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The examination of 93 specimens of the Phalotris bilineatus group housed in the major Argentinian collections allowed us to recognize three additional species of this group. We propose the resurrection of two names, Phalotris spegazzinii and Phalotris suspectus, previously placed in the synonymy of P. bilineatus, and describe a new species from northwestern Argentina. The new species is recognized based on a unique combination of morphological characters. In addition, we redescribe the resurrected taxa (providing variation in color patterns, meristic and morphometric characters) and present new data on their geographic distributions., El examen de 93 ejemplares del grupo de Phalotris bilineatus depositados en las principales colecciones de la Argentina permitió reconocer tres especies adicionales del complejo. Proponemos la revalidación de dos nombres, Phalotris spegazzinii y Phalotris suspectus, previamente ubicados en la sinonimia de Phalotris bilineatus, y se describe una nueva especie del noroeste de la Argentina. La nueva especie se reconoce con base en una combinación única de caracteres. Adicionalmente, redescribimos los taxones revalidados (brindando información sobre patrones de coloración, caracteres morfológicos y merísticos) y presentamos nuevos datos sobre su distribución geográfica., Asociación Herpetológica Argentina

Genus Xenoxybelis Machado, 1993 resurrected Type species: Coluber argenteus Daudin, 1803, by original designation. Synonyms: Oxybelis Wagler, 1830; Philodryas Wagler, 1830. Content: (two species) Xenoxybelis argenteus (Daudin, 1803); Xenoxybelis boulengeri (Procter, 1923) species name revalidated. Diagnosis: Xenoxybelis can be distinguished from the other genera of Philodryadini by the following combination of characters: markedly elongated snout, forming an acuminated and sharp head shape; large number of prediastemal maxillary teeth (16-21), followed by one or two grooved postdiastemal teeth; short heart-shaped hemipenes with a well-defined papillate longitudinal crest, formed by the confluence of the body calyces, that runs medially in the asulcate surface; lateral surfaces of hemipenes covered with two to four rows of well-developed enlarged lateral spines (Fig. 4). Geographical distribution: Both species occur in the Amazonian region,from Colombia and Guianas to Bolivia and Paraguay (Cunha & Nascimento, 1978; Prudente et al., 2008). Etymology: The generic name of the South American Vine snake Xenoxybelis is formed by the Greek words Xenos- (adjective, ξέΝΟς, meaning "different"), -oxy- (adjective, Οξύς, meaning "sharp"), and -belos (noun, βέΛΟς, meaning "dart"), in reference to their external similarity with the unrelated Neotropical Vine snake genus Oxybelis Wagler, 1830. Comment: Zaher et al. (2009) and Grazziotin et al. (2012) placed Xenoxybelis and Pseudablabes under the synonymy of Philodryas. With this nomenclatural act, Philodryas boulengeri Werner, 1909 and Philodryas boulengeri (Procter, 1923) became secondary homonyms (Grazziotin et al., 2012),resulting in the proposition of the new replacement name georgeboulengeri for the latter species.According to our phylogenetic results, returning Philodryas georgeboulengeri (Procter,1923) to the revalidated genus Xenoxybelis eliminates the homonymy. Therefore, we revalidate the species name Xenoxybelis boulengeri (Procter, 1923)., Published as part of Arredondo, Juan Camilo, Grazziotin, Felipe G., Scrocchi, Gustavo J., Rodrigues, Miguel Trefaut, Bonatto, Sandro Luís & Zaher, Hussam, 2020, Molecular phylogeny of the tribe Philodryadini Cope, 1886 (Dipsadidae: Xenodontinae): Rediscovering the diversity of the South American Racers, pp. 1-14 in Papéis Avulsos de Zoologia (Pap. Avulsos Zool., S. Paulo) (Pap. Avulsos