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Rhyacophila macedonica Karaouzas, Valladolid & Ibrahimi (n. sp.) This species has been found in continental Greece, North Macedonia, Kosovo and Serbia. Etymology. Named after the Greek region Macedonia, where the new species was first collected and identified. Type material. Holotype ♂: GREECE. Serres, Elaionas (41.142N, 23.566E, 556 m a.s.l.), 20/ix/2018 (I. Karaouzas) (KPC) (GenBank accession number MZ 645045). Paratypes: 5 ♂♂ P + 2 ♀♀ P + 2 IP + 1L, same locality as holotype, 20/ix/2018 (I. Karaouzas) (KPC). 2 ♂♂ P + 1 ♀ P + 1L, Serres, Agios Ioannis, 01/v/2013 (I. Karaouzas) [1 ♂ P MNCN _ Ent 296080 (MNCN); 1 ♂ P + 1 ♀ P (KPC); 1L MNCN _Ent 296081 (MNCN)]; 1 ♂ P + 1 IP + 1 L, 21/ix/2018 (I. Karaouzas) (KPC). 2 ♂♂ P + 4 ♀♀ P + 2 IP + 3L, Drama, 21/ix/2018 (I. Karaouzas) [1 ♀ P MNCN _ Ent 296082 (MNCN); 2 ♂♂ P + 4 ♀♀ P + 2 IP + 3 L (KPC)]. REPUBLIC OF NORTH MACEDONIA. 1 ♂, Banjaska Reka river, above Banjane village, 11/viii/2018 (J. Hinić & V. Slavevska Stamenković) [NMK _ SCG005, (MNCI)]. KOSOVO. 1 ♂, Mjak, 17/v/2017 (A. Bilalli & M. Musliu) (KTC). 1 ♂ + 2 ♀♀, Samakovë, 21/v/2017 (H. Ibrahimi) [1♀ MNCN _ Ent 296085 (MNCN); 1 ♂ + 1 ♀ (KTC)]. 3 ♂♂, Lugu i Kopilaqës, 3/xi/2019, (H. Ibrahimi) [1 ♂ MNCN _ Ent 296083 (MNCN); 2 ♂♂ (KTC)]. 1 ♂ + 1 ♀, Malishevë, Gjilan, 16/vii/2020, (H. Ibrahimi) (GJTC). 4 ♂♂, Kamenicë, 19/ix/2020 (A. Bilalli & H. Ibrahimi) (GOTC). 2 ♂♂, Kamenicë municipality, 19/ix/2020 (H. Ibrahimi) (GOTC). 2 ♂, Shushtë, 17/vii/2017 (A. Bilalli, H. Ibrahimi & M. Musliu) [1 ♂ MNCN _ Ent 296084 (MNCN); 1 ♂ (KTC)]. 1 ♂, Dubovë, 20/v/2020 (H. Ibrahimi) (BNTC). SERBIA. 1♂ + 1 ♀, Jastrebac, 25/ix/2020 (A. Bilalli & M. Musliu) (STC). Description of the larva (Figs 2–6, 12.) Biometrics of last instar larva and prepupa: Length: 12.02–18.41 mm ( = 14.61, n = 7). Maximum width of larva at metathorax: 2.53–3.38 mm ( = 3.04, n = 7). Specimens preserved in ethanol reddish brown, dorsum dark, on thorax two pale dorsolateral areas; on abdomen, two stripes of anterolateral circular pale spots, central stripe of irregular pale spots. Sclerotized areas of head and thorax pale with dark areas and spots. Head (Figs 2, 12a). Length: 1.73–1.95 mm ( = 1.82, n = 7). Width: 1.11–1.28 mm ( = 1.21, n = 7). Maximum width at posterior third of head, lateral margins slightly converging anteriorly, rounded posteriorly. Cephalic capsule (Figs 2a–2e), dorsally and dorsolaterally light brown, except anterior third, areas adjacent to frontoclypeal and anterior coronal sutures and anterior dorsolateral areas paler (Fig. 2a), in lighter specimens (Fig. 2c) pale in general, only posterior frontoclypeus and small spotted areas brown; ventral side pale; anteroventral edge and submentum reddish brown, posterior edge black or brown with black areas; laterally (Fig. 2b) with dark band in posterodorsal half with darker spots, triangular brown area with dark spots in middle, in lighter specimens (Fig. 2d) with only triangular brown area. Frontoclypeal apotome (Fig. 2e) pale; pair of oval pale brown submesal muscle attachment spots in anterior third, between setae #4; dark pigmentation in posterior third somewhat heart-shaped, with truncate anterior edges, central area paler brown, pointed anteriorly, with 4–6 oval darkly bordered brown muscle attachment spots, dark area including pair of pits #4 but not setae #6. Two-thirds of posterior dorsal area of head light brown, or pale in lighter specimens, marked with darker muscle attachment spots; at midlength one pair of light brown patches, wing-shaped, outside and paralleling posterior edge of frontal suture, in anterior extension, 2 or 3 spots parallel to suture, in middle of each patch two rows of spots perpendicular to suture; light circular area on juncture of