 2) Figs 3A–G, 4A–B Material examined (all sequenced) PHILIPPINES • 1 lv; 15°20′ N, 121°37′ E; depth 593–600 m; AURORA 2007, stn CP2729; MNHN- IM-2009-13451. SOUTH CHINA SEA • 2 lv; 20°01′ N, 115°02′ E; depth 700–723 m; NanHai 2014, stn CP4118; MNHN- IM-2013-44574, MNHN-IM-2013-44605 • 1 lv; 20°29′ N, 116°08′ E; depth 590–633 m; DongSha 2014, stn CP4129; MNHN-IM-2013-50215. Description MEASUREMENTS (largest specimen). SL 65.2 mm, AL (with canal) 35.1 mm, AL (without canal) 21.0 mm, SW 18.5 mm. SHELL. Moderately thick, strong except for fragile outer aperture lip, fusiform, with high spire and long, narrow, straight siphonal canal. Protoconch small, globose, of about 1.5 strongly convex, microshagreened whorls, eroded or missing in all specimens. Early teleoconch whorls weakly to moderately angular at shoulder. Largest available specimen (SL 65.2 mm) of 10.3 teleoconch whorls. Suture shallowly impressed, subsutural ramp broad, weakly concave to flat. All teleoconch whorls with a subsutural row of nodules, corresponding in shape to upper parts of thickened growth lines. Nodules more distinct on upper whorls, increasing in number, up to 30 on penultimate whorl and 37 on last whorl in largest specimen. Subsutural ramp may be completely smooth, or sometimes with very weak spiral striation or dendritic lines. Shoulder with pronounced thickened and rounded nodules, reaching lower suture, weakly opisthocline on upper whorls and more strongly inclined and confluent with growth lines on lower whorls. Nodules evanesce on last and even penultimate whorls at SL over 50 mm and shoulder becomes evenly rounded; their number increase with size from 14–15 on upper whorls to 21–22 on lower whorls. Spiral sculpture of 3–6 moderately pronounced, closely spaced narrow cords on shoulder, crossing shoulder nodules on spire whorls. Last whorl below shoulder with 40–45 cords varying in width, their intervals 0.5–2 times width of cords, also distinct on canal. Shell base gradually narrowing towards long, narrow, nearly straight siphonal canal. Aperture narrow, constricted posteriorly, with narrow and thin parietal callus, outer lip with rounded angle at shoulder, weakly convex below shoulder and weakly concave at transition to canal. Anal sinus moderately deep, subsutural, broadly arcuate, confluent with large forward extension of outer lip. Shell off-white with very light yellowish subsutural band, protoconch pale tan. Periostracum persists on part of shell, light yellowish. ANATOMY. Foregut similar to that of Sibogasyrinx cf. pyramidalis 2 (see below), except for presence of a large oval accessory salivary gland, adjoining oesophagus, with rather thick duct, entering proboscis and following along its wall. RADULA (n = 2; MNHN-IM-2009-13451, MNHN-IM-2013-50215). Comprising approximately 40 rows of teeth, 15 nascent, short, length ca 2.5 mm (15% of AL without canal). Radula width up to 530 μm (3.3% of AL without canal). Central tooth with broad, subrectangular basal plate and anteriorly broadly concave, borders distinct and with narrow but rather long, sharply pointed cusp. Marginal teeth with strongly thickened posterior edges, folded longitudinally. When immature, teeth nearly flat with elevated posterior edge, on developing part of radular tooth folding clearly visible (on Fig. 4A not fully folded teeth are marked by white arrows), bringing posterior and anterior edges close together. During tooth maturation the edges, particularly posterior one, progressively thicken, so that fully formed tooth appears duplex (Fig. 4B). Distribution The confirmed distribution is the South China Sea and the Philippines Sea, at 590– 700 m., Published as part of Kantor, Yuri I. & Puillandre, Nicolas, 2021, Rare, deep-water and similar: revision of Sibogasyrinx (Conoidea: Cochlespiridae), pp. 19-60 in European Journal of Taxonomy 773 on pages 30-32, DOI: 10.5852/ejt.2021.773.1509, http://zenodo.org/record/5536301, {"references":["Schepman M. M. 1913. The Prosobranchia of the Siboga Expedition. Part 5. Toxoglossa. Siboga-Expeditie 49 (5): 365 - 452. Available from https: // www. biodiversitylibrary. org / page / 34450646 [accessed 24 Aug. 2021].","Powell A. W. B. 1969. The family Turridae in the Indo-Pacific. Part 2. The subfamily Turriculinae. Indo- Pacific Mollusca 2 (10): 215 - 416. Available from https: // www. biodiversitylibrary. org / page / 49824050 [accessed 24 Aug. 2021]."]}

Sibogasyrinx archibenthalis (Powell, 1969) Fig. 3R–T Leucosyrinx (Sibogasyrinx) archibenthalis Powell, 1969: 344, pl. 264 figs 6–7. Material examined Holotype PHILIPPINES • Mindanao I., Iligan Bay, Tabu Point; 08°16′45″ N, 124°02′49″ E; depth 924 m; R/V Albatross, stn 5513; USNM 238773. Description MEASUREMENTS (holotype). SL 41.8 mm. SHELL. Fusiform, narrow, with high, very narrow, flat-sided spire and relatively short, almost straight, un-notched siphonal canal. Protoconch small, globose, of 1.5 smooth whorls. Teleoconch of 12 whorls. Spire whorl profile quite straight and descending steeply to a narrowly rounded, flange-like, peripheral carina, abutting lower suture. Last whorl distinctly angled by peripheral carina, and base rapidly contracting toward moderately long anterior canal. Axial sculpture of rounded peripheral nodules, about 13 per whorl, but these become obsolete on last two whorls; subsutural nodules or folds lacking. Spiral sculpture of dense well defined and regular spiral cords, 11–12 on subsutural ramp. Anal sinus moderately deep, subsutural, broadly arcuate, confluent with large forward extension of outer lip. Colour opaque white, covered by a thin pale buff periostracum. Remarks The species is known only from the holotype. In shell shape it is most similar to S. pyramidalis. The differences are most obvious when comparing specimens of the same size (e.g., Fig.3 D–F and 3R); the last whorl is lower and the canal is proportionally much shorter in S. archibenthalis, while the spiral cords on the subsutural ramp are more pronounced and are of the same strength as on the remaining part of the last whorl. The last whorl is more angular in S. archibenthalis and the shell has a higher number of teleoconch whorls; in the holotype of S. archibenthalis the number of teleoconch whorls is 12 (SL 43 mm), compared with 9.5 (SL 46.8 mm) in samples of S. cf. pyramidalis 1 from the Philippines of similar shell size. Therefore, at present we consider S. archibenthalis to be a valid species., Published as part of Kantor, Yuri I. & Puillandre, Nicolas, 2021, Rare, deep-water and similar: revision of Sibogasyrinx (Conoidea: Cochlespiridae), pp. 19-60 in European Journal of Taxonomy 773 on page 35, DOI: 10.5852/ejt.2021.773.1509, http://zenodo.org/record/5536301, {"references":["Powell A. W. B. 1969. The family Turridae in the Indo-Pacific. Part 2. The subfamily Turriculinae. Indo- Pacific Mollusca 2 (10): 215 - 416. Available from https: // www. biodiversitylibrary. org / page / 49824050 [accessed 24 Aug. 2021]."]}