Zool., S. Paulo) 60 on page 8, DOI: 10.11606/1807-0205/2020.60.53, http://zenodo.org/record/4983748, {"references":["Machado, S. R. 1993. A new genus of Amazonian vine snake (Xenodontinae: Alsophiini). Acta Biologica Leopoldensia, 15: 99 - 108.","Wagler, J. G. 1830. Naturliches System der Amphibien, mit vorangehender Classification der Saugthiere und Vogel. Ein Beitrag zur vergleichenden Zoologie. Munchen, J. G. Cotta schen Buchhandlung. vi + 354 p.","Cunha, O. & Nascimento, E. F. 1978. Ofidios da Amazonia. X. - As cobras da regiao leste do Para. Publicacoes Avulsas Museu Paraense Emilio Goeldi, 31: 1 - 218.","Prudente, A. L. C.; Silva, M. A. A.; Rocha, W. A. & Franco, F. L. 2008. Morphological variation in Xenoxybelis boulengeri (Procter, 1923) (Serpentes, Xenodontinae, Philodryadini). Zootaxa, 1743: 53 - 61.","Zaher, H.; Grazziotin, F. G.; Cadle, J. E.; Murphy, R. W.; Moura-Leite, J. C. & Bonatto, S. L. 2009. Molecular phylogeny of the advanced snakes (Serpentes, Caenophidia), with an emphasis on South American Xenodontines: a revised classification and descriptions of new taxa. Papeis Avulsos de Zoologia, Sao Paulo, 49 (11): 115 - 153.","Grazziotin, F. G.; Zaher, H.; Murphy, R. W.; Scrocchi, G.; Benavides, M. A.; Zhang, Y. P. & Bonatto, S. L. 2012. Molecular phylogeny of the new world Dipsadidae (Serpentes: Colubroidea): areappraisal. Cladistics, 28: 437 - 59."]}

El patrón reproductivo es una de las principales características de la historia de vida de los organismos. Este patrón depende tanto de historia evolutiva de los organismos como de los factores externos. Se estudió la reproducción de Liolaemus espinozai en Campo El Arenal, Catamarca. Se analizó el ciclo reproductivo y de los cuerpos grasos de machos y hembras y sus relaciones con la temperatura, fotoperíodo y precipitación. La especie es ovípara y exhibe un patrón reproductivo estacional típico de primavera - verano, con ovulación y fecundación de primavera y puesta y eclosión de verano. La actividad reproductiva de las hembras empieza en noviembre con folículos vitelogénicos; desde diciembre pueden observarse huevos en oviducto y a mediados de enero hasta principios de febrero aparecen hembras con oviducto distendido. Los machos presentan actividad desde fines de septiembre con máximo volumen testicular hasta noviembre, luego estos valores descienden hasta su expresión mínima entre enero y marzo. El tamaño de la puesta varió de 2 a 7 huevos y se correlacionó con el tamaño de la hembra indicando una tendencia a que individuos mayores tengan posturas mayores. La especie muestra un patrón con características similares a lo encontrado para especies filogenéticamente cercanas. The reproductive pattern is a characteristic of major importance to understand the life history of a species, which depends of evolutionary history and ecology. We described the reproduction. We analyzed the reproductive and fat bodies cycle in males and females, and its relation with temperature, photoperiod and precipitation. Liolaemus espinozai is an oviparous species with a typical seasonal Spring-Summer reproductive pattern, starting its ovulation and fecundation in Spring, and its oviposition and hatching in Summer. Female reproductive activity begins with yolkled follicles in November; from December to middle January oviductal eggs can be observed and from middle January to early February females show distended oviduct. Male activity begins at the end of September, its testicle volume increases, and reaches the maximum in November. Later, the volume decreases and is minimal between January and March. The number of eggs varies from 2 to 7 and is correlated with the SVL, indicating that bigger females have more eggs. The pattern is similar to that of phylogenetically closer species.