frontal and coronal sutures, 3 or 4 pairs of spots aligned on both sides of coronal suture, two groups of spots on each side of posterior area. Three pair of pale spots around bases of setae #15, #16, and # 19–21 in darker specimens, one pale stripe in lighter specimens. Dorsal head pits with dark borders. Posterior occipital foramen black at midheight, dark brown surrounding remainder of foramen. Mandibles (Figs 3D, 3V) asymmetrical, as in other rhyacophilids; left mandible slightly larger than right one. Inner blade at base of left mandible slightly convex; right mandible inner blade at base straight with small tooth in middle. Maxillolabium and labrum as in other Rhyacophila species. Labrum with small transparent area in middle of anterior edge. Thorax (Figs 4, 12b): Anterior half of pronotum pale or very light coloured. Sinuous posterior margin bordered completely by dark band, reddish brown anterior of dark band at midheight of each half. Posterior half of pronotum (Fig. 4a) with three dark areas separated by light zones; (1) central dark region tapered posteriorly and reaching posterior edge of sclerite, anteriorly truncate near midlength of pronotum, wider than posteriorly, lateral edges concave, pair of submesal lines of dark brown spots inside this central dark region reaching posterior edge, several dark brown spots along posterior edge; (2) pair of lateral dark areas almost triangular, each with one long edge parallel to and as long as central dark region, one parallel to and slightly longer than posterior reddish brown region, and third edge diagonal between anterior subdorsal edge and ventral subposterior edge; (3) pair of circular spots anterolateral of anterior corners of triangular lateral dark areas. Each lateral dark area with two to three transverse rows of brown circular-oval spots with dark borders parallel to posterior border; four–six brown spots around setae #5, oval spot near diagonal edge of triangular dark area and curved row of pale dots from setae #5 extending toward anterolateral corner of pronotum, curving around circular spot (spot #3) on anterior half in darker specimens (Figs 4a, 4b); each side with narrow black band in anterolateral angle, sometimes extending posterad as fine brown band laterally to posterolateral margins; anterior margin light brown, with several brown spots near anterior angle (Fig. 12b). Legs: Similar to those of other Rhyacophila species. Colour yellowish, proximal regions of coxae, femora and distal regions of trochanters of fore- and midlegs black or dark brown, tibiae and tarsi light reddish; each inner and outer face of each femur with circular spots. Forelegs each ventrally with row of spicules, extending along from distal half of trochanter, and proximal half of femur, distal half of tibia and length of tarsus. Setae black or dark brown, some transparent in ventral area. Abdomen (Figs 5, 6, 12c, 12d): Similar to that of other species of Rhyacophila. In preserved larvae reddish brown dorsally, two stripes of anterolateral circular pale spots, central stripe of irregular pale spots, star shaped, ventral side pale. Widths of anterior abdominal segments similar, narrower in posterior segments. Lateral abdominal gills in tufts of 30–35 fine filaments. Dorsal sclerite of segment IX (Figs 5, 12c) light coloured, with continuous dark band on anterior edge and pair of discontinuous black bands on posterior edge; band of anterior edge black in lateral areas, brown in middle, with conspicuous pits in both sides; in darker specimens with semicircular coloured area in anterior middle; some dark spots near black bands on posterior edge, semicircular hyaline zone in middle posterior edge. Anal prolegs typical of group, each with long sword process (Figs 6a, 6b, 12d). Anal claws each with