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Appendix of: Vigne J.-D., Briois F., Guilaine J. (eds.), « Klimonas, an Early Pre-Pottery Neolithic village in Cyprus », Gallia Préhistoire & Paléorient, International Supplement 1, CNRS Éditions, Paris, 2023., Annexe de : Vigne J.-D., Briois F., Guilaine J. (dir.), « Klimonas, un village néolithique pré-céramique ancien à Chypre », Gallia Préhistoire & Paléorient, Supplément international 1,CNRS Éditions, Paris, 2023.

Appendix of: Vigne J.-D., Briois F., Guilaine J. (eds.), « Klimonas, an Early Pre-Pottery Neolithic village in Cyprus », Gallia Préhistoire & Paléorient, International Supplement 1, CNRS Éditions, Paris, 2023., Annexe de : Vigne J.-D., Briois F., Guilaine J. (dir.), « Klimonas, un village néolithique pré-céramique ancien à Chypre », Gallia Préhistoire & Paléorient, Supplément international 1,CNRS Éditions, Paris, 2023.

Traduction en Grec par Maria Rousou du chapitre 33 intitulé : Le village néolithique de Klimonas. Aspects techniques, sociaux et symboliques ; insularité et connectivité, signé par J.-D. Vigne, J. Guilaine, R. Hadad, L. Astruc, F. Briois, T. Cucchi, L. Gourichon, P. Mylona, S. Rigaud, M. Rousou de l'ouvrage suivant : Vigne J.-D., Briois F., Guilaine J. (dir.), « Klimonas, un village néolithique pré-céramique ancien à Chypre », Gallia Préhistoire, Supplément international 1, CNRS Éditions, Paris, 2023., Greek translation by Maria Rousou of Chapter 33 entitled: The Neolithic village of Klimonas: technical, social and symbolic aspects; insularity and connectivity co-authored by J.-D. Vigne, J. Guilaine, R. Hadad, L. Astruc, F. Briois, T. Cucchi, L. Gourichon, P. Mylona, S. Rigaud, M. Rousou of the following book: Vigne J.-D., Briois F., Guilaine J. (eds.), « Klimonas, an Early Pre-Pottery Neolithic village in Cyprus », Gallia Préhistoire, International Supplement 1, CNRS Éditions, Paris, 2023.

Annexe de : Vigne J.-D., Briois F., Guilaine J. (dir.), « Klimonas, un village néolithique pré-céramique ancien à Chypre », Gallia Préhistoire & Paléorient, Supplément international 1,CNRS Éditions, Paris, 2023., Appendix of: Vigne J.-D., Briois F., Guilaine J. (eds.), « Klimonas, an Early Pre-Pottery Neolithic village in Cyprus », Gallia Préhistoire & Paléorient, International Supplement 1, CNRS Éditions, Paris, 2023.

Includes bibliographical references. Number of sources in the bibliography: 977 Thesis (Ph. D.) -- University of Cyprus, Faculty of Letters, Department of History and Archaeology, 2022. The University of Cyprus Library holds the printed form of the thesis. Οι αρχαιολογικές έρευνες των τελευταίων δεκαετιών στην Κύπρο έφεραν στο φως εγκαταστάσεις κυνηγών/γεωργών, που ανάγονται στην 9η χιλιετία π.Χ. (Αρχική Προκεραμική Νεολιθική), τις οποίες διαδέχθηκαν θέσεις, στις οποίες το κυνήγι σταδιακά δίνει τη θέση του στην κτηνοτροφία, ενώ παράλληλα επιβεβαιώνονται γεωργικές πρακτικές (Πρώιμη Προκεραμική Νεολιθική). Η Ύστερη Προκεραμική Νεολιθική (7η-6η χιλιετία π.Χ.) στην Κύπρο χαρακτηρίζεται από την παρουσία ενός τοπικού νησιωτικού πολιτισμού (γνωστού και ως «Πολιτισμός της Χοιροκοιτίας»), ανεξάρτητου από ηπειρωτικές επιρροές, όπως υποδηλώνουν οικισμοί μερικές φορές μεγάλοι σε μέγεθος, που κατοικούνται από γεωργούς, κτηνοτρόφους και τεχνίτες. Αυτοί οι πρώιμοι οικισμοί εξελίχθηκαν και εκμεταλλεύτηκαν τα τοπικά οικοσυστήματα του νησιωτικού περιβάλλοντος, στο οποίο είχε παρέμβει ελάχιστα ο άνθρωπος. Η παρούσα μελέτη στοχεύει στον εντοπισμό των φυτικών πόρων και στην ανασύνθεση του παλαιοπεριβάλλοντος και της εξέλιξής του ανά τις χιλιετίες (9η-6η χιλιετία π.Χ.), όπως αυτό μεταβλήθηκε από την ανθρώπινη αλληλεπίδραση, μέσω της μελέτης των ανθρακολογικών καταλοίπων (κάρβουνων) που βρέθηκαν στις θέσεις Άγιος Τύχωνας-Κλήμωνας (8800 π.Χ.), Παρεκκλησσιά-Σιυλλουρόκαμπος (8500-6900 π.Χ.) και Χοιροκοιτία-Βούνοι (6800-5500 π.Χ.). Η ενδελεχής ανάλυση και των υπολειμμάτων καρπών άγριων ξυλωδών φυτών, ολοκληρώνει την ανθρακολογική μελέτη και επιτρέπει την προσέγγιση του ερωτήματος της στοχευμένης συλλογής καρπών. Οι συλλογές αναφοράς σύγχρονου βοτανικού υλικού που δημιουργήθηκαν στο πλαίσιο αυτής της έρευνας, επέτρεψαν την ταύτιση του πιο πάνω αρχαιοβοτανικού υλικού και οδήγησαν στον εμπλουτισμό των συλλογών του Εθνικού Μουσείου Φυσικής Ιστορίας στο Παρίσι και στη δημιουργία των πρώτων συλλογών αναφοράς στην Κύπρο (Πανεπιστήμιο Κύπρου). Επιπλέον, η μελέτη αυτή παρουσιάζει τη λεπτομερή μέθοδο ταυτοποίησης των καταλοίπων καρπών της Pistacia spp. Τα αποτελέσματα που προέκυψαν από την έρευνα δείχνουν την εκμετάλλευση από τους κατοίκους των οικισμών ενός ευρέος φάσματος τοπικών ξυλώδων φυτών στο άμεσο περιβάλλον των προαναφερθέντων θέσεων, το οποίο ήταν πιθανότατα πλουσιότερο και αρκετά διαφοροποιημένο σε ξυλώδη φυτά (μεσογειακή μακία, δασικοί σχηματισμοί) από ό,τι σήμερα. Επιπλέον, μας επιτρέπουν να σκιαγραφήσουμε την εξέλιξη της δυναμικής της βλάστησης σε μια περίοδο σχεδόν 3300 ετών και την σταδιακή υποβάθμισή της. Σημαντική αλλαγή παρατηρείται κατά την περίοδο της αναδιοργάνωσης του νεότερου οικισμού της Χοιροκοιτίας (επίπεδα Α, ΙΙΙ-Ι). Στο πλαίσιο αυτό, αξιολογείται ο ρόλος των φυσικών (κλιματικών) παραγόντων έναντι των ανθρωπογενών επιπτώσεων στους φυτικούς πληθυσμούς, με ιδιαίτερη έμφαση στο ρόλο των πυρκαγιών. Επίσης, η μελέτη αυτή προχωρεί σε κάποια συμπεράσματα για τη χρήση των ξυλωδών φυτών στην αρχιτεκτονική, ως καύσιμη ύλη, ζωοτροφή, για ιατρικές χρήσεις, καθώς και για την ανθρώπινη διατροφή. Επιπλέον, διερευνάται η βιογεωγραφία και η ιθαγένεια ορισμένων ξυλωδών φυτών, όπως η χαρουπιά και η αμυγδαλιά. Η παρούσα μελέτη συμπληρώνει προηγούμενες παλαιοπεριβαλλοντικές μελέτες και αναδεικνύει τη μακροπρόθεσμη αλληλεπίδραση ανθρώπου-περιβάλλοντος κατά την Προκεραμική Νεολιθική. Τέλος, δίνει νέες ερευνητικές κατευθύνσεις στην προσέγγιση της αλληλεπίδρασης ανθρώπου-περιβάλλοντος, την εκμετάλλευση, τη διαχείριση και τη χρήση των ξυλωδών φυτών στο νησιωτικό περιβάλλον της Κύπρου και στη Μεσόγειο γενικότερα. Archaeological research in Cyprus over the last decades has revealed the existence of settlements of hunters/early farmers since the 9th millennium BC (Initial Pre-Pottery Neolithic), followed by other sites, in which hunting progressively gives way to farming, and agricultural practices are confirmed (Early Pre-Pottery Neolithic). The Late Pre-Pottery Neolithic (7th-6th millennia BC) in Cyprus is marked by the development of a locally developed island culture (also known as the 'Khirokitia Culture'), independent of continental influences, as indicated by villages, sometimes large in size, populated by farmer-breeders and craftsmen. These early settlements in Cyprus developed and exploited the resources of the local ecosystems in an island environment previously little influenced by man. This study aims to localize the plant resources and to reconstruct the plant formations and evolution over time (9th-6th millennium BC), as resulted with human interaction, through the study of the anthracological remains (charcoal) recovered in the sites of Ayios Tychonas-Klimonas (8800 cal. BC), Parekklishia-Shillourokambos (8500-6900 cal. BC) and Khirokitia-Vounoi (6800-5500 cal. BC). A thorough analysis of the fruit remains targeting the exploitation of wild woody fruit taxa, completes the anthracological study and make possible to approach the question of purposeful fruit gathering. The reference collections of modern botanical material, established within the framework of this research, enabled the study of the above archaeobotanical material and led to the enrichment of the collections of the National Museum of Natural History in Paris, and to the creation of the first reference collections in Cyprus (University of Cyprus). This study also provides a detailed identification of the fruit remains of Pistacia spp. The results obtained, show the exploitation of a wide range of local woody resources by the inhabitants in the direct environment of the aforementioned sites, that was probably much richer and more diversified in woody taxa (Mediterranean maquis, forest formations) than today. Moreover, they allow us to outline the evolution of the vegetation dynamics over a period of nearly 3300 years and the degradation of the vegetation cover. A major change is observed at the time of the reorganisation of the recent village of Khirokitia (levels A, III-I). Within this framework, the role of natural (climatic) factors versus the anthropogenic impact on the plant communities is assessed, with a particular focus on the role of fire. This study also concludes for the use of woody resources for fuel, in architecture, for fodder, medical use, as well as in human diet. Besides, the biogeography and indigeneity of certain woody taxa, such as carob and almond are investigated. This study complements previous palaeoenvironmental studies and highlights human-environment interactions over the long term during the Pre-Pottery Neolithic. Finally, it opens up new research directions concerning human-environment interactions, exploitation, management, and use of woody plants on the island environment of Cyprus and in the Mediterranean in general. Les recherches archéologiques conduites récemment à Chypre ont mis en évidence la présence d’occupations humaines de chasseurs/agriculteurs depuis le 9ère millénaire av. n.-è. (Néolithique précéramique initial), suivies par des sites où la chasse laisse progressivement la place à l’élevage et où les pratiques agricoles se confirment (Néolithique précéramique ancien). Le Néolithique précéramique récent (7ème-6ème millénaires av. n.-è.) à Chypre est marqué par le développement d’une culture insulaire originale (dite aussi ‘Culture de Khirokitia’), désormais à l’écart des influences continentales, et s’exprimant au sein de villages, parfois de grande taille, peuplés d’agriculteurs-éleveurs et d’artisans. Ces premiers peuplements de Chypre ont évolué et exploité les ressources des écosystèmes locaux dans un environnement insulaire auparavant très peu influencé par l’Homme. Cette étude vise à reconstituer les aires d’approvisionnement des ressources végétales et proposer une reconstitution des formations végétales et leur évolution au cours du temps (9ème-6ème millénaire av. n.-è.), en interaction avec les activités humaines, à travers l’étude des vestiges anthracologiques (charbons de bois) collectés à Ayios Tychonas-Klimonas (8800 cal. BC), Parekklishia-Shillourokambos (8500-6900 cal. BC) et Khirokitia-Vounoi (6800-5500 cal. BC). Une étude carpologique approfondie ciblant plus particulièrement l’exploitation de taxons fruitiers ligneux sauvages, complète l’étude anthracologique et permet d’approcher la question de la cueillette des fruits. Les collections de référence de matériel botanique moderne, constituées dans le cadre de cette recherche ont facilité l’étude du matériel archéobotanique et ont abouti à l’enrichissement des collections du Muséum national d’Histoire naturelle et à la création des premières collections de référence archéobotaniques à l’Université de Chypre. Cette étude permet par ailleurs d’aborder en détail d’identification des restes fruitiers de Pistacia spp. Les résultats obtenus montrent l’exploitation d’un large éventail de ressources ligneuses locales, au sein d’environnements plus riches et diversifiés en taxons ligneux (maquis méditerranéen, formations forestières) que ceux observés actuellement autour des sites. De plus, ils permettent de reconstituer la dynamique de la végétation sur une période de près de 3300 ans en relevant des indices de dégradation du couvert végétal. Une rupture majeure observée au moment de la réorganisation du village récent de Khirokitia (niveaux A, III-I). Dans ce cadre, le rôle des facteurs naturels (climatiques) versus l’impact anthropique sur les communautés végétales est évalué, avec un focus particulier sur le rôle du feu. Cette étude explore aussi l’utilisation des ressources ligneuses à des fins de combustible, construction, fourrage, comme plantes médicinales, et dans l’alimentation humaine. De plus, nous examinons la question de la biogéographie et de l’indigénat de certains taxons ligneux, comme le caroubier et l’amandier. Notre travail permet de compléter les études paléo-environnementales antérieures et met en évidence les interactions homme-environnement sur la longue durée au cours du Néolithique Précéramique. Enfin, il ouvre des nouvelles directions de recherche concernant les interactions homme-environnement, l’exploitation, la gestion et l’utilisation des plantes ligneuses dans l’environnement insulaire de Chypre et dans la Méditerranée en général. Muséum National d’Histoire Naturelle