Bothrops sonene sp. nov. Figures 4 –7, Table 1 Bothrops gr. neuwiedi: Cadle et al. 2002: 102 (MUSM 33453, from Pampa Juliaca, department of Madre de Dios, Peru). Holotype. An adult female, MUBI 12319, collected on April 23, 2013 by J.A. Ochoa at Pampa Juliaca (12°57’46.79” S, 68°55’22.61” W; 206 m above sea level, asl hereafter), Pampas del Heath (Bahuaja-Sonene National Park), district of Tambopata, province of Tambopata, department of Madre de Dios, Peru (Figs. 5A, 5E, 6B). Paratypes. Three specimens, all from Pampa Juliaca, Pampas del Heath (Bahuaja-Sonene National Park), district of Tambopata, province of Tambopata, department of Madre de Dios, Peru. An adult male, MUBI 14679, collected on August 14, 2015 by J.A. Ochoa, G. Sebastián & K. Huamani (12°57’47.47” S, 68°54’57.01” W; 210 m asl) (Figs. 5B, 5F, 6C). An immature male, MUSA 4350, collected on November 13, 2015 by R. Santa Cruz (12°57’2.47” S, 68°54’51.05” W; 209 m asl) (Figs. 4, 5C, 6A, D). An immature male, MUSM 33453, collected on between June 02–25, 1996 by J. Cadle (12°59’20.10” S, 68°55’48.45” W; 208 m asl) (Fig. 5D). Diagnosis. Bothrops sonene can be distinguished from its congeners by the exclusive combination of the following morphological characters: dorsal color pattern of body consisting of small C-shaped blotches, apparently unique for the genus; postocular stripe originating posteriorly to the eye, two or two and a half scales width, covering three or four posterior supralabials; dorsum of the head with paired markings arranged symmetrically; venter cream heavily speckled with brown; lacunolabial scale absent, prelacunal discrete contacting second supralabial; prefoveals three to five; subfoveal single and usually present; postfoveals absent to two; canthals two; intersupraoculars seven; suboculars one or two; postoculars two or three; supralabials seven or eight; infralabials nine to eleven; interrictals 26–27; middorsal scales 23–25; ventrals 172 in the female, 169–173 in males; subcaudals divided, 45 in the female, 50 in males. Comparisons. Bothrops sonene is easily distinguished from its congeners by its unique dorsal color pattern of relatively small and C-shaped blotches. It is distinguished from B. atrox, B. barnetti, B. jararaca, B. jararacussu, B. pictus and B. taeniatus by the absence of a lacunolabial scale (= prelacunal fused with second supralabial), a condition shared with the species of the B. alternatus and B. neuwiedi groups. Similar C-shaped (or in the shape of a headphone) dorsal blotches are present in B. alternatus, B. cotiara, B. fonsecai, B. itapetiningae and B. jonathani (B. alternatus group excluding B. ammodytoides, whose dorsal blotches are quadrangular); however, the blotches of B. sonene are smaller and rounder than those species, the lower set of blotches are well separated from each other (except in B. itapetiningae), and blotches are bordered with black (vs. bordered with white). The new species can also be distinguished from some species of the B. alternatus group by the postocular stripe starting posteriorly to the eye (vs. below the eye in B. alternatus, B. jonathani and B. ammodytoides), number of prefoveals (3–5 vs. 3– 15), subfoveals (0–1 vs. 1–6), supralabials (8 vs. 8–12) and infralabials (9–11 vs. 10–15), length of hemipenes (relatively short vs. long), and fusiform and divergent hemipenial lobes (vs. parallel and subcylindrical lobes). Bothrops sonene is more similar to the B. neuwiedi group of species than to the rest of the species of the genus. They all have a symmetrical pattern of blotches in the dorsum of the head, a brown and relatively wide postocular stripe that extends posterior to the rictal region, a discrete prelacunal scale contacting the second supralabial, and similar number of prefoveals, subfoveals, supralabials and infralabials. They also bear resemblance in hemipenial morphology; the overall morphology of the hemipenis of the new