three or four teeth on ventral edge (Figs 6a, 6c), basoventral hook black basally and reddish brown apically (Figs 6b, 12d)., Published as part of Valladolid, Maria, Karaouzas, Ioannis, Ibrahimi, Halil, Arauzo, Mercedes, Stamenković, Valentina Slavevska, Dorda, Beatriz A., Hinić, Jelena, Bilalli, Astrit, Musliu, Milaim & Rey, Isabel, 2022, The Rhyacophila fasciata Group in Europe: Rhyacophila macedonica Karaouzas Valladolid & Ibrahimi (n. sp.) from Greece, Kosovo, Republic of North Macedonia and Serbia (Trichoptera: Rhyacophilidae), pp. 101-130 in Zootaxa 5125 (2) on pages 107-109, DOI: 10.11646/zootaxa.5125.2.1, http://zenodo.org/record/6424458

Rhyacophila fasciata Hagen 1859 Description of the imago. A complete description of the species is included in the publication by Valladolid et al. (2021). Here we describe small differences from the original description of R. fasciata found in male and female specimens from Kosovo. Male body length: 8.06–10.68 mm ( = 9.44, n = 22); each forewing: 10.11–12.52 ( = 11.32, n = 22); each hind wing: 8.94–10.91 mm ( = 9.87, n =22). Female body length: 8.92–12.99 mm ( = 10.75, n = 7); each forewing: 10.57–12.67 mm ( = 11.98, n = 7); each hind wing: 9.21–11.31 mm ( = 10.42, n = 7). Colour: Specimens preserved in ethanol reddish brown dorsally, yellowish ventrally, spurs reddish, setae golden brown, females generally darker than males; small black or dark brown spots in dorsal area; in lateral view, some specimens with large black spots marking border between dorsal and ventral areas. Forewings (Figs 8a, b) dark with small pale spots, two short transversal dark bands (fasciae) in Kosovo specimens (Fig. 8b), pale in preserved specimens, darker in females, hind wings pale (Fig. 8c). In specimens from Kosovo (Fig. 8b), forewings each with dark thick pterostigma area on leading edge between apices of subcostal (Sc) and first radial (R 1) or second radial (R 2) veins, each hind wing with pale thick pterostigma area between apices of subcostal (Sc) and second radial (R 2) veins, but not reaching second radial veins in some specimens; each forewing hyaline in irregular or triangular zone connecting medial (M) and medio-cubital (m-cu) veins and in distal area (arculus) of second cubital vein (Cu 2); in each hind wing (Fig. 8c) proximal segment of medial veins and connection area (M 1-4) not conspicuous, second cubital (Cu 2) veins not conspicuous proximally, proximal portion of cubital vein (Cu) and distal portions of first cubital vein (Cu 1a1b) thicker. Male genitalia (Fig. 15): Apical segment of each inferior appendage in lateral view (Fig. 15a; Valladolid et al. 2021, fig. 12A) with basal and distal edges diverging, dorsal and ventral edges diverging, posterior edge slightly concave dorsally and slightly convex ventrally, ventral edge slightly concave; apicodorsal vertex slightly angular (~117°), apicoventral angle rounded, projecting as thick lobe tapering to apex. Parameres in ventral view curved posteromesad in apical half (Fig. 15c; Valladolid et al. 2021, fig. 12BV). In lateral view (Fig. 15d; Valladolid et al. 2021, fig. 12BL) each constricted at short distance beyond base, dilated at middle (~38% as thick as length of paramere), with almost parallel posterodorsal and anteroventral margins in central area (S-shape), two files of long and thick spines on midventral margin, pointing outward (some Kosovo specimens with two files of sparse spines); midlateral surface covered by sparse fine spicules or setae, from middle anteroventral to posterodorsal edges and covering most of apex, absent in middle anteroventral and dorsal edges. Aedeagus (phallicata) in lateral view (Valladolid et al. 2021, fig. 12 CL) with anterodorsal margin concave and posterior corner of this concavity hooked anterad, posterior