Οι αρχαιολογικές έρευνες των τελευταίων δεκαετιών στην Κύπρο έφεραν στο φως εγκαταστάσεις κυνηγών/γεωργών, που ανάγονται στην 9η χιλιετία π.Χ. (Αρχική Προκεραμική Νεολιθική), τις οποίες διαδέχθηκαν θέσεις, στις οποίες το κυνήγι σταδιακά δίνει τη θέση του στην κτηνοτροφία, ενώ παράλληλα επιβεβαιώνονται γεωργικές πρακτικές (Πρώιμη Προκεραμική Νεολιθική). Η Ύστερη Προκεραμική Νεολιθική (7η-6η χιλιετία π.Χ.) στην Κύπρο χαρακτηρίζεται από την παρουσία ενός τοπικού νησιωτικού πολιτισμού (γνωστού και ως «Πολιτισμός της Χοιροκοιτίας»), ανεξάρτητου από ηπειρωτικές επιρροές, όπως υποδηλώνουν οικισμοί μερικές φορές μεγάλοι σε μέγεθος, που κατοικούνται από γεωργούς, κτηνοτρόφους και τεχνίτες.Αυτοί οι πρώιμοι οικισμοί εξελίχθηκαν και εκμεταλλεύτηκαν τα τοπικά οικοσυστήματα του νησιωτικού περιβάλλοντος, στο οποίο είχε παρέμβει ελάχιστα ο άνθρωπος. Η παρούσα μελέτη στοχεύει στον εντοπισμό των φυτικών πόρων και στην ανασύνθεση του παλαιοπεριβάλλοντος και της εξέλιξής του ανά τις χιλιετίες (9η-6η χιλιετία π.Χ.), όπως αυτό μεταβλήθηκε από την ανθρώπινη αλληλεπίδραση, μέσω της μελέτης των ανθρακολογικών καταλοίπων (κάρβουνων) που βρέθηκαν στις θέσεις Άγιος Τύχωνας-Κλήμωνας (8800 π.Χ.), Παρεκκλησσιά-Σιυλλουρόκαμπος (8500-6900 π.Χ.) και Χοιροκοιτία-Βούνοι (6800-5500 π.Χ.). Η ενδελεχής ανάλυση και των υπολειμμάτων καρπών άγριων ξυλωδών φυτών, ολοκληρώνει την ανθρακολογική μελέτη και επιτρέπει την προσέγγιση του ερωτήματος της στοχευμένης συλλογής καρπών. Οι συλλογές αναφοράς σύγχρονου βοτανικού υλικού που δημιουργήθηκαν στο πλαίσιο αυτής της έρευνας, επέτρεψαν την ταύτιση του πιο πάνω αρχαιοβοτανικού υλικού και οδήγησαν στον εμπλουτισμό των συλλογών του Εθνικού Μουσείου Φυσικής Ιστορίας στο Παρίσι και στη δημιουργία των πρώτων συλλογών αναφοράς στην Κύπρο (Πανεπιστήμιο Κύπρου). Επιπλέον, η μελέτη αυτή παρουσιάζει τη λεπτομερή μέθοδο ταυτοποίησης των καταλοίπων καρπών της Pistacia spp.Τα αποτελέσματα που προέκυψαν από την έρευνα δείχνουν την εκμετάλλευση από τους κατοίκους των οικισμών ενός ευρέος φάσματος τοπικών ξυλώδων φυτών στο άμεσο περιβάλλον των προαναφερθέντων θέσεων, το οποίο ήταν πιθανότατα πλουσιότερο και αρκετά διαφοροποιημένο σε ξυλώδη φυτά (μεσογειακή μακία, δασικοί σχηματισμοί) από ό,τι σήμερα. Επιπλέον, μας επιτρέπουν να σκιαγραφήσουμε την εξέλιξη της δυναμικής της βλάστησης σε μια περίοδο σχεδόν 3300 ετών και την σταδιακή υποβάθμισή της. Σημαντική αλλαγή παρατηρείται κατά την περίοδο της αναδιοργάνωσης του νεότερου οικισμού της Χοιροκοιτίας (επίπεδα Α, ΙΙΙ-Ι). Στο πλαίσιο αυτό, αξιολογείται ο ρόλος των φυσικών (κλιματικών) παραγόντων έναντι των ανθρωπογενών επιπτώσεων στους φυτικούς πληθυσμούς, με ιδιαίτερη έμφαση στο ρόλο των πυρκαγιών. Επίσης, η μελέτη αυτή προχωρεί σε κάποια συμπεράσματα για τη χρήση των ξυλωδών φυτών στην αρχιτεκτονική, ως καύσιμη ύλη, ζωοτροφή, για ιατρικές χρήσεις, καθώς και για την ανθρώπινη διατροφή. Επιπλέον, διερευνάται η βιογεωγραφία και η ιθαγένεια ορισμένων ξυλωδών φυτών, όπως η χαρουπιά και η αμυγδαλιά.Η παρούσα μελέτη συμπληρώνει προηγούμενες παλαιοπεριβαλλοντικές μελέτες και αναδεικνύει τη μακροπρόθεσμη αλληλεπίδραση ανθρώπου-περιβάλλοντος κατά την Προκεραμική Νεολιθική. Τέλος, δίνει νέες ερευνητικές κατευθύνσεις στην προσέγγιση της αλληλεπίδρασης ανθρώπου-περιβάλλοντος, την εκμετάλλευση, τη διαχείριση και τη χρήση των ξυλωδών φυτών στο νησιωτικό περιβάλλον της Κύπρου και στη Μεσόγειο γενικότερα., Archaeological research in Cyprus over the last decades has revealed the presence of human settlements of hunter/early farmers since the 9th millennium BC (Initial Pre-Pottery Neolithic), followed by other sites, in which hunting progressively gives way to farming, and agricultural practices are confirmed (Early and Late Pre-Pottery Neolithic, 8th-6th mill. BC). This study aims to localize the plant resources and to reconstruct the plant formations and evolution over a period of nearly 3300 years, in interaction with human activities, through the study of anthracological remains (charcoal) recovered at the sites of Ayios Tychonas-Klimonas (8800 cal. BC), Parekklishia-Shillourokambos (8500-6900 cal. BC) and Khirokitia-Vounoi (6800-5500 cal. BC). A thorough analysis of the fruit remains targeting the exploitation of wild woody fruit taxa, completes the anthracological study and makes is possible to approach the question of fruit gathering. This study also provides a detailed identification of the fruit remains of Pistacia spp. The reference collections of modern botanical material, established within the framework of this research, enabled the study of the above archaeobotanical material and led to the enrichment of the collections of the National Museum of Natural History and to the creation of the first reference collections at the University of Cyprus. The results obtained show the exploitation of a wide range of local woody resources in the direct environment of the aforementioned sites, that was probably much richer and more diversified in woody taxa (Mediterranean maquis, forest formations) than today. Moreover, they allow us to assess the role of natural factors versus the anthropogenic impact on the degradation of the vegetation cover, with a particular focus on the role of fire. In addition, the use of woody resources for various purposes (fuel, architecture, fodder, human diet) is also explored. The biogeography and indigeneity of certain woody taxa (carob, almond) are also investigated. This work highlights the long-term human-environment interactions during the Pre-Pottery Neolithic and opens up new research directions concerning human-environment interactions, exploitation, management, and use of woody plants on the island environment of Cyprus and in the Mediterranean in general., Les recherches archéologiques conduites récemment à Chypre ont mis en évidence la présence d’occupations humaines de chasseurs/agriculteurs depuis le 9ème millénaire av. n.-è., suivies par d’autres sites où la chasse laisse progressivement la place à l’élevage et l’agriculture (Néolithique précéramique ancien et récent, 8ème-6ème millénaires av. n.-è.). Cette étude vise à reconstituer les aires d’approvisionnement des ressources végétales et proposer une reconstitution des formations végétales et leur évolution au cours du temps sur une période de près de 3300 ans, en interaction avec les activités humaines, à travers l’étude des vestiges anthracologiques (charbons de bois) collectés à Ayios Tychonas-Klimonas (8800 cal. BC), Parekklishia-Shillourokambos (8500-6900 cal. BC) et Khirokitia-Vounoi (6800-5500 cal. BC). Une étude carpologique approfondie ciblant l’exploitation de taxons fruitiers ligneux sauvages, complète l’étude anthracologique et permet d’approcher la question de la cueillette des fruits. Cette étude permet par ailleurs d’aborder en détail l’identification des restes fruitiers de Pistacia spp. Les collections de référence de matériel botanique moderne, constituées dans le cadre de cette recherche ont facilité l’étude du matériel archéobotanique et ont abouti à l’enrichissement des collections du Muséum national d’Histoire naturelle et à la création des premières collections de référence archéobotaniques à l’Université de Chypre. Les résultats obtenus montrent l’exploitation d’un large éventail de ressources ligneuses locales, au sein d’environnements riches et diversifiés en taxons ligneux (maquis méditerranéen, formations forestiers). De plus, ils permettent d’évaluer le rôle des facteurs naturels versus l’impact anthropique sur la dégradation du couvert végétal, avec un focus particulier sur le rôle du feu. Par ailleurs, le rôle des ressources ligneuses à des fins diverses (combustible, construction, alimentation humaine et animale) est aussi exploré. La question de la biogéographie et de l’indigénat de certains taxons ligneux (caroubier, amandier) est aussi examinée. Cette étude met en évidence les interactions homme-environnement sur la longue durée au cours du Néolithique Précéramique et ouvre des nouvelles directions de recherche concernant les interactions homme-environnement, l’exploitation, la gestion et l’utilisation des plantes ligneuses dans l’environnement insulaire de Chypre et dans la Méditerranée en général.