Sibogasyrinx maximei sp. nov. (= PSH 9) urn:lsid:zoobank.org:act: EEC38F7C-DD00-46B9-9D56-3915EB0B2CD2 Figs 8E, 10A–C Diagnosis Shell medium-sized, reaching 41.6 mm in length, fusiform, subsutural ramp weakly concave on early teleoconch whorls, flat on later ones. It is nearly smooth (except for axial riblets). Shoulder of teleoconch whorls with nodules, absent on last whorl. Spiral sculpture weak, of indistinct closely spaced cords on and below shoulder. Radula with central tooth and longitudinally folded marginal teeth with weakly thickened edges. Etymology This species is named after Maxime, the son of the second author. Material examined Holotype SOLOMON SEA • off Marshall Bennett I.; 08°38′ S, 151°46′ E; depth 720 m; MADEEP, stn DW4323; MNHN-IM-2013-45883. Description (holotype) MEASUREMENTS. SL 41.6 mm, AL (with canal) 23.2 mm, AL (without canal) 15.7 mm, SW 12.4 mm. SHELL. Moderately thick, strong except for fragile and partially broken outer aperture lip, fusiform, with high, conical spire and moderately long, broad, straight siphonal canal. Protoconch small, globose, of about 1.5 strongly convex, microshagreened whorls. Protoconch/teleoconch transition marked by strongly arcuate axial rib, corresponding to shape of growth line. Protoconch diameter 1.0 mm, height 0.83 mm. Spire whorls weakly angled at shoulder, last whorl with hardly discernible shoulder. Total teleoconch whorls 8.5. Suture shallowly impressed, subsutural ramp broad, weakly concave on first four teleoconch whorls, flat on later whorls. Subsutural ramp with short, distinct, prosocline axial wrinkles, corresponding to upper parts of thickened growth lines, extending from suture to upper ⅓ of ramp, increasing in number from 20 on first whorl to 34 on penultimate and 43 on last whorl. Subsutural ramp with only traces of indistinct spiral striation. Shoulder of teleoconch whorls (except last one) with a row of distinct elongated nodules, these nearly orthocline on upper whorls and weakly opisthocline on penultimate whorl, 14–17 per whorl, reaching lower suture and intersected by spiral cords. Starting from 2 nd teleoconch whorl, spiral cords appearing on shoulder, these intersecting nodules and becoming progressively stronger, from two on 2 nd whorl to nine on penultimate one. Cords very closely spaced, separated by narrow grooves. On last whorl cords cover entire shell surface below indistinct shoulder, about 50 cords in total, of which 25 on canal. Cords weak, slightly wavy, very closely spaced just below suture and with intervals 0.5–1.0 times width of cords on shell base and canal. Shell base gradually narrowing towards moderately broad, almost straight siphonal canal. Aperture narrow, constricted posteriorly, with narrow and very thin parietal callus, outer lip partially broken, evenly convex and weakly concave at transition to canal. Anal sinus moderately deep, subsutural, broadly arcuate, confluent with large forward extension of outer lip. Shell very light yellowish, protoconch light tan. Periostracum smooth, tightly adhering. RADULA (n = 1; MNHN-IM-2013-45883) (Fig. 8E). Medium-long, comprising 40 rows of teeth, 12 nascent. Radula length 1.8 mm (11.4% of AL without canal), width up to 290 μm (1.85% of AL without canal). Central tooth with basal plate, having distinct borders and long, narrow, sharp cusp, anterior margin overlapped by preceding row, posterior margin formed by two straight sections meeting at obtuse angle in midline. Marginal teeth flat when formed, becoming trough-shaped with weakly thickened edges during maturation, folded longitudinally when fully formed, with both margins overlapping at tooth tip. Resulting folded tooth moderately broad with sharply pointed tip, border between both margins present as a narrow slit at anterior edge. Tooth folding occurring within 14 th row of teeth (counting from rear). Remarks The species is known from the holotype alone and is very similar to Sibogasyrinx clausura sp. nov. (for comparison see remarks under the following species). In shell outline it is very similar to S. cf. pyramidalis 1 and 2, but differs in having fewer nodules on the shoulder; the holotype has 17 nodules on the penultimate whorl, whereas specimens of S. cf. pyramidalis 1 and 2 of similar size have up to 26 nodules. There are additional differences in radular morphology. Distribution Known only from the type locality., Published as part of Kantor, Yuri I. & Puillandre, Nicolas, 2021, Rare, deep-water and similar: revision of Sibogasyrinx (Conoidea: Cochlespiridae), pp. 19-60 in European Journal of Taxonomy 773 on pages 46-48, DOI: 10.5852/ejt.2021.773.1509, http://zenodo.org/record/5536301

Sibogasyrinx filosa Ardovini, 2021 (= PSH 6) Figs 8C, 9A–H Sibogasyrinx filosus Ardovini, 2021: 5–6, textfigs (erroneous gender agreement of specific epithet). Material examined Holotype PAPUA NEW GUINEA • lv; NE of Bougainville I.; 04°30′ S, 157°20′ E; depth 1100–1200 m; MNHN-IM-2000-37629 (originally R. Ardovini collection). Paratype PAPUA NEW GUINEA • 1 lv; same collection data as for holotype; R. Ardovini collection. Other material (all sequenced) PAPUA NEW GUINEA • 1 lv; New Ireland; 02°33′ S, 150°45′ E; depth 150–170 m; KAVIENG 2014, stn DW4500; MNHN-IM-2013-59044. SOLOMON ISLANDS • 1 lv; off Choiseul I., Papua New Guinea Exclusive Economic Zone; 06°37′ S, 156°13′ E, depth 508–522 m; SALOMON 2, stn CP2227; MNHN-IM-2007-42498 • 1 lv; Guadalcanal I.; 09°19′ S, 160°06′ E; depth 416–425 m; SALOMONBOA 3, stn CP2767; MNHN-IM-2009-16831. Description MEASUREMENTS. Largest available specimen (MNHN-IM-2007-42498): SL 49.9 mm, AL (with canal) 28 mm, AL (without canal) 16.7 mm, SW 13.5 mm. Holotype has SL 52 mm. SHELL. Moderately thick, strong except for fragile and often partially broken outer aperture lip, narrowly fusiform, with high spire and long, narrow, straight siphonal canal. Protoconch small, globose, of just over 1.5 strongly convex, microshagreened whorls (MNHN-IM-2009-16831). Protoconch/teleoconch transition indistinct, marked by appearance of shoulder carina. Protoconch diameter 0.89 mm, height 0.72 mm. Spire whorls strongly angled at shoulder, last whorl with more rounded shoulder. Total teleoconch whorls just under 9 in largest specimen. Suture shallowly impressed on last whorl and rather deep on spire ones, subsutural ramp moderately broad, strongly concave. Subsutural region with a row of distinct narrow axial ribs, confluent with growth lines and forming small nodules at intersections with spiral cords, absent on most of first teleoconch whorl, about 20 on second–third whorls, about 35 on penultimate and 45 on last whorl. Subsutural ramp with spiral cords, first visible on second whorl and distinct on