species is identical to that of the species of the B. neuwiedi group, which is characteristic of that group. Differences in coloration and scalation between Bothrops sonene and the species of the B. neuwiedi group are summarized in Table 2. Description of the holotype. Adult female; snout-vent length 950 mm; tail length 123 mm; caudal spine length 8.9 mm; head length 44.8 mm; maximum head width 36.5 mm; interorbital space 7.2 mm; internasal space 2.4 mm; dorsal scale rows 23–24–19; ventral scales 172; subcaudal scales 45/45; rostral trapezoidal; canthus rostralis elevated forming a ridge; internasals 1/1; canthals 1/1, similar size than internasals; intercanthals, keeled, 4–5–5; intersupraoculars, keeled, 7; scales around supraoculars 11/10; supralabials 8/7; infralabials 10/11; postnasal in contact (left side) or not in contact (right side) with first supralabial; loreal subtriangular; prefoveals 5/ 5; subfoveal 1/0; postfoveals 1/1; prelacunal contacting the second supralabial; sublacunals 1/1; supralacunals 1/1; suboculars 2/1; postoculars 3/2; upper preoculars contributing to the canthus rostralis, right preocular divided; upper and medial preoculars in contact with the orbit; inferior preoculars not in contact with the orbit; scales between suboculars and 3 rd –4 th supralabials 2/2; scales between suboculars and 4 th supralabial 2/2; scales between suboculars and 4 th –5 th supralabials 2/2; interrictals 26; gulars 4. Dorsal background color of body light brown; dorsum with dark brown, black-edged blotches, each blotch composed of dorsal and ventral C-shaped forms; ventral surface of body cream, heavily speckled with dark brown spots; dorsum of head with symmetrically arranged brown and dark-edged blotches, including a single blotch on the snout, a pair of blotches near supraoculars, a pair of parietal stripes that extend posterior-laterally, and a pair of occipital stripes that extend posteriorly; head with lateral, brown and black-bordered postocular stripes originating posterior to each eye. Color pattern variation. Dorsal background color of head and body light brown with olive green tonality in live specimens; dorsum of body with 28–31 C-shaped, dark brown and black-edged, dorsal blotches; ventral portion of blotches formed by a pair of mostly-fused round blotches, forming a single curved blotch; ventral surface of body cream, speckled with dark brown spots, the spots forming a transversal stripe in some ventral scales; dorsum of head with brown, dark-edged, symmetrically arranged blotches; postocular stripes extend 1.5–3 scales posterior to rictal region, encroaching 3–4 supralabials and 1–2 infralabials; supralabials with dark brown spots, more conspicuous between 3 rd –5 th supralabials; infralabials mostly bordered with dark brown pigments. Gular region of males cream with irregular dark brown markings, absent in the female of the type-series suggesting a putative sexual dimorphism in this character; tip of tail pale ventrally in adult female and male (MUBI 14679); both dorsal and ventral surface of tip of tail pale in subadult males (MUSA 4350, MUSM 33453). Hemipenis morphology. Based on MUBI 14679 and MUSA 4350. Organ strongly bilobed; hemipenial lobes fusiform and divergent, 67% of total hemipenial length; hemipenial body 33% of total hemipenial length; capitulum longer on sulcate side, occupying 62% of lobe; base of capitulum covered with spinulate calyces; hookshaped spines distributed asymmetrically on lobes, big sized spines located proximally; small, thin and curved spines present in the intralobular region, located distally on the lobes; hemipenial body covered with spinules; microornamentation on intrasulcar region absent; sulcus spermaticus bifurcating proximal to capitulum and ornamented with spinules (Fig. 7). Distribution and natural history. Known to occur in Pampas del Heath, in the Bahuaja-Sonene National Park, province of Tambopata, department