International audience

Coastal hypoxia is a serious environmental problem that needs to be addressed at a global level. The phenomenon of hypoxia is characterized by low Dissolved Oxygen (DO) levels in the water column that causes detrimental effects on aquatic organisms. Anthropogenic activities such as intensive agriculture practices and point-source nutrient loading are considered the main causes of coastal hypoxia. This study utilizes data-driven modelling based on Artificial Neural Networks (ANNs), and specifically Feed-Forward ANNs, to predict surface DO levels. Several surface water quality parameters such as water temperature, nitrogen species (ammonium, nitrite and nitrate), phosphorus, pH, salinity, electrical conductivity, and chlorophyll-a served as the ANN’s input parameters. These parameters were measured at several sampling sites in the coastal waters of Cyprus and some of the sites were located near areas with anthropogenic activities, during the period 2000-2021. An ANN with a 9-5-1 topology was developed and ANN managed to predict with good accuracy the DO levels, with the Coefficient of determination (r 2) as high as r 2=0.991 for the test set. Additionally, sensitivity analysis was performed to measure the impact of each input parameter on the DO level, and it was estimated that the water temperature is the most influential factor. The “Weights” sensitivity analysis algorithm was used for this purpose. The ANN-based method proposed can be used as a management tool for predicting the DO levels and prevention of hypoxia. Therefore, this work has a positive impact on marine sciences and marine information technology.

Laonice bahusiensis Söderström, 1920 (Figs 2, 3, 4, 5A, 18A, 19A, Table 3) Laonice bahusiensis Söderström, 1920: 4–7, 81–83, 93, 98, 99, 110, 114, 128, 134, 195, 223, figs 78–82. Maciolek 2000: 534- 535, table 1. Jelsing 2003: 244, figs 1H, 3E, F. Sikorski 2003 (Part.): 320–325, figs 3A–I, 4A, B, 5A, B, 6F. Jelsing & Eibye-Jacobsen 2010: 376–377, figs 1–4. Type locality. SWEDEN, Skagerrak, Gullmarfjord, Bohuslän, approximately 58.25°N, 11.00°E, Zool. Stn. 1893. Type material. SMNH 4637 (lectotype), 4636, 4638 (11 paralectotypes); ZMMU Pl 973 (2 paralectotypes); UUZM 153 c, d, f, g (70 paralectotypes). Adult morphology. Up to 60 mm long, 1.2 mm wide for 120 chaetigers. Lectotype complete specimen in three fragments, totally 41 mm long, 1.0 mm wide for 107 chaetigers. Pigmentation on body and palps absent. Prostomium triangular, anteriorly wide, usually broadly rounded, occasionally truncate to slightly concave, fused with fronto-lateral margin of peristomium (Fig. 2A, G), posteriorly extending over 32 chaetigers (to end of chaetiger 26 in lectotype) as a low narrow caruncle, shorter in small individuals (Fig. 4A). Nuchal organs U-shaped ciliary bands on sides of caruncle (Fig. 2A). Length of nuchal organs was strongly correlated with individual number of branchiate chaetigers (Fig. 4C, r 2 = 0.7247, n = 76). Well-developed occipital antenna present in the lectotype (Fig. 3B) and in some specimens (21 of the 69 paralectotypes deposited at the UUZM), but may often be small or even greatly reduced (Fig. 3A). Two pairs of red eyes (appearing very dark red, almost black, in large formalin-fixed specimens) arranged trapezoidally, comprising one pair of large median eyes and one pair of small lateral eyes situated anteriorly and set wider apart; lateral eyes well separated from median pair but often deeply embedded inside epithelium and hardly discernible, especially in large individuals. Palps as long as 5–15 chaetigers, with deep frontal longitudinal groove lined with cilia. Chaetiger 1 with well-developed capillary chaetae and small postchaetal lamellae in both rami; notopodial lamellae triangular; neuropodial lamellae rounded. All notopodia with capillary chaetae only. Low prechaetal lamellae present in noto- and neuropodia on anterior chaetigers after chaetiger 1. Notopodial postchaetal lamellae large, leaf-like on branchiate chaetigers, greatly diminishing in size on posterior abranchiate chaetigers; lamellae on anterior branchiate chaetigers with upper tips pointed (Fig. 2C–E). Neuropodial postchaetal lamellae ear-like on branchiate chaetigers, greatly diminishing in size on posterior abranchiate chaetigers. Branchiae from chaetiger 2, up to 35 pairs (on chaetigers 2–32 in lectotype); first pair shorter or similar in length to notopodial postchaetal lamellae of chaetiger 2; from chaetigers 5–6 branchiae up to twice as long as notopodial postchaetal lamellae (Figs 2E, 3A, B), gradually diminishing in size on succeeding chaetigers, posteriorly present on 0–10 chaetigers after end of nuchal organs (Figs 4A, C, 19A). Branchiae free from lamellae, slightly flattened, with surfaces oriented perpendicular to body axis, with ciliation along inner and outer edges. Afferent and efferent branchial blood vessels forming a loop and interconnected by numerous circular capillaries giving branchiae annulate appearance. Individual number of branchiae was strongly correlated with length of nuchal organs (Fig. 4C). Dorsal transverse crests connecting notopodial postchaetal lamellae present, one per chaetiger, beginning on a chaetiger posterior to caruncle and continuing up to the 21 succeeding chaetigers (Fig. 2B) (on chaetigers 27–47 in lectotype). Lateral interneuropodial pouches from chaetigers 8–25 (from chaetiger 17 in lectotype) to end of the body. Anterior start of pouches was weakly correlated with individual number of branchiate chaetiger (Fig. 18A, r 2 = 0.3928, n = 77). Sabre chaetae in neuropodia from chaetigers 10–20 (from chaetiger 19 in lectotype), from more anterior chaetigers in small individuals (Fig. 4B), 1–3 in a tuft; chaetae with weak granulation on shaft. First appearance of sabre chaetae in neuropodia was weakly correlated with individual number of branchiae (Fig. 4D, r 2 = 0. 2879, n = 76). Hooded hooks in neuropodia from chaetigers 18–32 (from chaetiger 29 in lectotype), from more anterior chaetigers in small individuals (Fig. 4B), up to 14 in a series. Hooks tridentate or quinquedentate, with one or two pairs of upper teeth and a very small median superior tooth above main fang (Fig. 3D). Start of hooks in neuropodia was moderately correlated with the individual number of branchiae (Fig. 4D, r 2 = 0.6559, n = 76). Pygidium with up to seven pairs of cirri arranged around terminal anus, comprising one pair of ventral cirri and up to six pairs of thinner and longer dorsal cirri (Fig. 3C); fewer cirri in small individuals. Digestive tract without gizzard-like structure. Reproduction. Laonice bahusiensis is dioecious. The lectotype is a female with small (apparently not yet fully developed) oocytes. Methylene green staining. Tips of notopodial postchaetal lamellae from chaetigers 4–5 to chaetigers 6–12 usually well stained (Fig. 2G). Narrow bands of light staining present dorsally along anterior edge of prostomium and peristomium (Fig. 2G) and ventrally around mouth (Fig. 2F). Remarks. Most species of Laonice typically have one occipital antenna on the prostomium, inserted between the palp bases. In the L. bahusiensis examined in our study, the length of the occipital antennae varied a great deal, with more than half of the type specimens having neither occipital antenna nor scars to suggest that they were broken. In other type and non-type specimens possessing all the other characteristics of the species, a well-developed antenna was not always present between the palps. Distribution. Norwegian Sea, from Kolvereidvågen (64.9°N) south to Kattegat, Västra, Sweden and off Frederikshavn, Denmark (Fig. 5A). At 14–373 m depth., Published as part of Sikorski, Andrey V., Radashevsky, Vasily I., Castelli, Alberto, Pavlova, Lyudmila V., Nygren, Arne, Malyar, Vasily V., Borisova, Polina B., Mikac, Barbara, Rousou, Maria, Martin, Daniel, Gil, João, Pacciardi, Lorenzo & Langeneck, Joachim, 2021, Revision of the Laonice bahusiensis complex (Annelida: Spionidae) with a description of three new species, pp. 253-283 in Zootaxa 4996 (2) on pages 261-264, DOI: 10.11646/zootaxa.4996.2.2, http://zenodo.org/record/5069822, {"references":["Soderstrom, A. (1920) Studien uber die Polychatenfamilie Spionidae. Inaugural-Dissertation. Almquist & Wicksells, Uppsala, 286 pp.","Maciolek, N. J. (2000) New species and records of Aonidella, Laonice, and Spiophanes (Polychaeta: Spionidae) from shelf and slope depths of the Western North Atlantic. Bulletin of Marine Science, 67 (1), 529 - 547.","Jelsing, J. (2003) Ultrastructural studies of dorsal ciliated organs in Spionidae (Annelida: Polychaeta). Hydrobiologia, 496 (1 - 3), 241 - 251. https: // doi. org / 10.1023 / A: 1026105200983.","Sikorski, A. V. (2003) Laonice (Polychaeta, Spionidae) in the Arctic and the North Atlantic. Sarsia, 88 (5), 316 - 345. https: // doi. org / 10.1080 / 00364820310002551","Jelsing, J. & Eibye-Jacobsen, D. (2010) Ultrastructure of the extensively developed nuchal organs of Laonice bahusiensis (Annelida: Canalipalpata: Spionidae). Journal of Morphology, 271 (3), 376 - 382. https: // doi. org / 10.1002 / jmor. 10813"]}