fourth whorl. Last, penultimate and antepenultimate whorls with 4–5 distinct cords on subsutural ramp, their intervals equal to width of cords, followed below shoulder by 5–7 more narrow, closely spaced cords. Shoulder with row of pronounced rounded nodules, more distinct on upper teleoconch whorls and absent on last whorl, 15–17 on penultimate and antepenultimate whorls. Spiral sculpture in addition to cords on subsutural ramp of distinct narrow cords, covering entire shell surface, including shoulder nodules. On last whorl about 45 cords below shoulder, 25 of which on canal. Cords weakly rounded or flat on top, their intervals mostly narrower than cords themselves, rarely equal to or even slightly wider than cords. Shell base gradually narrowing towards long, narrow, almost straight siphonal canal. Aperture narrow, constricted posteriorly, with very narrow and thin parietal callus, outer lip with rounded angle at shoulder, weakly convex below shoulder, weakly concave at transition to canal. Anal sinus moderately deep, subsutural, broadly arcuate, confluent with large forward extension of outer lip. Shell light orange with lighter middle part of last whorl, in sequenced specimens off-white, with very light yellowish subsutural ramp and irregular darker blotches on subsutural ramp. Protoconch light tan. ANATOMY (n = 1; MNHN-IM-2009-16831). Head with moderately long, conical tentacles, rounded on tips and with closely spaced bases, large eyes situated on small lobes at tentacle base. Proboscis conical, moderately long, with broad base, rapidly narrowing towards tip. Proboscis retractors not defined, entire posterior part of very thin proboscis sheath base weakly muscular. Buccal mass very large and broad, posterior to proboscis base, constituting about half of proboscis length, oesophagus forming a very short loop before nerve ring. Radular sac with small odontophore, opening dorso-laterally at right side of buccal mass. Venom gland thick, moderately long and convoluted, opening into oesophagus within nerve ring. Muscular bulb moderately large, elongated and folded in posterior part. Salivary glands fused, relatively large, of irregular shape, acinous. Small ovate accessory salivary gland situated dorsally at nerve ring. RADULA (n = 1; MNHN-IM-2009-16831) (Fig. 8C). Relatively short, consisting of 41 rows of teeth, 15 nascent. Radula length 1.55 mm (12% of AL without canal), width up to 225 μm (1.8% of AL without canal). Central tooth with subrectangular basal plate, having distinct borders and moderately long cusp. Anterior margin overlapped by preceding row, posterior margin evenly and weakly rounded. Marginal teeth flat when formed, becoming trough-shaped with weakly thickened edges during maturation, folded longitudinally when fully formed, with both margins overlapping at tooth tip. Resulting folded tooth moderately broad, with sharp pointed tip, anterior edge with a narrow slit between folded tooth margins. Remarks Our specimens are very similar to the holotype and paratype (except lighter shells) and were collected close to the type locality. Our specimens were collected at much shallower depths (150–508 m) as compared to the holotype and paratype (1100 m). Nevertheless, the strong similarity of the shell shape and sculpture, especially obvious when comparing the holotype with sequenced specimen of the same size (Fig. 9A–B and C–D) suggests the conspecifity of our and the type specimens. Ardovini (2021) himself considered the specimen MNHN-IM-2007-42498 as belonging to his newly described species based on the photograph on the MNHN website. Smaller specimens (SL 34.4 and 29.9 mm) retain smooth, flaking, yellowish periostracum, better preserved in interspaces between cords. This species is most similar to Sibogasyrinx subula sp. nov., but differs in having more strongly developed spiral cords on the subsutural ramp, a relatively narrower ramp resulting in a higher shoulder position on the spire whorls, a different radula and the presence of eyes. In shell outline the new species resembles some species of Leucosyrinx, particularly Leucosyrinx sp. A (Fig. 13H), but the shell is broader with more numerous nodules on the shoulder. Compared to Leucosyrinx sp. F (Fig. 13K), the shell of S. filosa is larger, broader and has a coarser sculpture. Distribution This species is found off the Solomon Islands Archipelago and New Ireland, at a broad depth range of 150–1100 m., Published as part of Kantor, Yuri I. & Puillandre, Nicolas, 2021, Rare, deep-water and similar: revision of Sibogasyrinx (Conoidea: Cochlespiridae), pp. 19-60 in European Journal of Taxonomy 773 on pages 41-43, DOI: 10.5852/ejt.2021.773.1509, http://zenodo.org/record/5536301, {"references":["Ardovini R. 2021. Descrizione di Sibogasyrinx filosus n. sp. (Neogastropoda, Conoidea, Cochlespiridae) da Papua Nuova Guinea, Oceano Pacifico Meridionale. Malacologia Mostra Mondiale 111: 5 - 6.","Powell A. W. B. 1969. The family Turridae in the Indo-Pacific. Part 2. The subfamily Turriculinae. Indo- Pacific Mollusca 2 (10): 215 - 416. Available from https: // www. biodiversitylibrary. org / page / 49824050 [accessed 24 Aug. 2021].","Kantor Y. I., Fedosov A. E. & Puillandre N. 2018. New and unusual deep-water Conoidea revised with shell, radula and DNA characters. Ruthenica, Russian Malacological Journal 28: 47 - 82."]}

Sibogasyrinx lolae sp. nov. (= PSH 8) urn:lsid:zoobank.org:act: 73404289-ED57-4ABD-A0D0-05939F095C40 Figs 8D, 9I–Q Diagnosis Shell medium-sized, reaching 43 mm in length, fusiform, with weakly concave, nearly smooth subsutural ramp and nodules on shoulder of upper teleoconch whorls, absent on later whorls of large specimens. Spiral sculpture weak, of closely spaced cords covering entire shell except subsutural ramp in some specimens. Radula with central tooth and longitudinally folded marginal teeth with weakly thickened edges. Etymology The species is named after Lola, the daughter of the second author. Material examined Holotype SOLOMON ISLANDS • W of San Cristobal I.; 10°26′ S, 161°20′ E; depth 506–567 m; SALOMONBOA 3, stn CP2839; MNHN-IM-2007-42537. Other material (all sequenced) NEW CALEDONIA • 1 lv; Bank de L’Orne/Walpole; 22°22′ S, 168°57′ E; depth 460–708 m; EXBODI, stn CP3864; MNHN-IM-2009-29230 • 1 lv; 21°58′ S, 167°07′ E; depth 511–1050 m; TERRASSES, stn CP3098; MNHN-IM-2009-29311 • 1 lv; Fairway Ridge; 21°39′ S, 162°42′ E; depth 787 m; KANADEEP, stn CP4923; MNHN-IM-2013-48156. Description MEASUREMENTS (holotype). SL 37.3 mm, AL (with canal) 22.9 mm, AL (without canal) 14.5 mm, SW 11.3 mm. SHELL (holotype). Moderately thick, glossy, strong except for very fragile and partially broken outer aperture lip, narrowly fusiform, with high spire and long, narrow, straight siphonal canal. Protoconch small, globose, corroded, of about 1.5 whorls. Protoconch