edge of aedeagus straight, apicoventral tip projected posterodorsad, subapically concave dorsally and convex ventrally.Ventral lobe of aedeagus triangular in ventral view (Fig. 15c; Valladolid et al. 2021, fig. 12BV), lateral edges slightly convex; lateroventral lobes of phallus straight, posterior edges strongly convex, subapicolateral margins slightly convex (Valladolid et al. 2021, fig. 12CD). Apicodorsal lobe of segment IX slightly dilated subapicolaterally, almost round in dorsal view (Fig. 15b; Valladolid et al. 2021, fig. 12DD), with apicomesal excision in some specimens; preanal appendages round, shorter than apicodorsal lobe. In ventral view (Valladolid et al. 2021, fig. 12DV), apical band V-shaped, nearly as wide as long, non-sclerotized inside area with posteromesal excision, posterior edges rounded, anal sclerites triangular. Female genitalia (Fig. 17): In lateral view (Fig. 17L, Valladolid et al. 2021, figs 13La; 13Lb-1, -2, -3), posterodorsal margin of segment VIII valves on each side irregular (in Kosovo specimens concave), with subdorsal projection, convex on ventral edge, posterolateral margin irregular, with 1 or 2 posterior projections, ventral projection more conspicuous, in some specimens slightly indented posteriorly, in Kosovo specimens only one ventral projection, rounded apically. In dorsal view (Fig. 17D; Valladolid et al. 2021, fig. 13Da), with indentation between segment VIII valves, and rounded apical and subapical corners, delimiting oval space, small projection in apical corner in some specimens. In ventral view (Fig. 17V; Valladolid et al. 2021, fig. 13V) segment VIII valves forming two elongate sclerites, narrowly separate basally, basal 1/4 of mesal edges parallel, distal 3/4 of mesal edges irregular, slightly concave, diverging; intersegmental membrane with pair of dark, ovoid or slightly elongate sclerites, ovoid in Kosovo specimens., Published as part of Valladolid, Maria, Karaouzas, Ioannis, Ibrahimi, Halil, Arauzo, Mercedes, Stamenković, Valentina Slavevska, Dorda, Beatriz A., Hinić, Jelena, Bilalli, Astrit, Musliu, Milaim & Rey, Isabel, 2022, The Rhyacophila fasciata Group in Europe: Rhyacophila macedonica Karaouzas Valladolid & Ibrahimi (n. sp.) from Greece, Kosovo, Republic of North Macedonia and Serbia (Trichoptera: Rhyacophilidae), pp. 101-130 in Zootaxa 5125 (2) on pages 106-107, DOI: 10.11646/zootaxa.5125.2.1, http://zenodo.org/record/6424458, {"references":["Hagen, H. (1859) Die Phryganiden Pictet's. Entomologische Zeitung, 20 (4 - 6), 131 - 170. Available from: https: // www. biodiversitylibrary. org / item / 35933 page / 141 / mode / 1 up (accessed 2 August 2021)","Valladolid, M., Arauzo, M., Chertoprud, M., Chvojka, P., Czachorowski, S., Dorda, B. A, Hinic, J., Ibrahimi, H., Karaouzas, I., Krpac, V., Kucinic, M., Lodovici, O., Salokannel, J., Slavevska Stamenkovic, V., Stojanovic, K., Wallace, I. & Rey, I. (2021) The Rhyacophila fasciata Group in Europe: Rhyacophila fasciata Hagen 1859 and formerly synonymized species (Trichoptera: Rhyacophilidae), with new description of Rhyacophila fasciata and Rhyacophila septentrionis McLachlan 1865 (stat. prom.). Zootaxa, 4975 (1), 1 - 57. https: // doi. org / 10.11646 / zootaxa. 4975.1.1"]}