Laonice grimaldii Sikorski, Nygren & Mikac n. sp. http://zoobank.org:act: D21FC5F2-B9E2-4840-B75F-2D24EBBC2F45 (Figs 10, 11, 12A–C, 13, 14A, 18C, 19B, Table 3) Aonides cirrata: Fauvel, 1909 (Part.): 4–5. Not M. Sars 1851. Laonice cirrata: Fauvel, 1927 (Part.): 37–38, fig. 12a–e. Not M. Sars 1851. Type locality. ITALY, Ligurian Sea, st. GAS 10 ter., 43.6368°N, 10.0983°E, 70–80 m, muddy sand. Type material. MOM INV-0022706 (holotype), INV-0022681–0022693, 0022698, 0022699 (35 paratypes); HUJI NVPOLY-2950 (4 paratypes); MIMB 39037–39039, 39047 (23 paratypes); MNCNM 16.01 /18549–18566 (144 paratypes). Adult morphology. Holotype largest complete specimen 40 mm long, 0.8 mm wide for 109 chaetigers. Smallest complete paratype 13 mm long, 0.26 mm wide for 61 chaetigers. Pigmentation absent on body and palps. Gatherings of glandular cells on branchiae and postchaetal lamellae appearing dark in some formalin-fixed specimens. Prostomium triangular, anteriorly wide, truncate or broadly rounded, fused with anterior margin of peristomium (Figs 10A, B, F, 12A), extending posteriorly to chaetiger 37 (to end of chaetiger 33 in holotype) as a low caruncle (Figs 10A, 12A), shorter in small individuals (Fig. 13A). Posterior end of caruncle flat and often indistinct (Fig. 12B). Nuchal organs U-shaped ciliary bands on sides of caruncle (Fig. 10A). Length of nuchal organs was strongly correlated with individual number of branchiate chaetigers (Fig. 13C, r 2 = 0.8927, n = 30). Occipital antenna present, usually small (Fig. 10A, B, F, 12A). One pair of medium-sized median eyes as transverse short wide bands or oblong oval spots oriented obliquely (Figs 10A, F, 12A). Palps as long as 10–14 chaetigers, with deep frontal longitudinal groove lined with cilia. Chaetiger 1 with short capillaries and moderate sized postchaetal lamellae in both rami. Capillaries in each anterior parapodium arranged in two vertical rows. All notopodia with capillary chaetae only. Notopodial postchaetal lamellae with pointed upper tips usually on 3–4 anterior chaetigers (Fig. 10A), occasionally on 5–9 anterior chaetigers (Fig. 11A–D), with rounded to truncate upper edge on succeeding chaetigers (Fig. 11E, F). Notopodial postchaetal lamellae of chaetiger 3 slightly larger than those of chaetiger 4. Notopodial lamellae and branchiae on anterior chaetigers usually overlapping middorsally, thus obscuring observation of nuchal organs. Notopodial postchaetal lamellae on last branchiate chaetiger and on about seven succeeding postbranchial chaetigers with lower part expanded ventrally (Fig. 11G). Neuropodial postchaetal lamellae until chaetiger 4 with upper part acute (Fig. 11A, B), on succeeding chaetigers with rounded edges (Fig. 11C–G). Branchiae from chaetiger 2 to chaetigers 12–45 (on chaetigers 2–38 in holotype), fewer in smaller individuals (Fig. 13A). Branchiae short on anterior chaetigers, full-sized and slightly longer than notopodial postchaetal lamellae from chaetigers 4–5 onwards (Fig. 11B–F). Individual number of branchiae was strongly correlated with length of nuchal organs (Fig. 13C). Dorsal transverse crests absent on chaetigers with nuchal organs (Fig. 12B). Two dorsal crests usually present on each of 2–3 last branchiate chaetigers (Fig. 10D). Anterior crest on each of those chaetigers appearing as an extension of posterior sides of lateral interneuropodial pouches onto dorsum (Fig. 10C, D), becoming prominent from chaetiger 16 (Fig. 10C) and forming complete dorsal crests on 3–4 last branchiate chaetigers (Fig. 10D). Posterior crests appearing 1–2 chaetigers after first start of anterior crests and interconnecting notopodial postchaetal lamellae (Fig. 10D). Anterior and posterior crests almost equal in maximum height on 2–3 last branchiate chaetigers. Anterior crests reduced in height on succeeding chaetigers and disappearing completely after 2–6 postbranchiate chaetigers. Posterior crests on approximately 20 postbranchiate chaetigers (until chaetiger 57 in holotype) becoming nearly as high as notopodial postchaetal lamellae. Lateral interneuropodial pouches from chaetigers 7–26 (from chaetiger 14 in holotype) to body end. Anterior start of pouches was moderately correlated with individual number of branchiate chaetigers (Fig. 18C, r 2 = 0.5278, n = 29). Sabre chaetae in neuropodia from chaetigers 8–18 (from chaetiger 15 in holotype), from more anterior chaetigers in smaller individuals (Fig. 13B), 1–2 in a tuft below vertical row of capillaries or hooded hooks (Fig. 11F, G). Anterior start of sabre chaetae was weakly correlated with individual number of branchiate chaetigers (Fig. 13D, r 2 = 0.2854, n = 30). Hooded hooks in neuropodia from chaetigers 13–36 (from chaetiger 34 in holotype), from more anterior chaetigers in smaller individuals (Fig. 13B), up to 13 in a series (Fig. 11G). Hooks tri- or quinquedentate, with one or two pairs of small apical teeth above main fang (Figs 11H, 12C); superior pair of teeth tiny and hardly discernible, not developed in some hooks. Anterior start of hooks was strongly correlated with individual number of branchiate chaetigers (Fig. 13D, r 2 = 0.9189, n = 31). Pygidium with one pair of short ventral cirri and up to four pairs of longer and thinner thread-like dorsal cirri with swollen bases (Fig. 10E). Digestive tract without gizzard-like structure. Methylene green staining. Intensely stained narrow band along frontal edge of prostomium and peristomium (Fig. 10F); usually stained as well the posterior surfaces, lateral margins and sometimes frontal surfaces of distal parts of notopodial postchaetal lamellae from chaetiger 4 to chaetiger 9 (rarely up to chaetiger 19), and basal parts of pygidial cirri (Fig. 10F). Characteristic diffused staining on ventral body surface, most intensely from approximately chaetiger 21 to chaetigers 33–35 (Fig. 10G). Remarks. Adult L. grimaldii n. sp. appear similar to L. bahusiensis in having the prostomium fused with the frontal margin of the peristomium, dorsal crests only on postbranchiate chaetigers and a narrow band along the frontal edge of prostomium stained with Methylene Green. They differ from all other species of Laonice in having double dorsal crests on chaetigers near the last branchiate chaetiger. Within the L. bahusiensis complex, L. grimaldii n. sp. differs from the other species in having rounded instead of acute notopodial postchaetal lamellae after chaetigers 4–9, and branchiae being slightly longer instead of considerably longer (from 1.3 to 2 times longer) than notopodial postchaetal lamellae. Moreover, the adults of L. grimaldii n. sp. characteristically have a caruncle flattened and therefore indistinct on its posterior end (Fig. 12B), contrary to adults of most other species, which have distinct caruncles that terminate clearly and abruptly. The adults of L. grimaldii n. sp. are similar to those of L. antipoda Sikorski, 2011 from South Africa in various numeric characteristics, especially in the presence of branchiae and prominent dorsal crests on several chaetigers posterior to nuchal organs. In the original description of L. antipoda, Sikorski (2011) noticed the absence of an occipital antenna in the specimens examined but assumed that it might be due to damage. Recent examinations of additional material of L. antipoda from Angola and Gabon (Sikorski, unpublished), however, confirmed the lack of an occipital antenna in this species (Fig. 12D). Thus, L. grimaldii n. sp. differs from L. antipoda by having smaller adults (maximum body width ≤ 0.8 mm in L. grimaldii n. sp. versus ≥ 1.0 mm in L. antipoda), double dorsal crests on 1–2 last branchiate chaetiger(s), an occipital antenna on the prostomium, and notopodial postchaetal lamellae of chaetiger 4 smaller than those of chaetiger 3 (instead of much bigger as in L. antipoda). Etymology. The name of the species refers to the House of Grimaldi, the princely family of Monaco and, more specifically, it is dedicated to Prince Albert I, who collected some of the specimens used for the present species description. Distribution. Mediterranean Sea (Fig. 14A). At 4–80 m depth., Published as part of Sikorski, Andrey V., Radashevsky, Vasily I., Castelli, Alberto, Pavlova, Lyudmila V., Nygren, Arne, Malyar, Vasily V., Borisova, Polina B., Mikac, Barbara, Rousou, Maria, Martin, Daniel, Gil, João, Pacciardi, Lorenzo & Langeneck, Joachim, 2021, Revision of the Laonice bahusiensis complex (Annelida: Spionidae) with a description of three new species, pp. 253-283 in Zootaxa 4996 (2) on pages 269-273, DOI: 10.11646/zootaxa.4996.2.2, http://zenodo.org/record/5069822, {"references":["Fauvel, P. (1909) Deuxieme note preliminaire sur les polychetes provenant des campagnes de l'Hirondelle et de la Princesse- Alice, ou deposees dans la Musee Oceanographique de Monaco. Bulletin de l'Institute Oceanographique de Monaco, 142, 1 - 76.","Sars, M. (1851) Beretning om en i Sommeren 1849 foretagen zoologisk Reise i Lofoten og Finmarken. Nyt Magazin for Naturvidenskaberne, 6 (2), 121 - 211.","Fauvel, P. (1927) Polychetes sedentaires. Addenda aux Errantes, Archiannelides, Myzostomaires. Faune de France, 16, 1 - 494.","Sikorski, A. V. (2011) Review of Laonice (Spionidae, Annelida) with remarks on several species and a description of a new species from South Africa. Italian Journal of Zoology, 78 (S 1), 201 - 214. https: // doi. org / 10.1080 / 11250003.2011.617218"]}

Laonice (Laonice) Malmgren, 1867 Laonice (Laonice): Sikorski, Gunton & Pavlova 2017: 962. Type species: Nerine cirrata M. Sars, 1851. By monotypy. Diagnosis. Prostomium fused with fronto-lateral margin of peristomium. Caruncle long, extending beyond half of branchiate chaetigers. Capillary chaetae on anterior chaetigers arranged in two vertical rows. Hooded hooks with main fang surmounted by paired apical teeth. Hooks absent in notopodia. Remarks. The nominotypical subgenus Laonice (Laonice) was recognized by Sikorski et al. (2017) based on a morphological analysis of Laonice species. Molecular analysis provided in the present study (Fig. 1) supports the monophyly of this group., Published as part of Sikorski, Andrey V., Radashevsky, Vasily I., Castelli, Alberto, Pavlova, Lyudmila V., Nygren, Arne, Malyar, Vasily V., Borisova, Polina B., Mikac, Barbara, Rousou, Maria, Martin, Daniel, Gil, João, Pacciardi, Lorenzo & Langeneck, Joachim, 2021, Revision of the Laonice bahusiensis complex (Annelida: Spionidae) with a description of three new species, pp. 253-283 in Zootaxa 4996 (2) on page 259, DOI: 10.11646/zootaxa.4996.2.2, http://zenodo.org/record/5069822, {"references":["Malmgren, A. J. (1867) Annulata Polychaeta Spetsbergiae, Gronlandiae, Islandiae et Scandinaviae Hactenus Cognita. Ex Officina Frenckelliana, Helsingforsiae, 127 pp. https: // doi. org / 10.5962 / bhl. title. 13358","Sikorski, A., Gunton, L. M. & Pavlova, L. (2017) Laonice species (Polychaeta, Spionidae) from the Whittard Canyon (NE Atlantic) with descriptions of two new species. Journal of the Marine Biological Association of the United Kingdom, 97 (5), 961 - 973. https: // doi. org / 10.1017 / s 0025315417000480","Sars, M. (1851) Beretning om en i Sommeren 1849 foretagen zoologisk Reise i Lofoten og Finmarken. Nyt Magazin for Naturvidenskaberne, 6 (2), 121 - 211."]}

Marine Non-Indigenous Species (NIS) are animals and plants introduced accidently or deliberately into the European seas, originating from other seas of the globe. About 800 marine non-indigenous species (NIS)currently occur in the European Union national marine waters, several of which have negative impacts on marine ecosystem services and biodiversity. Under the Marine Strategy Framework Directive (MSFD) Descriptor 2 (D2), EU Member States (MSs) need to consider NIS in their marine management strategies. The Descriptor D2 includes one primary criterion (D2C1: new NIS introductions), and two secondary criteria (D2C2 and D2C3). The D2 implementation is characterized by a number of issues and uncertainties which can be applicable to the Descriptor level (e.g. geographical unit of assessment, assessment period, phytoplanktonic, parasitic, oligohaline NIS, etc.), to the primary criterion D2C1 level (e.g. threshold values, cryptogenic, questionable species, etc), and to the secondary criteria D2C2 and D2C3. The current report tackles these issues and provides practical recommendations aiming at a smoother and more efficient implementation of D2 and its criteria at EU level. They constitute a solid operational output which can result in more comparable D2 assessments among MSsand MSFD regions/subregions. When it comes to the policy-side, the current report calls for a number of different categories of NIS to be reported in D2 assessments, pointing the need for the species to be labelled/categorised appropriately in the MSFD reporting by the MSs. These suggestions are proposed to be communicated to the MSFD Working Group of Good Environmental Status (GES) and subsequently to the Marine Strategy Coordination Group (MSCG) of MSFD. Moreover, they can serve as an input for revising the Art. 8 Guidelines.