preserved in other juvenile specimen (MNHN-IM-2013-48156), comprising just over 1.5 strongly convex, microshagreened whorls. Protoconch/teleoconch transition indistinct, marked by appearance of axial ribs. Protoconch diameter 1.0 mm, height 0.74 mm. Spire whorls angled at shoulder, last whorl with less angular shoulder. Total teleoconch whorls just under 8. Suture shallowly impressed on last whorl and rather deep on spire whorls, subsutural ramp moderately broad, weakly concave on upper teleoconch whorls and nearly flat on penultimate and last whorls. Subsutural ramp with distinct prosocline axial wrinkles, extending from suture to upper ⅔ of ramp, 22–23 on first and second whorls, 24 on penultimate and 31 on last whorl, and without spiral sculpture on upper four whorls and later with indistinct spiral cords, three on last whorl. Shoulder of teleoconch whorls (except last one) with a row of distinct, opisthoclinely elongated nodules, intersected by weak spiral cords. Seventeen nodules on first teleoconch whorl, 18 on antepenultimate and penultimate whorls. Weak spiral cords on and below shoulder, starting from 3 rd teleoconch whorls, about twice as wide below shoulder. Penultimate whorl with four cords on shoulder and four below shoulder. Shoulder smooth on last whorl, but with about 55 cords below shoulder, of which 25 on canal. Cords weak, slightly wavy and closely spaced, their intervals about half the width of cords. Shell base gradually narrowing towards long, narrow, nearly straight siphonal canal. Aperture narrow, constricted posteriorly, with narrow, very thin parietal callus, outer lip badly broken, distinctly impressed at shoulder, weakly convex below shoulder and shallowly concave at transition to canal. Anal sinus judging from growth lines shallow, subsutural, broadly arcuate, confluent with forward extension of outer lip. Shell very light yellowish, protoconch very light tan (in holotype) and light brown in MNHN-IM-2013-48156. ANATOMY (n = 1; MNHN-IM-2009-29311). Male. Penis tip obliquely truncated, with long and very narrow papilla, surrounded by circular fold, much larger in diameter than papilla itself. Proboscis conical, moderately long, with broad base, anterior half rapidly narrowing towards tip. Proboscis retractors distinct, arranged in two symmetrical lateral bundles, attached to inner proboscis walls at border of its posterior third. Buccal mass with small radular sac, odontophore situated within proboscis in its broader posterior part; elongate oval, occupying slightly less than half of proboscis length. Single small accessory salivary gland present. Salivary glands separate, acinous, irregular in shape. Venom gland long, moderately thick and convoluted, opening into oesophagus within the nerve ring. RADULA (n = 1; MNHN-IM-2009-29311) (Fig. 8D). Short, comprising 28 rows of teeth, 13 nascent. Radula length 1.15 mm (7.9% of AL without canal), width up to 255 μm (1.8% of AL without canal), tooth length 160 μm (1.10% of AL without canal). Central tooth with subrectangular basal plate, having distinct borders and long, narrow, sharp cusp. Anterior margin overlapped by preceding row, posterior margin formed by two straight sections meeting at obtuse angle in midline. Marginal teeth flat when formed, becoming trough-shaped with weakly thickened edges during maturation, folded longitudinally when fully formed, with both margins overlapping at tooth tip. Resulting folded tooth moderately broad, with sharp pointed tip, anterior edge with a narrow slit between tooth margins. Tooth folding occurring within 15 th row of teeth. Remarks The largest specimen attains 43.1 mm. Other specimens are similar to the holotype in shell shape, although some have much more pronounced spiral sculpture, especially MNHN-IM-2009-29230, which also retains spiral cords on the shoulder of the last whorl. In shell shape and sculpture the new species is most similar to Sibogasyrinx filosa, some specimens being almost indistinguishable (e.g., holotype of S. filosa and MNHN-IM-2009-29230). Sibogasyrinx lolae sp. nov. can be distinguished from S. filosa due to its generally less distinct spiral sculpture and less concave, nearly flat subsutural ramp. Other conchologically similar species are S. cf. pyramidalis 1 and 2, but these differ in their radular morphology. In both these species and S. lolae sp. nov., the nodules on the shoulder become less pronounced or disappear on later whorls. Nevertheless, the nodules are still pronounced on the shoulder of the last whorl in specimens of S. cf. pyramidalis 1 and 2 of the same size as specimens of S. lolae sp. nov. which already lack nodules on the last whorl. Distribution This species is recorded from the Solomon Islands, the Coral Sea and southern New Caledonia, at depths of 460– 787 m., Published as part of Kantor, Yuri I. & Puillandre, Nicolas, 2021, Rare, deep-water and similar: revision of Sibogasyrinx (Conoidea: Cochlespiridae), pp. 19-60 in European Journal of Taxonomy 773 on pages 43-46, DOI: 10.5852/ejt.2021.773.1509, http://zenodo.org/record/5536301, {"references":["Ardovini R. 2021. Descrizione di Sibogasyrinx filosus n. sp. (Neogastropoda, Conoidea, Cochlespiridae) da Papua Nuova Guinea, Oceano Pacifico Meridionale. Malacologia Mostra Mondiale 111: 5 - 6."]}

Genus Sibogasyrinx Powell, 1969 Leucosyrinx (Sibogasyrinx) Powell, 1969: 343. Type species Surcula pyramidalis Schepman, 1913 (original designation). Diagnosis Shell small to large, adult length from 27 to 65 mm, narrowly fusiform, rarely pagodiform, with concave to nearly flat subsutural ramp. Spiral sculpture variously developed, always present below shoulder, comprising narrow, close-set cords, often also on subsutural ramp, Shoulder with a row of strong nodules, often obsolete on last whorl. Venom gland does not pass through nerve ring and opens into oesophagus within or posterior to nerve ring. Radula usually with central tooth, absent in one species, marginal tooth morphology variable, folded longitudinally., Published as part of Kantor, Yuri I. & Puillandre, Nicolas, 2021, Rare, deep-water and similar: revision of Sibogasyrinx (Conoidea: Cochlespiridae), pp. 19-60 in European Journal of Taxonomy 773 on page 29, DOI: 10.5852/ejt.2021.773.1509, http://zenodo.org/record/5536301, {"references":["Powell A. W. B. 1969. The family Turridae in the Indo-Pacific. Part 2. The subfamily Turriculinae. Indo- Pacific Mollusca 2 (10): 215 - 416. Available from https: // www. biodiversitylibrary. org / page / 49824050 [accessed 24 Aug. 2021].","Schepman M. M. 1913. The Prosobranchia of the Siboga Expedition. Part 5. Toxoglossa. Siboga-Expeditie 49 (5): 365 - 452. Available from https: // www. biodiversitylibrary. org / page / 34450646 [accessed 24 Aug. 2021]."]}