of Madre de Dios, Peru, at between 206–210 m asl (Fig. 2). The area is situated in southeastern Peru, near the border with Bolivia, between the Tambopata and Heath Rivers. Bothrops sonene is known from four close localities situated inside of the savannas of the Peruvian Pampas del Heath. The specimens were collected at the end of the rainy season and during the dry season. They have diurnal activity; during the day they were observed on the ground and above grass at 500 mm above ground, sometimes slithering rapidly. They are associated with water bodies, which are widespread along the savannas of Pampas del Heath in rainy seasons (the soil is flooded up to 250 mm from the ground) and restricted to the “Aguajales” (Palm Tree areas) in dry seasons (Fig. 8). Etymology. The specific epithet “sonene” is derived from the tacana language, and is the name used by the Ese’ejja for the Heath River. The Ese’ejja are an Amazonian community that inhabits the areas surrounding the Madre de Dios, Tambopata, and Heath Rivers, in the border between Peru and Bolivia. We named the new species “sonene” because its type locality, the Peruvian Pampas del Heath, is located near the Heath River. B. sonene B. cf. sonene B. neuwiedi B. aff. neuwiedi B. mattogrossensis B. aff. B. aff. B. aff. mattogrossensis 1 mattogrossensis 2 mattogrossensis 3 BP C-shaped blotches Trapezoidal or C- Trapezoidal or Trapezoidal or Trapezoidal Trapezoidal Trapezoidal Trapezoidal shaped blotches subtriangular subtriangular blotches blotches blotches blotches blotches blotches PM Stripes Stripes or rounded Stripes Stripes Rounded blotches Rounded blotches Rounded blotches Rounded blotches blotches POSw 2–2.5 2 – 3 2.5 – 3 1.5 – 3 2 – 2.5 2 – 3 1.75 – 3 2 – 2.5 x=2.2 x=2.4 x=2.8 x=2.3 x=2.2 x=2.2 x=2.2 SLPOS 3–4 3 – 4 2 2 – 3 3 – 4 2 – 4 2 – 4 2 – 3 x =3.2 x=3.3 x= 2 x =3.8 x= 3 x = 3 x =2.7 V 164–172 173 – 178 166 – 172 172 – 184 183 – 190 163 – 173 155 – 165 167 – 177 x=169.5 x=175 x=169 x=177.9 x=186.5 x=167.5 x=160 x=173 MD 23–25 25 – 27 25 – 27 24 – 28 23 – 25 21 – 25 21 – 23 21 – 25 x = 24 x = 26 x =26.3 x=25.7 x=23.5 x= 23 x =22.6 x=22.8 SL 8 8 – 9 8 – 11 8 – 9 7 – 10 7 – 8 7 – 8 7 – 9 x = 8 x = 9 x =8.4 x=8.3 x=7.6 x=7.6 x=8 IL 10–11 10 – 11 11 – 13 10 – 13 11 – 12 9 – 11 8 – 10 9 – 12 x=10.5 x= 10 x =11.7 x= 11 x =11.5 x= 10 x =9.4 x=10.8 ICm 4–6 4 – 6 6 4 – 6 3 – 4 3 – 5 3 – 5 3 – 7 x =5.2 x=5.2 x= 5 x =3.7 x=3.7 x=4.2 x=5 ISPO 7 7 8 – 11 7 – 11 5 – 8 5 – 7 4 – 7 6 – 8 x =9.3 x= 8 x = 7 x =6.3 x=5.6 x=7.3 PF 3–5 4 5 – 7 2 – 6 3 – 6 2 – 5 2 – 4 4 – 5 x = 4 x =6.3 x=3.9 x=4.5 x=3.2 x=3.2 x=4.6 SBF 1 0 – 1 1 – 3 0 – 2 0 – 2 0 – 1 0 – 1 1 – 3 x =0.3 x= 2 x = 1 x =0.8 x=0.5 x=0.6 x=1.6 ……continued on the next page TABLE 2. (Continued) TABLE 2. Continues. ……continued on the next page TABLE 2. (Continued)

The feeding habits of a population of Liolaemus laurenti in northwest Argentina were studied in an annual period of activity. The objectives of the study were to determine the composition of the diet of the species, the trophic diversity and variations between sexes and among dry and wet seasons. From the analysis of 42 stomach contents, it was determined that ants constitute the main prey category of the diet, they comprise 93.15% of all present prey; it was also the most frequent category (100% of stomach contents). Lizard population also feeds on other insects and chelates. The trophic diversity was higher in females and in both in males and females increases during the wet season. Seasonal differences could be attributed to the fact that in arid ecosystems rainfall strongly affects productivity primary. Therefore the supply of resources for arthropods in general and therefore, for their predators., Asociación Herpetológica Argentina

Apostolepis multicincta, a fossorial snake known only from Bolivia, is recorded in Argentina, extending its known range 450 km south of the previous closest record.