Marine Non-Indigenous Species (NIS) are animals and plants introduced accidently or deliberately into the European seas, originating from other seas of the globe. About 800 marine non-indigenous species (NIS) currently occur in the European Union national marine waters, several of which have negative impacts on marine ecosystem services and biodiversity. Under the Marine Strategy Framework Directive (MSFD) Descriptor 2 (D2), EU Member States (MSs) need to consider NIS in their marine management strategies. The Descriptor D2 includes one primary criterion (D2C1: new NIS introductions), and two secondary criteria (D2C2 and D2C3). The D2 implementation is characterized by a number of issues and uncertainties which can be applicable to the Descriptor level (e.g. geographical unit of assessment, assessment period, phytoplanktonic, parasitic, oligohaline NIS, etc.), to the primary criterion D2C1 level (e.g. threshold values, cryptogenic, questionable species, etc), and to the secondary criteria D2C2 and D2C3. The current report tackles these issues and provides practical recommendations aiming at a smoother and more efficient implementation of D2 and its criteria at EU level. They constitute a solid operational output which can result in more comparable D2 assessments among MSs and MSFD regions/subregions. When it comes to the policy-side, the current report calls for a number of different categories of NIS to be reported in D2 assessments, pointing the need for the species to be labelled/categorised appropriately in the MSFD reporting by the MSs. These suggestions are proposed to be communicated to the MSFD Working Group of Good Environmental Status (GES) and subsequently to the Marine Strategy Coordination Group (MSCG) of MSFD. Moreover, they can serve as an input for revising the Art. 8 Guidelines.

Marine Non-Indigenous Species (NIS) are animals and plants introduced accidently or deliberately into the European seas, originating from other seas of the globe. About 800 marine non-indigenous species (NIS) currently occur in the European Union national marine waters, several of which have negative impacts on marine ecosystem services and biodiversity. Under the Marine Strategy Framework Directive (MSFD) Descriptor 2 (D2), EU Member States (MSs) need to consider NIS in their marine management strategies. The Descriptor D2 includes one primary criterion (D2C1: new NIS introductions), and two secondary criteria (D2C2 and D2C3). The D2 implementation is characterized by a number of issues and uncertainties which can be applicable to the Descriptor level (e.g. geographical unit of assessment, assessment period, phytoplanktonic, parasitic, oligohaline NIS, etc.), to the primary criterion D2C1 level (e.g. threshold values, cryptogenic, questionable species, etc), and to the secondary criteria D2C2 and D2C3. The current report tackles these issues and provides practical recommendations aiming at a smoother and more efficient implementation of D2 and its criteria at EU level. They constitute a solid operational output which can result in more comparable D2 assessments among MSs and MSFD regions/subregions. When it comes to the policy- side, the current report calls for a number of different categories of NIS to be reported in D2 assessments, pointing the need for the species to be labelled/categorised appropriately in the MSFD reporting by the MSs. These suggestions are proposed to be communicated to the MSFD Working Group of Good Environmental Status (GES) and subsequently to the Marine Strategy Coordination Group (MSCG) of MSFD. Moreover, they can serve as an input for revising the Art. 8 Guidelines.

1. Ampelisca brevicornis (Costa, 1853) Materials examined. 97 individuals; Stations Vas.1(8), 3(6), 9(1), 11(1), 13(1), 16(2), 21(1), 22(7), 24(2), 28(21), 29(12), 32(20), 37(15). Distribution. Atlanto-Mediterranean. The species was reported from Cyprus by Kocatas et al. (2001). In Vasiliko Bay (current study), it is considered to be a common species (FI: 30.95 %). Ecology. Ampelisca brevicornis was found in soft-bottom sediments at 5 m depth (Kocatas et al. 2001). In Vasiliko Bay A. brevicornis was found at depths of 4���13 m (7 + 3 m) mainly in sandy but also muddy sand substrates, with organic matter ranging between 1.86���7.93% (3.23 + 1.55 %), and in association with C. prolifera, C. nodosa and P. oceanica. It was found in stations classified according to the Bentix Index as in: ���high��� (6 stations, 39 individuals), ���good��� (5 stations, 41 individuals) and ���moderate��� (2 stations, 17 individuals). Individuals were recorded at the ports (38 individuals) and at the freshwater underwater and the power station outlet stations (9 individuals)., Published as part of Rousou, Maria, Plaiti, Wanda, Lowry, Jim, Charalambous, Stephanos & Chintiroglou, Chariton Charles, 2020, Amphipoda species (Suborders: Amphilochidea and Senticaudata) from Vasiliko Bay, Cyprus: New records, information on their biogeography and an annotated checklist from the coasts of Cyprus, pp. 373-408 in Zootaxa 4896 (3) on page 382, DOI: 10.11646/zootaxa.4896.3.3, http://zenodo.org/record/4383886, {"references":["Costa, A. (1853) Relazione sulla memoria del Dottor Achille Costa, di ricerche su' crostacei amfipodi del regno di Napoli. Rendiconto della Societa Reale Borbonica, Accademia delle Scienze, New Series, 2, 167 - 178. https: // doi. org / 10.5962 / bhl. title. 2070","Kocatas, A., Katagan, T. & Huseyin. A. B. (2001) Contribution to the knowledge of the crustacean fauna of Cyprus. Israel Journal of Zoology, 47 (2), 147 - 160. https: // doi. org / 10.1560 / YQL 8 - 4 PBT- 12 W 2 - 82 HV"]}

4. Peltocoxa gibbosa (Schiecke, 1977) Materials examined. 2 individuals; Stations Vas. 33(1), 41(1). Distribution. Atlanto-Mediterranean (Bellan-Santini & Ruffo 2003; Christodoulou et al. 2013). The species is reported in Cyprus for the first time. In Vasiliko Bay it is considered to be a rare species (FI: 4.76 %). Ecology. In Vasiliko Bay it was found at depths of 32 and 40 m (36 + 6 m), in muddy sand substrate with organic matter ranging between 8.848 % and 11.119 % (9.983 + 1.605 %) and in association with C. prolifera. It was found in stations classified as ���moderate��� according to the Bentix Index., Published as part of Rousou, Maria, Plaiti, Wanda, Lowry, Jim, Charalambous, Stephanos & Chintiroglou, Chariton Charles, 2020, Amphipoda species (Suborders: Amphilochidea and Senticaudata) from Vasiliko Bay, Cyprus: New records, information on their biogeography and an annotated checklist from the coasts of Cyprus, pp. 373-408 in Zootaxa 4896 (3) on page 383, DOI: 10.11646/zootaxa.4896.3.3, http://zenodo.org/record/4383886, {"references":["Schiecke U. (1977) Zwei neue Vertreter der Cyproideinae (Amphipoda: Amphilochidae) aus dem Mittelmeer: Pseudopeltocoxa gibbosa n. g., n. sp. und Peltocoxa mediterranea n. sp. Bolletino del Museo Civico di Storia Naturale di Verona, 4, 525 - 542.","Bellan-Santini, D. & Ruffo, S. (2003) Biogeography of benthic marine amphipods in Mediterranean Sea. Biogeographia-The Journal of Integrative Biogeography, 24 (1), 273 - 292. https: // doi. org / 10.21426 / B 6110176","Christodoulou, M., Paraskevopoulou, S., Syranidou, E. & Koukouras, A. (2013) The amphipod (Crustacea: Peracarida) fauna of the Aegean Sea, and comparison with those of the neighbouring seas. Journal of the Marine Biological Association of the United Kingdom, 93 (5), 1303 - 1327. https: // doi. org / 10.1017 / S 002531541200183 X"]}

42. Jassa marmorata Holmes, 1905 Materials examined. 7 individuals; Stations Vas. 2(1), 3(1), 5(2), 7(1), 11(1), 35(1). Distribution. Atlanto-Mediterranean. The species was reported from Cyprus by Kocatas et al. (2001) and Christodoulou et al. (2013). In Vasiliko Bay (current study), it is considered as a rare species (FI: 14.28 %). Ecology. Jassa marmorata was found in association with algae, on rocks and stones and on sandy substrates at 3 m depth (Kocatas et al. 2001). In Vasiliko Bay it was found at 5���58 m depth (25 + 20 m), mainly in muddy sand and also sand substrates, with organic matter ranging between 1.859 ���8.416 % (5.237 + 2.606 %) and in association with C. prolifera, C. cylindracea and P. oceanica. It was found in stations classified according to the Bentix Index as in: ���high��� (1 station, 1 individual), ���good��� (3 stations, 4 individuals) and ���moderate��� (2 stations, 2 individuals). Three individuals were recorded at a fish farm station., Published as part of Rousou, Maria, Plaiti, Wanda, Lowry, Jim, Charalambous, Stephanos & Chintiroglou, Chariton Charles, 2020, Amphipoda species (Suborders: Amphilochidea and Senticaudata) from Vasiliko Bay, Cyprus: New records, information on their biogeography and an annotated checklist from the coasts of Cyprus, pp. 373-408 in Zootaxa 4896 (3) on page 392, DOI: 10.11646/zootaxa.4896.3.3, http://zenodo.org/record/4383886, {"references":["Holmes, S. J. (1905) The Amphipoda of southern New England. Bulletin of the United States. Bureau of Fisheries, 24, 457 - 529. [https: // www. biodiversitylibrary. org / page / 25135693]","Kocatas, A., Katagan, T. & Huseyin. A. B. (2001) Contribution to the knowledge of the crustacean fauna of Cyprus. Israel Journal of Zoology, 47 (2), 147 - 160. https: // doi. org / 10.1560 / YQL 8 - 4 PBT- 12 W 2 - 82 HV","Christodoulou, M., Paraskevopoulou, S., Syranidou, E. & Koukouras, A. (2013) The amphipod (Crustacea: Peracarida) fauna of the Aegean Sea, and comparison with those of the neighbouring seas. Journal of the Marine Biological Association of the United Kingdom, 93 (5), 1303 - 1327. https: // doi. org / 10.1017 / S 002531541200183 X"]}