Sibogasyrinx sangeri Kantor, Fedosov & Puillandre, 2018 (= PSH 7) Figs 7, 8A–B Sibogasyrinx sangeri Kantor, Fedosov & Puillandre, 2018: 58, figs 5e–n, 6c–d. Material examined Holotype PAPUA NEW GUINEA • 07°52′ S, 148°03′ E; depth 575–655 m; BIOPAPUA, stn CP3729; MNHN- IM-2009-17022 (sequenced). Other material (all sequenced) PAPUA NEW GUINEA • 2 lv; 04°04′ S, 151°56′ E; depth 585–601 m; BIOPAPUA, stn CP3671; MNHN-IM-2009-16989, MNHN-IM-2013-52052 • 1 lv; 04°24′ S, 151°50′ E; depth 788–805 m; BIOPAPUA, stn CP3674; MNHN-IM-2009-16995 • 1 lv; off Woodlark Is.; 09°08′ S, 152°19′ E; depth 448–470 m; BIOPAPUA, stn CP3742; MNHN-IM-2009-17057 • 1 lv; 05°39′ S, 153°59′ E; depth 654– 660 m; BIOPAPUA, stn CP3750; MNHN-IM-2009-17021 • 1 lv; N of Long I.; 05°10′ S, 147°03′ E; depth 724 m; PAPUA NIUGINI, stn CP3982; MNHN-IM-2013-19752 • 1 lv; Dampier Strait, E of Umboi I.; 05°35′ S, 148°13′ E; depth 630–870 m; PAPUA NIUGINI, stn CP4014; MNHN-IM-2013-19961. PHILIPPINES • 1 lv; 15°45′ N, 121°45′ E; depth 562 m; AURORA 2007, stn CP2663; MNHN- IM-2009-13434. SOLOMON ISLANDS • 1 lv; Sta Isabel I.; 08°47′ S, 159°40′ E; depth 645–840 m; SALOMON 2, stn CP2181; MNHN-IM-2009-16766 • 2 lv; Rendova I.; 08°36′ S, 157°27′ E; depth 509–520 m; SALOMON 2, stn CP2288; MNHN-IM-2007-42523, MNHN-IM-2009-16779. Description MEASUREMENTS. Holotype: SL 54.1 mm, AL (with canal) 31.3 mm, AL (without canal) 16.4 mm, SW 14.1 mm. Largest available specimen attains SL 55.8 mm (MNHN-IM-2009-16779; Fig. 7D–E). SHELL. Thin, fragile, variable in shape, from narrowly fusiform to moderately broad (SW/SL ratio from 0.22 to 0.27), with rather high spire and long, narrow, straight siphonal canal. Protoconch small, globose, of 1.75 strongly convex, microshagreened whorls. Protoconch/teleoconch transition indistinct. Protoconch diameter about 1.1 mm, height 1.2 mm. Early teleoconch whorls angular, usually in lower part just above suture, but sometimes at mid-whorl; last whorl strongly to very weakly angled or evenly rounded, even in specimens of the same size. Teleoconch comprising up to 10 whorls. Suture shallow, subsutural ramp with a row of distinct, closely spaced nodules, 10 on first whorl, 14 on second whorl, becoming more broadly spaced and less discernible on later whorls, sometimes obsolete on last whorl of largest specimens. Subsutural zone very weakly concave, nearly straight on penultimate and last whorls, smooth except for a few irregularly spaced and indistinct spiral threads (sometimes oblique), or several rather pronounced flattened spiral cords (Fig. 7J; MNHN-IM-2009-13434). Lower part of whorls with a row of larger, more pronounced nodules, just above suture in upper whorls or sometimes nearer midwhorl. In some specimens nodules absent on last whorl. Last whorl may bear low carina at periphery (in specimens with nodules absent), giving it very weakly angled outline. Below periphery 2–3 distinct spiral cords on penultimate whorl and about 30 cords below carina on last whorl and 20 on canal. Shell base gradually narrowing towards long, nearly straight siphonal canal. Aperture narrow, constricted posteriorly with broad, very thin parietal callus, outer lip partially broken, convex and weakly angled in upper part and slightly convex below shoulder, and shallowly concave at transition to canal. Anal sinus moderately deep, subsutural, broadly arcuate, growth lines confluent with large forward extension of outer lip. Growth lines thin but distinct. Shell uniformly off-white, protoconch light tan. ANATOMY (n = 1, MNHN-IM-2009-16995). Proboscis of moderate length in contracted stage, buccal mass situated outside proboscis. Oesophagus very broad, forming short loop before passing through nerve ring. Venom gland opening into oesophagus ventrally and immediately posterior to nerve ring. RADULA (n = 1, MNHN-IM-2009-16995) (Fig. 8A–B). Relatively short, comprising ca 40 rows of teeth, with long nascent part (15–16 rows). Radula length 2.2 mm (16% of AL without canal), width up to 365 μm (2.7% of AL without canal). Central tooth with subrectangular basal plate, very shallowly arcuate anteriorly, with distinct borders and weak cusp. Marginal teeth flat when formed, becoming trough-shaped during maturation, folded longitudinally when fully formed. On developing part of radula, folding of teeth occurs abruptly, within one subsequent row (on Fig. 8B white arrow indicates last still unfolded tooth, while black arrow with white outline indicate the first folded tooth). Folding evident at 17 th row in radula studied. Resulting folded tooth moderately broad, with sharp pointed tip, formed by overlapping of both thickened margins (on Fig. 8A these overlapping parts are marked by hollow white arrows). Distribution This species is known from the Solomon Islands, Papua New Guinea and the Philippines, over a relatively broad bathymetric range, from 448 to 788 m., Published as part of Kantor, Yuri I. & Puillandre, Nicolas, 2021, Rare, deep-water and similar: revision of Sibogasyrinx (Conoidea: Cochlespiridae), pp. 19-60 in European Journal of Taxonomy 773 on pages 38-40, DOI: 10.5852/ejt.2021.773.1509, http://zenodo.org/record/5536301, {"references":["Kantor Y. I., Fedosov A. E. & Puillandre N. 2018. New and unusual deep-water Conoidea revised with shell, radula and DNA characters. Ruthenica, Russian Malacological Journal 28: 47 - 82."]}

Sibogasyrinx subula sp. nov. (= PSH 3) urn:lsid:zoobank.org:act: 6A992BBF-7C62-4F15-BFCC-AD9F347ED379 Figs 4E–F, 6 Sibogasyrinx sp. – Bouchet et al. 2011: figs 11b, 12b–c. Diagnosis Shell large, reaching 57 mm in length, narrowly fusiform, with concave, nearly smooth subsutural ramp with darker subsutural band on posterior half of last whorl and strong nodules on shoulder. Spiral sculpture of distinct, closely spaced cords below subsutural ramp. Radula with central tooth and duplex marginal teeth. Etymology Latin ‘ subula ’ (noun in apposition), meaning ‘awl’, with reference to the general appearance of the spire. Material examined Holotype SOLOMON ISLANDS • SE of Sta Isabel I.; 08°17′ S, 160°00′ E; depth 482–604 m; SALOMON 2, stn CP2187; MNHN-IM-2007-42530. Other material (all sequenced) PAPUA NEW GUINEA • 1 lv; 05°39′ S, 153°59′ E; depth 654–660 m; BIOPAPUA, stn CP3750; MNHN-IM-2009-17048 • 1 lv; New Ireland; 02°13′ S, 150°12′ E; depth 564–743 m; KAVIENG 2014, stn CP4448; MNHN-IM-2013-58409. SOLOMON ISLANDS • 2 lv; SE of Sta Isabel I.; 08°17′ S, 160°00′ E; depth 464–523 m; SALOMON 2, stn CP2184; MNHN-IM-2009-13567, MNHN-IM-2009-13568 • 1 lv; SW of Sta Isabel I.; 08°25′ S, 159°27′ E; depth 440–521 m; SALOMON 2, stn CP2194; MNHN-IM-2007-42533 • 1 lv; 08°41′ S, 157°38′ E; depth 814–980 m; SALOMON 2, stn CP2276; MNHN-IM-2007-42517. VANUATU • 4 lv; 15°00′ S, 166°55′ E; depth 630–705 m; BOA1, stn CP2432; MNHN-IM-2007-17701 to IM-2007-17704. Description MEASUREMENTS (holotype). SL 55 mm, AL (with canal) 31 mm, AL (without canal) 17.5 mm, SW 