Taxonomy, assessment information, geographic range, distribution map, population, habitat and ecology, use and trade, threats and conservation actions of Cercosaura steyeri., Facultad de Ciencias Naturales y Museo

Hemos revisado las colecciones del Instituto de Herpetología de la Fundación Miguel LiIlo, Tucumán (FML), del Museo Argentino de Ciencias Naturales "Bernardino Rivadavia". Buenos Aires (MACN), de la Cátedra de Anatomía Comparada de la Universidad Nacional del Nordeste (UNNEC) y de la Colección "Félix de Azara", actualmente depositada en el Museo Argentino de Ciencias Naturales "Bernardino Rivadavia". En base a estos datos confirmamos la presencia de la especie en Santa Fe y brindar la distribución exacta conocida en Argentina. Además, describimos con mis exactitud el rango de variación de los principales caracteres de escamación y describimos los hemipenes evertidos, corrigiendo algunos errores de la descripción existente.

Se presenta los principales caracteres y la distribución de Psomophis genimaculatus y P. obtusus, las dos especies del género presentes en Argentina. Se discute brevemente la variación de los caracteres con respecto a la descripción original. The taxonomic characters and geographic distribution of the two species of the genus Psomophis presents in Argentina are described. The variation of characters is discussed briefly.

El 3 de febrero de 2005 murió en San Miguel de Tucumán el Dr. Raymond Ferdinand Louis-Philippe Laurent, a quien la herpetología y los herpetólogos debemos tanto. Al recordarlo aquí celebramos su vida, que tuvo visos de aventura y con ese espíritu fue vivida hasta el final.

A más de una década de la primera Lista Roja de herpetofauna amenazada propuesta por la Asociación Herpetológica Argentina (AHA 2000), se recategorizaron las serpientes a partir de nueva información taxonómica, biogeográfica y bio-ecológica, además de modificaciones metodológicas respecto a la evaluación anterior. Mediante la participación de 18 especialistas de toda la Argentina se reevaluaron 136 taxones de serpientes (130 en la anterior) incluyendo varios cambios taxonómicos (8 taxones nuevos para Argentina y 2 sinonimizados), obteniéndose como resultado la inclusión de 49 especies en la lista roja (5 En Peligro, 17 Amenazadas, 27 Vulnerables), 15 Insuficientemente Conocidas y 72 No Amenazadas. En relación con la categorización anterior de la AHA: un taxón descendió de Vulnerable a No Amenazado, 11 No amenazados y 4 Insuficientemente Conocidos fueron elevados a distintas categorías de amenaza, 7 taxones Vulnerables fueron elevados a Amenazados, un taxón fue elevado de Amenazado a En Peligro. De 8 taxones no evaluados en 2000, uno categorizó No Amenazado, 4 Insuficientemente Conocidos, uno Vulnerable y 2 Amenazados. Estas modificaciones son el resultado de: (1) Mayor información sistemática, biogeográfica y bio-ecológica disponible para la evaluación; (2) Cambios en cuanto a las presiones antrópicas sobre las especies o sus hábitats; (3) Modificaciones metodológicas que incluyeron instructivos para aplicar los conceptos, la discusión y consenso entre especialistas y el análisis de las incertidumbres., After more than a decade from the first red list of threatened herpetofauna proposal by the Asociación Herpetológica Argentina (2000), we re-categorized snakes from new taxonomic, biogeographical and bio-ecological information as well as methodological changes in the former evaluation. Through the participation of 18 specialists from all over Argentina, 136 taxa of snakes (130 in the previous) were re-evaluated including several taxonomic changes (8 new taxa added to Argentina, and 2 sinonimies). The results were the inclusion of 49 species in the red list (5 Endangered, 17 Threatened, 27 Vulnerable), 15 Insufficiently