45. Maera grossimana (Montagu, 1808) Materials examined. 22 individuals; Stations Vas. 2(12), 5(1), 14(4), 20(1), 35(3), 39(1). Distribution. Atlanto-Mediterranean (Bellan-Santini & Ruffo 2003). The species was reported from Cyprus by Kocatas et al. (2001). In Vasiliko Bay (current study), it is considered as a rare species (FI: 14.28 %). Ecology. Maera grossimana was found in association with algae and seagrasses, on rocks and stones, in sponges and in artificial substrata at 35 m depth (Kocatas et al. 2001). In Vasiliko Bay it was found at 20���48 m depth (32 + 9 m), mainly in muddy sand and also sand substrates, with organic matter ranging between 4.621 ���8.416 % (6.933 + 1.456 %) and in association with C. prolifera, C. cylindracea, C. nodosa and P. oceanica. It was found in stations classified according to the Bentix Index as in: ���good��� (2 stations, 5 individuals) and ���moderate��� (4 stations, 17 individuals). It was also recorded at fish farm stations (14 individuals)., Published as part of Rousou, Maria, Plaiti, Wanda, Lowry, Jim, Charalambous, Stephanos & Chintiroglou, Chariton Charles, 2020, Amphipoda species (Suborders: Amphilochidea and Senticaudata) from Vasiliko Bay, Cyprus: New records, information on their biogeography and an annotated checklist from the coasts of Cyprus, pp. 373-408 in Zootaxa 4896 (3) on pages 392-393, DOI: 10.11646/zootaxa.4896.3.3, http://zenodo.org/record/4383886, {"references":["Montagu, G. (1808) Description of several marine animals found on the south coast of Devonshire. Transactions of the Linnean Society of London, 9, 81 - 114. https: // doi. org / 10.1111 / j. 1096 - 3642.1818. tb 00327. x","Bellan-Santini, D. & Ruffo, S. (2003) Biogeography of benthic marine amphipods in Mediterranean Sea. Biogeographia-The Journal of Integrative Biogeography, 24 (1), 273 - 292. https: // doi. org / 10.21426 / B 6110176","Kocatas, A., Katagan, T. & Huseyin. A. B. (2001) Contribution to the knowledge of the crustacean fauna of Cyprus. Israel Journal of Zoology, 47 (2), 147 - 160. https: // doi. org / 10.1560 / YQL 8 - 4 PBT- 12 W 2 - 82 HV"]}

20. Synchelidium maculatum Stebbing, 1906 Materials examined. 6 individuals; Stations Vas. 17(1), 24(2), 30(1), 33(2). Distribution. Atlanto-Mediterranean (Bellan-Santini & Ruffo 2003). The species is reported in Cyprus (Vasiliko bay) for the first time. In Vasiliko Bay it is considered to be a rare species (FI: 9.52 %). Ecology. In Vasiliko Bay it was found at depths of 12 and 35 m (23 + 12 m), in muddy sand and sand substrates with organic matter ranging between 2.2���8.848 % (5.123 + 3.028 %), in association with C. prolifera and C. nodosa. It was found in stations classified according to the Bentix Index as in: ���high��� (1 station, 2 individuals), ���good��� (1 station, 1 individual) and ���moderate��� (2 stations, 3 individuals)., Published as part of Rousou, Maria, Plaiti, Wanda, Lowry, Jim, Charalambous, Stephanos & Chintiroglou, Chariton Charles, 2020, Amphipoda species (Suborders: Amphilochidea and Senticaudata) from Vasiliko Bay, Cyprus: New records, information on their biogeography and an annotated checklist from the coasts of Cyprus, pp. 373-408 in Zootaxa 4896 (3) on pages 386-387, DOI: 10.11646/zootaxa.4896.3.3, http://zenodo.org/record/4383886, {"references":["Stebbing, T. R. R. (1906) Amphipoda. I. Gammaridea. Das Tiereich, 21, 1 - 806.","Bellan-Santini, D. & Ruffo, S. (2003) Biogeography of benthic marine amphipods in Mediterranean Sea. Biogeographia-The Journal of Integrative Biogeography, 24 (1), 273 - 292. https: // doi. org / 10.21426 / B 6110176"]}

41. Ericthonius punctatus (Spence Bate, 1857) Materials examined. 24 individuals; Stations Vas. 9(7), 13(4), 17(1), 30(1), 32(8), 37(1), 40(2). Distribution. Atlanto-Mediterranean. The species was reported from Cyprus by Kocatas et al. (2001) and Christodoulou et al. (2013). In Vasiliko Bay (current study), it is considered as a rare species (FI: 16.6 %). Ecology. In Vasiliko Bay it was found at depths of 7���35 m (14 + 10 m), in sandy and muddy sand substrates with organic matter ranging between 2.2���6.462 % (3.996 + 1.725 %) and in association with C. nodosa. It was found in stations classified according to the Bentix Index as in: ���high��� (2 stations, 10 individuals), ���good��� (3 stations, 12 individuals) and ���moderate��� (2 stations, 2 individuals)., Published as part of Rousou, Maria, Plaiti, Wanda, Lowry, Jim, Charalambous, Stephanos & Chintiroglou, Chariton Charles, 2020, Amphipoda species (Suborders: Amphilochidea and Senticaudata) from Vasiliko Bay, Cyprus: New records, information on their biogeography and an annotated checklist from the coasts of Cyprus, pp. 373-408 in Zootaxa 4896 (3) on page 392, DOI: 10.11646/zootaxa.4896.3.3, http://zenodo.org/record/4383886, {"references":["Spence Bate, C. S. (1857) On a new amphipod: Iphimedia eblanae. The Natural History Review, 4, 229 - 230.","Kocatas, A., Katagan, T. & Huseyin. A. B. (2001) Contribution to the knowledge of the crustacean fauna of Cyprus. Israel Journal of Zoology, 47 (2), 147 - 160. https: // doi. org / 10.1560 / YQL 8 - 4 PBT- 12 W 2 - 82 HV","Christodoulou, M., Paraskevopoulou, S., Syranidou, E. & Koukouras, A. (2013) The amphipod (Crustacea: Peracarida) fauna of the Aegean Sea, and comparison with those of the neighbouring seas. Journal of the Marine Biological Association of the United Kingdom, 93 (5), 1303 - 1327. https: // doi. org / 10.1017 / S 002531541200183 X"]}

34. Apherusa chiereghinii Giordani-Soika, 1950 Materials examined. 25 individuals; Stations Vas. 2(2), 5(5), 7(1), 9(1), 12(2), 13(2), 14(2), 17(2), 18(1), 21(2), 24(2), 36(1), 39(1), 41(1). Distribution. Mediterranean (Bellan-Santini & Ruffo 2003), Black Sea (WoRMS). The species was reported from Cyprus by Kocatas et al. (2001). In Vasiliko Bay (current study), it is considered as a common species (FI: 33.3 %). Ecology. Apherusa chiereghinii was found in association with algae and on soft sediments (mud, mud and sand mixture) at 0���45 m depth (Kocatas et al. 2001). In Vasiliko Bay it was found at 4���58 m depth (30 + 18 m), in sandy and muddy sand substrates with organic matter ranging between 2.2���6���11.119 % (5.543 + 2.826 %) and in association with C. prolifera, C. cylindracea, C. nodosa, Dictyota spp. and H. stipulacea. It was found in stations classified according to the Bentix Index as in: ���high��� (2 stations, 4 individuals), ���good��� (7 stations, 14 individuals) and ���moderate��� (5 stations, 7 individuals). It was recorded at the power station outlet (2 individuals) and fish farm stations (8 individuals)., Published as part of Rousou, Maria, Plaiti, Wanda, Lowry, Jim, Charalambous, Stephanos & Chintiroglou, Chariton Charles, 2020, Amphipoda species (Suborders: Amphilochidea and Senticaudata) from Vasiliko Bay, Cyprus: New records, information on their biogeography and an annotated checklist from the coasts of Cyprus, pp. 373-408 in Zootaxa 4896 (3) on page 390, DOI: 10.11646/zootaxa.4896.3.3, http://zenodo.org/record/4383886, {"references":["Giordani-Soika, A. (1950) Gli Anfipodi gammarini della laguna di Venezia. Archivio di Oceanograja e Limnologia, 6 (2 - 3), 165 - 212.","Bellan-Santini, D. & Ruffo, S. (2003) Biogeography of benthic marine amphipods in Mediterranean Sea. Biogeographia-The Journal of Integrative Biogeography, 24 (1), 273 - 292. https: // doi. org / 10.21426 / B 6110176","Kocatas, A., Katagan, T. & Huseyin. A. B. (2001) Contribution to the knowledge of the crustacean fauna of Cyprus. Israel Journal of Zoology, 47 (2), 147 - 160. https: // doi. org / 10.1560 / YQL 8 - 4 PBT- 12 W 2 - 82 HV"]}

30. Microdeutopus bifidus Myers, 1977 Materials examined. 72 individuals; Stations Vas. 14(41), 18(1), 31(3), 40(27). Distribution. Mediterranean Sea (WoRMS). The species was reported from Cyprus by Pavloudi et al. (2016). In Vasiliko Bay (current study), it is considered as a rare species (FI: 9.52 %). Ecology. In the Eastern Mediterranean Sea M. bifidus was found in association with sponges (Pavloudi et al. 2016), in association with P. oceanica and sea anemones and in sandy substrates (Voutsiadou-Koukoura & Stefanidou, 1995). In Vasiliko Bay it was found at depths of 17���51 m (29 + 15 m), in muddy sand substrate, with organic matter ranging between 5.621 ���8.183 % (6.888 + 1.099 %) and in association with C. prolifera and C. cylindracea. It was found in stations classified according to the Bentix Index as in: ���high��� (1 station, 27 individuals), ���good��� (2 stations, 42 individuals) and ���moderate��� (1 station, 3 individuals)., Published as part of Rousou, Maria, Plaiti, Wanda, Lowry, Jim, Charalambous, Stephanos & Chintiroglou, Chariton Charles, 2020, Amphipoda species (Suborders: Amphilochidea and Senticaudata) from Vasiliko Bay, Cyprus: New records, information on their biogeography and an annotated checklist from the coasts of Cyprus, pp. 373-408 in Zootaxa 4896 (3) on page 389, DOI: 10.11646/zootaxa.4896.3.3, http://zenodo.org/record/4383886, {"references":["Myers, A. A. (1977) Two new species of the amphipod genus Microdeutopus Costa from the Mediterranean Sea. Bolletino del Museo Civico di Storia Naturale di Verona, 4, 475 - 478.","Pavloudi, C., Christodoulou, M. & Mavidis, M. (2016) Macrofaunal assemblages associated with the sponge Sarcotragus foetidus Schmidt, 1862 (Porifera: Demospongiae) at the coasts of Cyprus and Greece. Biodiversity Data Journal, 4, e 8210. https: // doi. org / 10.3897 / BDJ. 4. e 8210"]}