15.6 mm. SHELL (holotype). Moderately thick, strong except for fragile outer lip of aperture, fusiform, with high spire and long, narrow, straight siphonal canal. Protoconch small, globose, of about 1.5 strongly convex, microshagreened whorls. Protoconch/teleoconch transition marked by arcuate axial rib and appearance of definitive sculpture. Protoconch diameter 1.0 mm, height 0.79 mm. Teleoconch whorls angular at shoulder. Teleoconch comprising just under 10 whorls in total. Suture shallowly impressed, subsutural ramp broad, strongly concave. Subsutural region with a row of distinct sigmoidal nodules that correspond in shape to upper parts of thickened growth lines, about 20 on first whorl, 17–19 on second–fourth whorls, 32 on penultimate and 30 on last whorl. Subsutural ramp smooth except for subsutural row of nodules, few thickened growth lines and 3–4 very weak spiral threads on last and penultimate whorls. Shoulder with pronounced thickened and rounded nodules, weakly opisthocline on upper whorls, but more strongly inclined and confluent with growth lines on last whorl; 17 on penultimate and antepenultimate whorls and 19 on last whorl. Spiral sculpture of 4–5 moderately pronounced, narrow cords on shoulder, seen both on nodules and in their intervals, and rather distinct, weakly rounded cords below shoulder, some flattened. Single cord appears on 3rd whorl, two cords on other whorls, including penultimate one. On last whorl cords occupy entire shell below shoulder, varying in width, their intervals 1–2 times width of cords, in total 48 cords on last whorl, of which 28 on canal. Shell base gradually narrowing towards long, narrow, nearly straight siphonal canal. Aperture narrow, constricted posteriorly, with very narrow and thin parietal callus, outer lip partially broken, with rounded angle at shoulder, weakly convex below shoulder and weakly concave at transition to canal. Anal sinus moderately deep, subsutural, broadly arcuate, confluent with large forward extension of outer lip. Shell off-white, with very light yellowish subsutural band and darker subsutural band on posterior half of last whorl, protoconch light tan. ANATOMY (n = 1, MNHN-IM-2009-13568). Head with long cylindrical tentacles, rounded on tips with closely spaced bases, eyes absent. Specimen male, with long penis with obliquely truncated tip. Seminal papilla occupies entire tip and is surrounded by circular fold. Proboscis conical, with broad base, but very narrow along most of its length. Proboscis retractors large, arranged in two bundles attached to proboscis base. Buccal mass very large and broad, nearly as long as proboscis, oesophagus forming short loop before nerve ring. Venom gland thick, medium-long and convoluted, opening dorsally into oesophagus just posterior to nerve ring. Muscular bulb very large. RADULA (n = 3; MNHN-IM-2007-17701, MNHN-IM-2007-17703, MNHN-IM-2009-13568). All specimens with similar radular morphology, radula comprising 38–40 rows of teeth, 11–17 nascent, short, length ca 1.8 mm (11% of AL without canal). Radula width up to 310 μm (1.9% of AL without canal). Central tooth with broad, subrectangular basal plate, anteriorly overlapped by preceding tooth, borders distinct, with narrow, sharply pointed cusp. Marginal teeth with strongly thickened posterior edges, folded longitudinally (Fig. 4E–F), in all details similar to those of Sibogasyrinx cf. pyramidalis 1. Remarks Largest specimen attains an SL of 57.4 mm. The other specimens are rather similar to the holotype in shell shape and sculpture. The entire subsutural ramp may be pale yellowish on the last whorl, but the subsutural zone of the posterior part of last whorl in larger specimens is always more darkly coloured than the rest of the shell. The subsutural ramp may be completely smooth or with indistinct spiral threads, the shoulder angulation on the last whorl is sometimes less distinct than in the holotype, as may be the cords below the shoulder on the penultimate whorl (up to 4). In shell outline, the new species resembles its congener Sibogasyrinx filosa, but differs in having much less distinct spiral sculpture on the subsutural ramp, as well as in its larger size (maximal SL 57.4 mm in S. subula sp. nov. and 52 mm in S. filosa) and radular morphology. The shell shape of the new species also strongly resembles that of some species of Leucosyrinx, particularly L. verrillii (Dall, 1881) (Fig. 13A–C) and to a lesser extent the smaller specimens of Leucosyrinx sp. B and Leucosyrinx sp. G (Fig. 13D and 13I, respectively). All these species of Leucosyrinx, however, have a different radular morphology, shorter siphonal canal and, according to the molecular tree (Fig. 1) and other published Conoidea phylogenies (Puillandre et al. 2011; Abdelkrim et al. 2018), belong to a different family. Distribution This species is known from the Solomon Islands, Papua New Guinea and Vanuatu, and has a relatively broad bathymetric range, from 440 to 814 m., Published as part of Kantor, Yuri I. & Puillandre, Nicolas, 2021, Rare, deep-water and similar: revision of Sibogasyrinx (Conoidea: Cochlespiridae), pp. 19-60 in European Journal of Taxonomy 773 on pages 35-38, DOI: 10.5852/ejt.2021.773.1509, http://zenodo.org/record/5536301, {"references":["Bouchet P., Kantor Y. I., Sysoev A. & Puillandre N. 2011. New operational classification of the Conoidea (Gastropoda). Journal of Molluscan Studies 77: 273 - 308. https: // doi. org / 10.1093 / mollus / eyr 017","Puillandre N., Kantor Y. I, Sysoev A., Couloux A., Meyer C., Rawlings T., Todd J. A. & Bouchet P. 2011. The dragon tamed? A molecular phylogeny of the Conoidea (Mollusca, Gastropoda). Journal of Molluscan Studies 77 (3): 259 - 272. https: // doi. org / 10.1093 / mollus / eyr 015","Abdelkrim J., Aznar-Cormano L., Fedosov A., Kantor Y. I., Lozouet P., Phuong M., Zaharias P. & Puillandre N. 2018. Exon-capture based phylogeny and diversification of the venomous gastropods (Neogastropoda, Conoidea). Molecular Biology and Evolution 35 (10): 2355 - 2374. https: // doi. org / 10.1093 / molbev / msy 144"]}