Known and 72 Not Threatened. Compared to the former categorization of the AHA: one taxon descended from Vulnerable to Not Threatened, 11 Not Threatened and 4 Insufficiently Known were elevated to different categories of threat, 7 taxa were elevated from Endangered to Vulnerable, one from Vulnerable to Endangered. From the 8 taxa not evaluated in 2000, one categorized Not Threatened, 4 Insufficiently Known, one Vulnerable, and 2 Threatened. These changes are the result of: (1) increased systematic, biogeographical and bio- ecological information available for the evaluation, (2) Changes in human pressures on the species or their habitats, (3) methodological changes that included recommendations to apply concepts, discussion and consensus among specialists and the analysis of uncertainties., Asociación Herpetológica Argentina (AHA)

Liolaemus goetschi M��ller and Hellmich, 1938 (Fig. 1���3) 1938. Liolaemus goetschi. M��ller and Hellmich. Zoologischer Anzeiger, 123 (5���6): 130���142. 1977. Liolaemus melanops. Cei, J. M. and J. A. Scolaro. Physis (Buenos Aires), 36: 225���226. 2003. Liolaemus goetschi. Cei and Scolaro. Facena, 19: 163. 2006. Liolaemus goetschi. Avila et al. Biol. J. Linnean Soc., 89: 241���275. 2007. Liolaemus goetschi. Abdala. Zootaxa, 1538: 1���84. 2008. Liolaemus goetschi. Pincheira Donoso, et al. Zootaxa, 1800: 1���85. 2010. Liolaemus goetschi. Nori et al. Check List, 6 (1): 6 - 4. Holotype. ZSM 4. Adult male, Laguna Playa, approximately 20 km north of General Roca, R��o Negro Province, Argentina, W. Goetsch, 1938. Paratype. ZSM 5. Adult female, Laguna Playa, approximately 20 km north of General Roca, R��o Negro Province, Argentina, W. Goetsch, 1938. Type locality. In the original description of Liolaemus goetschi, the authors designated Laguna Playa as the type locality of L. goetschi (approximately 20 km north of General Roca city, the site where Prof. Dr. Goetsch collected all the individuals corresponding to the type series) as the type locality of L. goetschi. Because the toponym ���Laguna Playa��� is not included in maps and records of the area and is unknown to local people, we conducted extensive surveys among rural people to locate the site. Finally, an elderly inhabitant of the nearest city (General Roca) informed us about the place and following his directions, we were able to find the exact place, located at 30 �� 52 ��� 59 ��� S and 67 �� 34 ��� 08��� W. The Liolaemus goetschi individuals were collected in subsequent intensive field surveys. Diagnosis. Liolaemus goetschi belongs to the L. boulengeri group because of the presence of the femoral patch of enlarged scales in the posterior surface of the femur (Etheridge 1995). Within this group, L. goetschi is distinguished from the L. anomalus group because of the hypertrophy of the puboischiotibialis muscle (Abdala et al. 2006), higher percentage of tail autotomy, tail longer than snout-vent length (SVL), head longer than wide, outer cilliaries not projecting and higher number of precloacal pores in males. It also differs from the species of the L. wiegmannii group in having a row of lorilabials between subocular and supralabials and four scales surrounding the mental (Etheridge 2000). Liolaemus goetschi differs from species of the L. laurenti group (Abdala 2007), L. abaucan, L. albiceps, L. calchaqu��, L. chacoensis, L. crepuscularis, L. darwinii, L. espinozai, L. grosseorum, L. irregularis L. koslowskyi, L. laurenti, L. lavillai, L. olongasta, L. ornatus, L. quilmes and L. uspallatensis in having posterior teeth with strongly cusped crowns and expanded margins, and in the presence of barely evident sexual dichromatism (Abdala 2007). Females of L. goetschi lack precloacal pores; this character distinguishes the species from