33. Bathyporeia guilliamsoniana (Spence Bate, 1857) Materials examined. 25 individuals; Stations Vas. 1(7), 3(12), 6(1), 9(5) Distribution. Atlanto-Mediterranean (Bellan-Santini & Ruffo 2003). The species was reported from Cyprus by Kocatas et al. (2001). In Vasiliko Bay (current study), it is considered as a rare species (FI: 9.52 %). Ecology. In the Eastern Mediterranean Sea, B. guilliamsoniana was found in sandy substrates at depths 3���5 m (Voultsiadou-Koukoura & Stefanidou 1995; Kocatas et al. 2001). In Vasiliko Bay it was found at depths of 5���10 m (6 + 2 m), in sandy substrates with organic matter ranging between 1.859 ���2.757 % (2.203 + 0.409 %) and in association with C. prolifera and C. cylindracea. It was found in stations classified according to the Bentix Index as in: ���high��� (2 stations, 8 individuals) and ���good��� (2 stations, 17 individuals). It was recorded in underwater outlets (7 individuals)., Published as part of Rousou, Maria, Plaiti, Wanda, Lowry, Jim, Charalambous, Stephanos & Chintiroglou, Chariton Charles, 2020, Amphipoda species (Suborders: Amphilochidea and Senticaudata) from Vasiliko Bay, Cyprus: New records, information on their biogeography and an annotated checklist from the coasts of Cyprus, pp. 373-408 in Zootaxa 4896 (3) on page 390, DOI: 10.11646/zootaxa.4896.3.3, http://zenodo.org/record/4383886, {"references":["Spence Bate, C. S. (1857) On a new amphipod: Iphimedia eblanae. The Natural History Review, 4, 229 - 230.","Bellan-Santini, D. & Ruffo, S. (2003) Biogeography of benthic marine amphipods in Mediterranean Sea. Biogeographia-The Journal of Integrative Biogeography, 24 (1), 273 - 292. https: // doi. org / 10.21426 / B 6110176","Kocatas, A., Katagan, T. & Huseyin. A. B. (2001) Contribution to the knowledge of the crustacean fauna of Cyprus. Israel Journal of Zoology, 47 (2), 147 - 160. https: // doi. org / 10.1560 / YQL 8 - 4 PBT- 12 W 2 - 82 HV","Voultsiadou-Koukoura, E. & Stefanidou, D. (1995) An Account of Our Knowledge of the Amphipod Fauna of the Aegean Sea, Crustaceana, 68 (5), 597 - 615. https: // doi. org / 10.1163 / 156854095 X 00845"]}

10. Leucothoe lilljeborgi Boeck, 1861 Materials examined. 13 individuals; Stations Vas. 12(1), 20(1), 25(2), 30(9). Distribution. Atlanto-Mediterranean (Bellan-Santini & Ruffo 2003). The species was reported from Cyprus by Kocatas et al. (2001). In Vasiliko Bay (current study), it is considered as a rare species (FI: 9.52 %). Ecology. Leucothoe lilljeborgi was found in soft-bottom sediments (mud, mud-and mixture) at 69 m depth (Koca-tas et al. 2001). In Vasiliko Bay it was found at depths of 35���45 m (38 + 5 m), in muddy sand substrate with organic matter ranging between 4.656���6.54 % (5.926 + 0.877 %) and in association with C. prolifera and C. cylindracea. It was found in stations classified as ���moderate��� according to the Bentix Index. It was recorded at the buoy station (2 individuals)., Published as part of Rousou, Maria, Plaiti, Wanda, Lowry, Jim, Charalambous, Stephanos & Chintiroglou, Chariton Charles, 2020, Amphipoda species (Suborders: Amphilochidea and Senticaudata) from Vasiliko Bay, Cyprus: New records, information on their biogeography and an annotated checklist from the coasts of Cyprus, pp. 373-408 in Zootaxa 4896 (3) on page 384, DOI: 10.11646/zootaxa.4896.3.3, http://zenodo.org/record/4383886, {"references":["Boeck, A. (1861) Bemaerkninger angaaende de ved de norske Kyster forekommende Amphipoder. Forhandl. Skand. Naturf. Ott. Mode i Kjopenhavn, 8, 631 - 677.","Bellan-Santini, D. & Ruffo, S. (2003) Biogeography of benthic marine amphipods in Mediterranean Sea. Biogeographia-The Journal of Integrative Biogeography, 24 (1), 273 - 292. https: // doi. org / 10.21426 / B 6110176","Kocatas, A., Katagan, T. & Huseyin. A. B. (2001) Contribution to the knowledge of the crustacean fauna of Cyprus. Israel Journal of Zoology, 47 (2), 147 - 160. https: // doi. org / 10.1560 / YQL 8 - 4 PBT- 12 W 2 - 82 HV"]}

49. Pereionotus testudo (Montagu, 1808) Materials examined. 16 individuals; Stations Vas. 5(1), 10(1), 18(1), 20(3), 23(1), 24(1), 25(2), 26(4), 30(1), 36(1). Distribution. Atlanto-Mediterranean (Bellan-Santini & Ruffo 2003). The species was reported from Cyprus by Kocatas et al. (2001). In Vasiliko Bay (current study), it is considered as a rare species (FI: 23.8 %). Ecology. Pereionotus testudo was found in association with algae, on rocks and stones and on soft-bottom sedi-ments (mud, mud and sand mixture) at 3���70 m depth (Kocatas et al. 2001). In Vasiliko Bay it was found at depths of 13���59 m (34 + 14 m), mainly in muddy sand and also sand substrates, with organic matter ranging between 3.179 ���9.414 % (6.012 + 1.617 %) and in association with C. prolifera, C. cylindracea, C. nodosa and Dictyota spp. It was found in stations classified according to the Bentix Index as in: ���high��� (1 station, 1 individual), ���good��� (5 stations, 8 individuals) and ���moderate��� (4 stations, 7 individuals). It was recorded at a fish farm station (1 individual) and at the buoy (2 individuals)., Published as part of Rousou, Maria, Plaiti, Wanda, Lowry, Jim, Charalambous, Stephanos & Chintiroglou, Chariton Charles, 2020, Amphipoda species (Suborders: Amphilochidea and Senticaudata) from Vasiliko Bay, Cyprus: New records, information on their biogeography and an annotated checklist from the coasts of Cyprus, pp. 373-408 in Zootaxa 4896 (3) on page 393, DOI: 10.11646/zootaxa.4896.3.3, http://zenodo.org/record/4383886, {"references":["Montagu, G. (1808) Description of several marine animals found on the south coast of Devonshire. Transactions of the Linnean Society of London, 9, 81 - 114. https: // doi. org / 10.1111 / j. 1096 - 3642.1818. tb 00327. x","Bellan-Santini, D. & Ruffo, S. (2003) Biogeography of benthic marine amphipods in Mediterranean Sea. Biogeographia-The Journal of Integrative Biogeography, 24 (1), 273 - 292. https: // doi. org / 10.21426 / B 6110176","Kocatas, A., Katagan, T. & Huseyin. A. B. (2001) Contribution to the knowledge of the crustacean fauna of Cyprus. Israel Journal of Zoology, 47 (2), 147 - 160. https: // doi. org / 10.1560 / YQL 8 - 4 PBT- 12 W 2 - 82 HV"]}

32. Microdeutopus versiculatus (Spence Bate, 1857) Materials examined. 46 individuals; Stations Vas. 9(5), 13(1), 17(1), 19(1), 24(3), 31(3), 32(2), 37(13), 40(17). Distribution. Atlanto-Mediterranean (Bellan-Santini & Ruffo 2003; Christodoulou et al. 2013), Black Sea (WoRMS). The species is reported in Cyprus for the first time. In Vasiliko Bay, it is considered as a rare species (FI: 21.42 %). Ecology. In Vasiliko Bay it was found at depths of 7���22 m (12 + 5 m), mainly in muddy sand and also sand substrates, with organic matter ranging between 2.2���7.728 % (3.84 + 1.82 %) and in association with C. prolifera, C. cylindracea and C. nodosa. It was found in stations classified according to the Bentix Index as in: ���high��� (3 stations, 22 individuals), ���good��� (4 stations, 8 individuals) and ���moderate��� (2 stations, 16 individuals). It was recorded at the ports (13 individuals)., Published as part of Rousou, Maria, Plaiti, Wanda, Lowry, Jim, Charalambous, Stephanos & Chintiroglou, Chariton Charles, 2020, Amphipoda species (Suborders: Amphilochidea and Senticaudata) from Vasiliko Bay, Cyprus: New records, information on their biogeography and an annotated checklist from the coasts of Cyprus, pp. 373-408 in Zootaxa 4896 (3) on page 389, DOI: 10.11646/zootaxa.4896.3.3, http://zenodo.org/record/4383886, {"references":["Spence Bate, C. S. (1857) On a new amphipod: Iphimedia eblanae. The Natural History Review, 4, 229 - 230.","Bellan-Santini, D. & Ruffo, S. (2003) Biogeography of benthic marine amphipods in Mediterranean Sea. Biogeographia-The Journal of Integrative Biogeography, 24 (1), 273 - 292. https: // doi. org / 10.21426 / B 6110176","Christodoulou, M., Paraskevopoulou, S., Syranidou, E. & Koukouras, A. (2013) The amphipod (Crustacea: Peracarida) fauna of the Aegean Sea, and comparison with those of the neighbouring seas. Journal of the Marine Biological Association of the United Kingdom, 93 (5), 1303 - 1327. https: // doi. org / 10.1017 / S 002531541200183 X"]}

28. Aora spinicornis Afonso, 1976 Materials examined. 6 individuals; Stations Vas. 9(1), 13(1), 17(1), 24(3). Distribution. Atlanto-Mediterranean (Bellan-Santini & Ruffo 2003). The species is reported in Cyprus (Vasiliko bay) for the first time. In Vasiliko Bay, it is considered as a rare species (FI: 9.52 %). Ecology. In Vasiliko Bay it was found at depths of 7���13 m (10 + 3 m), mainly in sandy and muddy sand substrates, with organic matter ranging between 2.2���3.179 % (2.79 + 0.43 %) and in association with C. nodosa. It was found in stations classified according to the Bentix Index as in: ���high��� (1 station, 3 individuals) and ���good��� (3 stations, 3 individuals)., Published as part of Rousou, Maria, Plaiti, Wanda, Lowry, Jim, Charalambous, Stephanos & Chintiroglou, Chariton Charles, 2020, Amphipoda species (Suborders: Amphilochidea and Senticaudata) from Vasiliko Bay, Cyprus: New records, information on their biogeography and an annotated checklist from the coasts of Cyprus, pp. 373-408 in Zootaxa 4896 (3) on pages 388-389, DOI: 10.11646/zootaxa.4896.3.3, http://zenodo.org/record/4383886, {"references":["Afonso, O. (1976) Amphipoda des Acores cueillis par scaphandrier autonome (avec la description d'une nouvelle espece). Publicacoes do Instituto de Zoologia \" Dr. Augusto Nobre \", Faculdade de Ciencias do Porto, 130, 8 - 38.","Bellan-Santini, D. & Ruffo, S. (2003) Biogeography of benthic marine amphipods in Mediterranean Sea. Biogeographia-The Journal of Integrative Biogeography, 24 (1), 273 - 292. https: // doi. org / 10.21426 / B 6110176"]}

14. Lepidepecreum longicornis (Spence Bate, 1862) Materials examined. 9 individuals; Stations Vas. 1(1), 2(1), 3(1), 30(2), 37(1), 38(3). Distribution. Atlanto-Mediterranean (Bellan-Santini & Ruffo 2003). The species is reported in Cyprus for the first time. In Vasiliko Bay it is considered to be a rare species (FI: 14.28 %). Ecology. In Vasiliko Bay it was found at depths of 5���35 m (19 + 14 m), mainly in muddy sand and also sand substrates, with organic matter ranging between 1.859 ���8.416 % (5.166 + 2.968 %), and in association with C. prolifera and C. cylindracea. It was found in stations classified according to the Bentix Index as in: ���high��� (1 station, 1 individual), ���good��� (1 station, 1 individual) and ���moderate��� (4 stations, 7 individuals). It was recorded at a fish farm station (1 individual) and the port (1 individual)., Published as part of Rousou, Maria, Plaiti, Wanda, Lowry, Jim, Charalambous, Stephanos & Chintiroglou, Chariton Charles, 2020, Amphipoda species (Suborders: Amphilochidea and Senticaudata) from Vasiliko Bay, Cyprus: New records, information on their biogeography and an annotated checklist from the coasts of Cyprus, pp. 373-408 in Zootaxa 4896 (3) on page 385, DOI: 10.11646/zootaxa.4896.3.3, http://zenodo.org/record/4383886, {"references":["Spence Bate, C. S. (1862) Catalogue of the Specimens of Amphipodous Crustacea in the Collection of the British Museum. British Museum (Natural History), London, iv + 399 pp., 56 pls. https: // doi. org / 10.5962 / bhl. title. 20901","Bellan-Santini, D. & Ruffo, S. (2003) Biogeography of benthic marine amphipods in Mediterranean Sea. Biogeographia-The Journal of Integrative Biogeography, 24 (1), 273 - 292. https: // doi. org / 10.21426 / B 6110176"]}