Sibogasyrinx pagodiformis sp. nov. (= PSH 5) urn:lsid:zoobank.org:act: DAA613D6-5AB6-4F89-A0AD-0C718A60307E Figs 8F, 12A–D Diagnosis Shell small, reaching 29 mm in length, pagodiform, with strongly concave subsutural ramp bearing two low spiral cords and distinct subsutural nodules. Axial sculpture of distinct opisthocline ribs extending to shell base, 10–11 per whorl. Spiral sculpture of closely spaced cords on and below shoulder. Radula lacking central tooth, marginal teeth trough-shaped with thickened edges. Etymology The name reflects the characteristic shell shape, resembling a pagoda. Material examined Holotype SOLOMON ISLANDS • NW of Malaita I.; 08°28′ S, 160°32′ E; depth 1250–1402 m; SALOMONBOA 3, stn CP2789; MNHN-IM-2009-16825. Other material (sequenced) VANUATU • 1 lv; E of Malekula I.; 16°00′ S, 167°38′ E; depth 1262–1285 m; SANTO 2006, stn AT139; MNHN-IM-2009-11327. Description MEASUREMENTS (holotype). SL 28.8 mm, AL (with canal) 15.2 mm, AL (without canal) 9.9 mm, SW 11.9 mm. SHELL (holotype). Moderately thick, except for fragile, partially chipped outer aperture lip; pagodiform, with moderately high spire and moderately long, narrow siphonal canal inclined to abaxially left. Protoconch missing (rendering exact an whorl count and protoconch measurements impossible), upper teleoconch whorls eroded. Teleoconch whorls rapidly increasing and strongly angled at shoulder, more than 8 remaining. Suture shallowly impressed, subsutural ramp broad, concave. A row of distinct, sharp, subsutural nodules on upper teleoconch whorls. Nodules less distinct on last three whorls and situated on short, slightly raised, prosocline axial ribs. Number of ribs on uppermost whorls indeterminate due to shell surface erosion, number of nodules per whorl increasing from 16–17 on first intact whorl to 22 on penultimate and 26 on last whorl. Subsutural ramp smooth except for nodules and two low, indistinct subsutural cords. Shoulder with pronounced, thickened, opisthocline axial ribs on all teleoconch whorls. Axial ribs on upper teleoconch whorls situated just above suture, extending to shell base on last whorl, number of ribs constant, 11–12 per whorl. Ribs intersected by closely spaced, rounded spiral cords, number increasing from 5 on uppermost remaining whorl to 10 on penultimate whorl. Last whorl with total of 38 cords, including 18 on canal. Cords narrow, their intervals 0.5–1.5 times cord width. Shell base rapidly narrowing towards narrow, moderately long siphonal canal. Aperture narrow, constricted posteriorly, with thin, moderately broad parietal callus, outer lip partially broken, convex on subsutural ramp, with rounded angle at shoulder, convex below shoulder and weakly concave at transition to canal. Anal sinus judging from growth lines shallow, subsutural, broadly arcuate. Shell flesh-coloured, with darker base, glossy. Periostracum thin, persisting in some cord intervals. ANATOMY (n = 1; MNHN-IM-2009-11327). Female. Proboscis relatively short, straight, cylindrical. Proboscis retractors large, in two bundles attached at proboscis base. Buccal mass within proboscis, rather long, about half length of proboscis, radular sac opening on right. Venom gland very thick, not long, opening with very short constriction just posterior to nerve ring. Venom bulb moderately large, elongate. RADULA (n = 2; MNHN-IM-2009-16825, AL 9.9 mm, IM-2009-11327, AL ca 9 mm). Relatively short, comprising 20–24 rows of teeth, 8–9 nascent. Radula length 1.1–1.6 mm (12–16% of AL without canal), tooth length 150 μm (1.7% of AL without canal). Central tooth absent. Marginal teeth trough-shaped when mature (Fig. 8F), with slightly thickened edges and sharp pointed tips. Teeth initially forming as a flat, elongate-oval plate, its edges gradually thickening along radular membrane and then by longitudinal folding of each tooth on 10–11 th row, counting from rear. Remarks The second live collected specimen has a broken shell base. In sculpture pattern it is very similar to the holotype. The species is distinct from all congeners in having a relatively broad, pagodiform shell with a shorter canal. Conchologically the species is quite similar to Comitas paupera (Watson, 1881), a highly variable species of uncertain generic affinity, but differing in lacking distinct spiral cords on the subsutural ramp. The lectotype and paralectotypes of the latter were illustrated by Sysoev (1996: figs 42–48). Superficially, the new species also strongly resembles a miniature copy of Leucosyrinx luzonica (Powell, 1969) (see Kantor et al. 2018: fig. 21), but differs in the shape of the marginal radular teeth and foregut anatomy. Another species of Leucosyrinx from the Solomon Islands, Leucosyrinx sp. D, is similar to the new species in shell outline (Fig. 13F), differing in radular morphology. Distribution This species was found in the Solomon Islands and Vanuatu at 1250–1262 m., Published as part of Kantor, Yuri I. & Puillandre, Nicolas, 2021, Rare, deep-water and similar: revision of Sibogasyrinx (Conoidea: Cochlespiridae), pp. 19-60 in European Journal of Taxonomy 773 on pages 51-54, DOI: 10.5852/ejt.2021.773.1509, http://zenodo.org/record/5536301, {"references":["Sysoev A. V. 1996. Deep-sea conoidean gastropods collected by the John Murray Expedition, 1933 - 34. Bulletin of the Natural History Museum of London, Zoology 62 (1): 1 - 30. Available from https: // www. biodiversitylibrary. org / page / 41008506 [accessed 7 Sep. 2021].","Powell A. W. B. 1969. The family Turridae in the Indo-Pacific. Part 2. The subfamily Turriculinae. Indo- Pacific Mollusca 2 (10): 215 - 416. Available from https: // www. biodiversitylibrary. org / page / 49824050 [accessed 24 Aug. 2021].","Kantor Y. I., Fedosov A. E. & Puillandre N. 2018. New and unusual deep-water Conoidea revised with shell, radula and DNA characters. Ruthenica, Russian Malacological Journal 28: 47 - 82."]}

Sibogasyrinx cf. pyramidalis (Schepman, 1913) 2 (= PSH 1) Figs 3J–Q, 4C–D, 5 Material examined (all sequenced) PAPUA NEW GUINEA • 2 lv; off Manus I.; 02°16′ S, 147°29′ E; depth 679–685 m; BIOPAPUA, stn CP3689; MNHN-IM-2009-16972, MNHN-IM-2009-16973 • 1 lv; N of Rabaul; 04°04′ S, 151°56′ E; depth 585–601 m; BIOPAPUA, stn CP3671; MNHN-IM-2009-16988 • 1 lv; N of Rabaul; 04°04′ S, 151°50′ E; depth 702–724 m; BIOPAPUA, stn CP3672; MNHN-IM-2009-17000 • 1 lv; Bismarck Sea, N of Long I.; 05°11′ S, 147°03′ E; depth 688 m; PAPUA NIUGINI, stn CP3981; MNHN-IM-2013-19716 • 1 lv; Bismarck Sea, N of Long I.; 05°10′ S, 147°03′ E; depth 724 m; PAPUA NIUGINI, stn CP3982; MNHN-IM-2013-19745 • 1 lv; Bismarck Sea, NW of Long I.; 05°12′ S, 146°59′ E; depth 470–508 m; PAPUA NIUGINI, stn DW3983; MNHN-IM-2013-19768 • 1 lv; New Ireland, 02°42′ S, 150°02′ E; depth 827–966 m; KAVIENG 2014, stn CP4483; MNHN-IM-2013-58845. Description MEASUREMENTS (largest specimen). SL 65.2 mm, AL (with canal) 34.7 mm, AL without canal 22.9 mm, SW 18.0 mm. SHELL. Extremely similar to that of Sibogasyrinx cf. pyramidalis 1. The only slight difference is the larger number of spiral cords crossing the shoulder nodules. Penultimate whorl of S. cf. pyramidalis 1 with 5–6 cords (increasing with SL), 6–8 in S. cf. pyramidalis 2. ANATOMY (n = 2, MNHN-IM-2009-16972, MNHN-IM-2009-17000). Head with long, cylindrical tentacles, rounded on tips with closely spaced bases, eyes absent.One specimen(MNHN-IM-2009-17000) male, with long penis with obliquely truncated tip. Seminal papilla occupies entire tip and is surrounded by circular fold. Proboscis long, coiled, with broad conical base, occupied by large bulbous buccal mass, spanning