Rhyacophila fasciata viteceki Valladolid & Kučinić (ssp. nov.) This subspecies has been found in Bosnia and Herzegovina. Etymology. The specific name is the genitive of Vitecek, given in honour of Dr. Simon Vitecek, biologist and trichopterologist of Wasser Cluster Lunz, Lunz am See, Austria. Type material. Holotype ♂: BOSNIA AND HERZEGOVINA, river Neretva, Bačevići (43.254154º N 17.824627º E, 35 m a.s.l.), 30/vi/17 (Stanić-Koštroman & M. Kučinić) [N1, MNCN _ Ent 269367 (MNCN)]. Paratypes: 2 ♂, same locality as holotype, 30/vi/17 (Stanić-Koštroman & M. Kučinić) [N2, no. 4455, (MCK) and N3, no. 1111 (UMBH)]. 2 ♂ and 1 ♀, river Sturba, 30/viii/18 (M. Kučinić) [1 ♂, S1, MNCN _ Ent 269368 (MNCN); 1 ♀, S3, MNCN _ Ent 269369 (MNCN); 1 ♂ S2, no. 4456 (MKC)]. 1 ♂, river Bunica, 04/vi/17 (StanićKoštroman & M. Kučinić) [B1, no. 1112 (UMBH)]. Description of the imago. Holotype (N1): Length from front of head to distal edge of segment IX 8.91 mm, each forewing 12.55 mm, each hind wing 10.54 mm. Males: Length 8.29–9.21 mm (x = 8.72, n = 6), each forewing 9.83–12.55 (x = 11.34, n = 6), each hind wing 8.85–10.54 mm (x = 9.79, n = 6). Female: Length from front of head to distal edge of segment VIII 11.26 mm (n =1), forewing 14.52 mm (n = 1), hind wing 12.15 mm (n = 1). In ethanol-preserved specimens colour generally pale brown, yellowish, with golden brown setae. Head, thorax, and abdomen dorsally pale brown, with small dark brown spots in dorsal area, some specimens in lateral view with long black spots marking borders between dorsal and ventral areas, abdomen ventrally yellowish; female darker than male, anterior 2/3 of abdominal segments pale brown or brown in darker specimens. Legs light, with spurs reddish brown. Forewings brown, spotted; hind wings pale. Male genitalia (Figs 3, 6 a–6e): Apical segment of each inferior appendage (Figs 3A, 6a) with basal and distal edges diverging, posterior edge oblique and slightly convex, ventral edge straight to slightly concave, at least two times longer that dorsal edge. Apicodorsal vertex angular, apicoventral angle projecting as thick lobe narrowing progressively to round apex (Fig 6a). Parameres (Figs 3 BV–3BL, 6c–6d) in ventral view curved posteromesad in apical 2/3 (Figs. 3 BV p, 6c). In lateral view (Figs. 3 BL, 6d) each slender at base, slightly dilated in middle, with rounded dorsal margin before midlength and almost straight ventral margin at midlength, pointed at apex; row of few thick spines on midventral margin (Figs 6 c–6d); midlateral surface covered by very fine spicules or setae, reaching from anteroventral to posterodorsal edges of paramere, absent from dorsal edge. Aedeagus (phallicata) in lateral view (Figs 3 CL, 6e) with dorsal margin deeply concave and posterior corner of concavity hooked anterad; upper posterior edge concave to obliquely truncate apex, ventral edge sinous. Lateroventral lobes of phallus (Fig. 3 CD lvl) convex, narrowing progressively towards apex, apicolateral margins round, posteromesal edge of each lobe concave. Ventral lobe of aedeagus nearly round, tapering and acute caudally (Fig 3 BV vl). Apicodorsal lobe of segment IX (Figs 3 DD, 6b) slightly dilated subapicolaterally, posterior edge semicircular, lateral edges converging anterad; preanal appendages (fig 3DD pa) shorter than apicodorsal lobe (Fig 3 DD al), obliquely subtruncate apicolaterally, with slightly concave posteromesal edges (Figs. 3 DD, 6b). In ventral view, apical band V-shaped, inner and outer edges converging in posterior third, pointed apically, longer than wide (Fig 3 DV ab); posterior edge of non-sclerotized ventral area slightly concave, with small mesal incision (Fig 3 DV va), anal sclerites subtriangular, concave apically (Fig 3 DV as)., Published as part of Valladolid, María, Kučinić, Mladen, Arauzo, Mercedes, Cerjanec, Darko, Ćuk, Renata, Dorda, Beatriz A., Lodovici, Omar, Stanić-Koštroman, Svjetlana, Vučković, Ivan & Rey, Isabel, 2020, The Rhyacophila fasciata Group in Croatia and Bosnia and Herzegovina: Rhyacophila f. fasciata Hagen 1859 and the description of two new subspecies, Rhyacophila fasciata delici Kučinić & Valladolid (ssp. nov.) from Croatia and Bosnia and Herzegovina and Rhyacophila fasciata viteceki Valladolid & Kučinić, pp. 51-75 in Zootaxa 4885 (1) on pages 65-66, DOI: 10.11646/zootaxa.4885.1.3, http://zenodo.org/record/4296372