39. Centraloecetes dellavallei (Stebbing, 1899) Materials examined. 7 individuals; Stations Vas. 14(1), 26(3), 31(3). Distribution. Atlanto-Mediterranean (Bellan-Santini & Ruffo 2003; Christodoulou et al. 2013). The species is reported in Cyprus for the first time. In Vasiliko Bay it is considered as a rare species (FI: 7.14 %). Ecology. In Vasiliko Bay it was found at 22���25 m depth (24 + 2 m), in muddy sand substrate with organic matter ranging between 6.79���8.18 % (7.419 + 0.707 %) and in association with C. prolifera, C. c y lindracea and Di ctyo ta spp. It was found in stations classified according to the Bentix Index as in: ���good��� (2 stations, 4 individuals) and ���moderate��� (1 station, 3 individuals)., Published as part of Rousou, Maria, Plaiti, Wanda, Lowry, Jim, Charalambous, Stephanos & Chintiroglou, Chariton Charles, 2020, Amphipoda species (Suborders: Amphilochidea and Senticaudata) from Vasiliko Bay, Cyprus: New records, information on their biogeography and an annotated checklist from the coasts of Cyprus, pp. 373-408 in Zootaxa 4896 (3) on page 391, DOI: 10.11646/zootaxa.4896.3.3, http://zenodo.org/record/4383886, {"references":["Stebbing, T. R. R. (1899) Revision of Amphipoda (continued). Annals and Magazine of Natural History, Series 7, 4 (21), 205 - 211. https: // doi. org / 10.1080 / 00222939908678185","Bellan-Santini, D. & Ruffo, S. (2003) Biogeography of benthic marine amphipods in Mediterranean Sea. Biogeographia-The Journal of Integrative Biogeography, 24 (1), 273 - 292. https: // doi. org / 10.21426 / B 6110176","Christodoulou, M., Paraskevopoulou, S., Syranidou, E. & Koukouras, A. (2013) The amphipod (Crustacea: Peracarida) fauna of the Aegean Sea, and comparison with those of the neighbouring seas. Journal of the Marine Biological Association of the United Kingdom, 93 (5), 1303 - 1327. https: // doi. org / 10.1017 / S 002531541200183 X"]}

7. Guernea (Guernea) coalita (Norman, 1868) Materials examined. 57 individuals; Stations Vas. 1(2), 9(2), 17(5), 24(10), 29(19), 32(5), 37(14). Distribution. Atlanto-Mediterranean (Bellan-Santini & Ruffo 2003). The species is reported in Cyprus for the first time. In Vasiliko Bay it is considered to be a rare species (FI: 14.28 %). Ecology. In Vasiliko Bay it was found at depths of 6���12 m (10 + 2 m), mainly in sandy, and also muddy sand substrates, with organic matter ranging between 1.933 ���4.204 % (2.823 + 0.793 %) and in association with C. nodosa. It was found in stations classified according to the Bentix Index as in: ���high��� (3 stations, 17 individuals), ���good��� (3 stations, 26 individuals) and ���moderate��� (1 station, 14 individuals). It was recorded at the ports (14 individuals) and at the underwater outlet station (2 individuals)., Published as part of Rousou, Maria, Plaiti, Wanda, Lowry, Jim, Charalambous, Stephanos & Chintiroglou, Chariton Charles, 2020, Amphipoda species (Suborders: Amphilochidea and Senticaudata) from Vasiliko Bay, Cyprus: New records, information on their biogeography and an annotated checklist from the coasts of Cyprus, pp. 373-408 in Zootaxa 4896 (3) on page 383, DOI: 10.11646/zootaxa.4896.3.3, http://zenodo.org/record/4383886, {"references":["Norman, A. M. (1868) On Crustacea Amphipoda new to science or to Britain. Annals and Magazine of Natural History, Series 4, 2 (12), 411 - 421. https: // doi. org / 10.1080 / 00222936808695843","Bellan-Santini, D. & Ruffo, S. (2003) Biogeography of benthic marine amphipods in Mediterranean Sea. Biogeographia-The Journal of Integrative Biogeography, 24 (1), 273 - 292. https: // doi. org / 10.21426 / B 6110176"]}

3. Nototropis guttatus Costa, 1853 Materials examined. 4 individuals; Stations Vas. 21(2), 33(2). Distribution. Atlanto-Mediterranean, Black Sea (WoRMS). The species is reported in Cyprus for the first time. In Vasiliko Bay it is considered to be a rare species (FI: 4.76 %). Ecology. In Vasiliko Bay it was found at depths of 4���32 m (18 + 19 m), in muddy sand and sandy substrates, with organic matter ranging between 2.446 ���8.848 % (5.647 + 4.527 %) and in association with C. prolifera. It was found in stations classified according to the Bentix Index as in: ���high��� (1 station, 2 individuals) and ���moderate��� (1 station, 2 individuals). It was recorded at the power station outlet (2 individuals)., Published as part of Rousou, Maria, Plaiti, Wanda, Lowry, Jim, Charalambous, Stephanos & Chintiroglou, Chariton Charles, 2020, Amphipoda species (Suborders: Amphilochidea and Senticaudata) from Vasiliko Bay, Cyprus: New records, information on their biogeography and an annotated checklist from the coasts of Cyprus, pp. 373-408 in Zootaxa 4896 (3) on page 382, DOI: 10.11646/zootaxa.4896.3.3, http://zenodo.org/record/4383886, {"references":["Costa, A. (1853) Relazione sulla memoria del Dottor Achille Costa, di ricerche su' crostacei amfipodi del regno di Napoli. Rendiconto della Societa Reale Borbonica, Accademia delle Scienze, New Series, 2, 167 - 178. https: // doi. org / 10.5962 / bhl. title. 2070"]}

51. Megamphopus cornutus Norman, 1869 Materials examined. 2 individuals; Station Vas. 37(2). Distribution. Atlanto-Mediterranean (Bellan-Santini & Ruffo 2003), Black Sea (WoRMS). The species is reported in Cyprus for the first time. In Vasiliko Bay, it is considered as a rare species (FI: 2.38 %). Ecology. In Vasiliko Bay it was found in a port at 10 m depth, in muddy sand substrate with 4.204 % organic matter. The station was in ���high��� ecological status according to the Bentix Index., Published as part of Rousou, Maria, Plaiti, Wanda, Lowry, Jim, Charalambous, Stephanos & Chintiroglou, Chariton Charles, 2020, Amphipoda species (Suborders: Amphilochidea and Senticaudata) from Vasiliko Bay, Cyprus: New records, information on their biogeography and an annotated checklist from the coasts of Cyprus, pp. 373-408 in Zootaxa 4896 (3) on page 394, DOI: 10.11646/zootaxa.4896.3.3, http://zenodo.org/record/4383886, {"references":["Norman, A. M. (1869) Last report on dredging among the Shetland Isles. Part 2. On the Crustacea, Tunicata, Polyzoa, Echinodermata, Actinozoa, Hydrozoa, and Porifera. In: Murray J., Report of the thirty-eighth meeting of the British Association for the Advancement of Science, London, 1869, pp. 247 - 336.","Bellan-Santini, D. & Ruffo, S. (2003) Biogeography of benthic marine amphipods in Mediterranean Sea. Biogeographia-The Journal of Integrative Biogeography, 24 (1), 273 - 292. https: // doi. org / 10.21426 / B 6110176"]}

8. Iphimedia obesa Rathke, 1843 Materials examined. 1 individual; Vas.18(1). Distribution. Atlanto-Mediterranean (Bellan-Santini & Ruffo 2003). The species was reported from Cyprus by Kocatas et al. (2001). In Vasiliko Bay (current study), it is considered to be a rare species (FI: 2.38 %). Ecology. Iphimedia obesa was found in association with algae and soft-bottom sediments (mud, mud-and mixture) at 92 m depth (Kocatas et al. 2001). In Vasiliko Bay it was found at 51 m depth, in muddy sand substrate with organic matter being 5.621 %, in association with C. prolifera, C. cylindracea and C. nodosa. The station was in ���good��� ecological status according to the Bentix Index., Published as part of Rousou, Maria, Plaiti, Wanda, Lowry, Jim, Charalambous, Stephanos & Chintiroglou, Chariton Charles, 2020, Amphipoda species (Suborders: Amphilochidea and Senticaudata) from Vasiliko Bay, Cyprus: New records, information on their biogeography and an annotated checklist from the coasts of Cyprus, pp. 373-408 in Zootaxa 4896 (3) on page 384, DOI: 10.11646/zootaxa.4896.3.3, http://zenodo.org/record/4383886, {"references":["Rathke, H. (1843) Beitrage zur Fauna Norwegens. Nova Acta Academiae Caesareae Leopoldino-Carolinae Naturae Curiosorum, Breslau & Bonn, 20, 1 - 264. https: // doi. org / 10.5962 / bhl. title. 120119","Bellan-Santini, D. & Ruffo, S. (2003) Biogeography of benthic marine amphipods in Mediterranean Sea. Biogeographia-The Journal of Integrative Biogeography, 24 (1), 273 - 292. https: // doi. org / 10.21426 / B 6110176","Kocatas, A., Katagan, T. & Huseyin. A. B. (2001) Contribution to the knowledge of the crustacean fauna of Cyprus. Israel Journal of Zoology, 47 (2), 147 - 160. https: // doi. org / 10.1560 / YQL 8 - 4 PBT- 12 W 2 - 82 HV"]}

12. Idunella nana (Schiecke, 1973) Materials examined. 2 individuals; Station Vas. 31(2). Distribution. Mediterranean (Bellan-Santini & Ruffo 2003). The species is reported in Cyprus (Vasiliko bay) for the first time and it is considered to be a rare species (FI: 2.38 %). Ecology. In Vasiliko Bay it was found at 22 m depth in muddy sand substrate with organic matter being 7.284%, in association with C. prolifera and C. cylindracea. The station was in ���moderate��� ecological status according to the Bentix Index., Published as part of Rousou, Maria, Plaiti, Wanda, Lowry, Jim, Charalambous, Stephanos & Chintiroglou, Chariton Charles, 2020, Amphipoda species (Suborders: Amphilochidea and Senticaudata) from Vasiliko Bay, Cyprus: New records, information on their biogeography and an annotated checklist from the coasts of Cyprus, pp. 373-408 in Zootaxa 4896 (3) on page 385, DOI: 10.11646/zootaxa.4896.3.3, http://zenodo.org/record/4383886, {"references":["Schiecke U. (1973) Ein Beitrag zur Kenntnis der Systematik, Biologie und Autokologie mariner Peracarida (Amphipoda, Isopoda, Tanadaicea) des Golfes von Neapel. Inaugural Dissertation. Mathematik Naturwissenschaften Fakulte Christian-Albrechts Universite, Kiel, 408 pp.","Bellan-Santini, D. & Ruffo, S. (2003) Biogeography of benthic marine amphipods in Mediterranean Sea. Biogeographia-The Journal of Integrative Biogeography, 24 (1), 273 - 292. https: // doi. org / 10.21426 / B 6110176"]}