posterior ¼ of proboscis and protruding backwards from proboscis. Proboscis sheath with very thin transparent walls. Remaining anterior part of proboscis narrow, gradually diminishing in diameter towards its tip. Proboscis retractors well defined, powerful and numerous, attached to posteriormost edge of rhynchodeum. Radular sac and short oval odontophore situated on right side of buccal mass, posterior to rear of proboscis. Oesophagus adjoining buccal mass slightly constricted and forming a moderately long loop before passing through nerve ring. Salivary glands irregularly shaped, covering dorsal portion of nerve ring. Accessory salivary gland not traced. Venom gland thick, long, strongly convoluted, opens dorsally into oesophagus, immediately posterior to nerve ring (Fig. 5B). Muscular bulb oval, of moderate size, situated to the right of oesophagus. RADULA (n = 2; MNHN-IM-2009-16972, MNHN-IM-2009-17000). Comprising approximately 40 rows of teeth, 15 nascent, short, length (measured only in MNHN-IM-2009-16972) ca 1.7 mm (14% of AL without canal). Radula width up to 300 μm (2.5% of AL without canal). Shape of teeth very similar to that of Sibogasyrinx cf. pyramidalis 1, central tooth has less rounded posterior margin and in one specimen (MNHN-IM-2009-17000, Fig. 4D) the cusp is obtuse. Remarks Despite the similarities in shell and radular morphology, Sibogasyrinx cf. pyramidalis 1 and Sibogasyrinx cf. pyramidalis 2 are clearly distinct from a molecular perspective. One morphological character of the foregut that separates the two species is the presence of an accessory salivary gland in the former. Distribution The confirmed distribution is Papua New Guinea, at 585– 827 m., Published as part of Kantor, Yuri I. & Puillandre, Nicolas, 2021, Rare, deep-water and similar: revision of Sibogasyrinx (Conoidea: Cochlespiridae), pp. 19-60 in European Journal of Taxonomy 773 on pages 32-35, DOI: 10.5852/ejt.2021.773.1509, http://zenodo.org/record/5536301, {"references":["Schepman M. M. 1913. The Prosobranchia of the Siboga Expedition. Part 5. Toxoglossa. Siboga-Expeditie 49 (5): 365 - 452. Available from https: // www. biodiversitylibrary. org / page / 34450646 [accessed 24 Aug. 2021].","Powell A. W. B. 1969. The family Turridae in the Indo-Pacific. Part 2. The subfamily Turriculinae. Indo- Pacific Mollusca 2 (10): 215 - 416. Available from https: // www. biodiversitylibrary. org / page / 49824050 [accessed 24 Aug. 2021]."]}

Sibogasyrinx elbakyanae Kantor, Puillandre & Bouchet sp. nov. (= PSH 4) urn:lsid:zoobank.org:act: F9B671D9-4F84-4DF4-BAA0-3EF30AF06512 Fig. 12E–H Diagnosis Shell small, reaching 27.5 mm in length, fusiform, subsutural ramp strongly concave with distinct, broadly-spaced spiral cords and dense subsutural nodules. Axial sculpture of distinct opisthocline ribs extending to shell periphery. Spiral sculpture of distinct cords on and below shoulder. Etymology Philippe Bouchet joins us in dedicating this species to honor Alexandra Elbakyan, creator of the famous sci-hub portal. Material examined Holotype SOLOMON ISLANDS • S of Malaita I.; 09°55′ S, 161°33′ E; depth 1136–1750 m; SALOMONBOA 3, stn CP2817; MNHN-IM-2009-16834. Description (holotype) MEASUREMENTS. SL 27.3 mm, AL (with canal) 15.0 mm, AL (without canal) 9.6 mm, SW 8.9 mm. SHELL. Moderately thick, except for fragile outer aperture lip, partially chipped; fusiform, with high spire and long, narrow, straight siphonal canal. Protoconch eroded and partially decollated, thus rendering whorl count and measurements impossible. Teleoconch whorls strongly angled at shoulder, about 8 whorls in total. Suture deeply impressed, subsutural ramp moderately broad, strongly concave. Subsutural region with a row of dense, distinct, sigmoidal axial ribs that correspond in shape to upper parts of thickened growth lines and form small rounded nodules at intersections with spiral cords. Rib counts on uppermost whorls not possible due to shell surface erosion, about 24 on antepenultimate whorl, 27 on penultimate and 34 on last whorl. Subsutural ramp with thin but distinct spiral cords, 3 on first preserved teleoconch whorl and 5 on last three whorls, one immediately subsutural followed by another with interval about three times cord width, then three more closely spaced ones separated from second cord by an interval of about five times cord width. Upper two cords forming nodules at intersections with axial ribs. Shoulder with pronounced, thickened, opisthocline axial ribs on all teleoconch whorls, about 12 on upper whorls, including penultimate one and 14 on last whorl. Ribs extending to abapical suture on spire whorls, intersected by spiral cords, uppermost 2–3 very thin and closely spaced followed by 5–8 more broadly spaced. Last whorl with about 35 cords on shoulder and below, including 15 on canal. Cords narrow and separated by intervals 1–3 times the width of cords on axial ribs on shoulder, more broadly spaced on shell base and canal. Shell base gradually narrowing towards narrow, moderately long, nearly straight siphonal canal. Aperture narrow, constricted posteriorly, with moderately broad, thin parietal callus, outer lip partially broken, with rounded angle at shoulder, weakly convex below shoulder and weakly concave at transition to canal. Anal sinus shallow, subsutural, broadly arcuate. Shell base off-white, upper part of last whorl and teleoconch whorls very light yellow. Periostracum thin, persisting in intervals between cords and ribs. ANATOMY. Adult male, penis long, gradually narrowing towards tip, obliquely truncated at tip with a small but rather long conical papilla, surrounded by circular fold. Proboscis not long, conical, proboscis retractors not defined, entire posterior part of proboscis base muscular. Salivary glands fused, small. Single oval accessory salivary gland present at level of anterior part of proboscis. Venom gland large, thick and strongly convoluted, very constricted before opening into oesophagus within nerve ring. Muscular bulb moderately large. RADULA. Minute, comprising about 15 rows of teeth. Radula length 650 μm (6.8% of AL without canal), width up to 100 μm (1% of AL without canal). Central tooth present, with long cusp. Marginal teeth trough-shaped. Due to poor preparation it was impossible to study tooth morphology in detail. Remarks This species is most similar to Sibogasyrinx filosa, but differs in having a more stout shell with a relatively shorter canal, fewer, more distinct axial ribs on the shoulder (35 vs 46 on last whorl), and thinner, more widely spaced spiral cords. In shell outline the new species resembles some species of Leucosyrinx, particularly Leucosyrinx verrillii (Dall, 1881) (Fig. 13A–C), but differs in its smaller size, less numerous shoulder nodules and radular morphology. Distribution Known only from the type locality.