L. albiceps, L. calachaqui, L. crepuscularis, L. irregularis, L. lavillai and L. ornatus because a high percentage of females of all these species have 1���6 precloacal pores (Abdala 2007). Within the L. melanops group (Abdala 2007), L. goetschi differs from L canqueli, L. fitzingerii, L. melanops, L. rothi, L. sagei, and L. xanthoviridis in having a shorter snout-vent-length (SVL) (max SVL 74.25 mm vs. 89.00��� 106.00 mm) and because of the presence of pre and postscapular spots (absent in these species). It also differs from L. canqueli and L. melanops because of the abscence of the cephalic melanism typical of these species It differs from L. rothi and L. sagei in the more conspicuous gular melanism (Abdala 2007) and from L. morenoi in its smaller size and lower number of scales around the midbody (72���85 Mean = 79.00 vs. 62���72 Mean = 66.35). Liolaemus goetschi differs from Liolaemus inacayali by the presence of pre and postscapular spots and a lower number of scales from occiput to tigh (73���84, Mean = 77.50 vs. 83���96, Mean = 88.75) (Abdala 2003). It differs from L. cheuachekenk in having a shorter maximum SVL (74.25 mm vs. 98.30 mm), in the presence of four to six scales in contact with the mental scale (L. cheuachekenk always presents four), in the absence of abdominal and pectoral melanism and in the presence of a different color pattern (Avila et al. 2008). L. goetschi differs from L. puelche in its smaller size (max SVL 74.25 mm vs. 89.00 mm in L. puelche), lower number of scales around midbody (62���72, Mean = 66.35 vs. 67���76, Mean = 70.75), gular melanism and two series of well defined black paraventral spots, absent in L. puelche (Avila et al. 2007). Within the Liolaemus cuyanus clade (Abdala 2007), composed of L. mapuche, L. cuyanus, and L. donosobarrosi, L. goetschi differs from the first two species in having smaller SVL (max SVL 74.25 mm vs. 79.00 and 102.00 mm, respectively) and a clearly different color pattern that never has a black antehumeral arch (Abdala 2002). It differs from L. donosobarrosi in having longer SVL (max SVL 74.25 mm vs. 60.80 mm), a lower number of scales around the midbody (62���72, Mean = 66.30 vs. 79���95, Mean = 85.40) and a different color pattern (Abdala 2005, 2007). Liolaemus goetschi differs from L. boulengeri, L. josei, L. loboi, L. senguer, L. tehuelche and L. telsen, in having a faint prescapular spot and a larger postscapular spot; two black spots usually band -or line- shaped on each side of the gular region; belly in males and females white or light pink, never yellow, red or bright blue; anterior throat in males never melanic and sexual dichromatism absent or barely marked. It also differs from L. martorii in having a longer SVL (max SVL 74.25 mm vs. 67.10 mm), and four to six scales in contact with mental scale (L. martorii always has four); L. goetschi also has a higher number of light blue scales in tail and body; also scapular spots and spots on the sides of body that are larger and more marked (Table 1). Character L. goetschi L. martorii L. melanops External morphology. Maximum SVL 74.25 mm (56.20 mm ��� 74.25 mm; Mean = 65.80 mm); tail length (93.00 mm ��� 105.90 mm; Mean = 98.20 mm); 1.4 times longer than SVL. Head 1.28 times longer (12.10 mm ��� 17.55 mm; x = 14.54 mm) than wide (9.20 mm ��� 12.55 mm; Mean = 11.31 mm). Head height (6.70 mm ��� 14.10 mm, Mean = 8.59 mm). Humerus length (7.10 mm ��� 10.70 mm, Mean = 8.27 mm). Radius length (6.60 mm ��� 9.20 mm; Mean = 8.11 mm). Hand length (7.60 mm ��� 9.90 mm, Mean = 8.26 mm). Tibial length (10.00 mm ��� 14.15 mm, Mean = 11.68 mm). Dorsal head scales smooth, convex and bulged. Scale organs distributed in postrostrals, internasals, frontonasals and prefrontals. Rostral pentagonal wider than long. Two postrostrals in contact with anterior supralabials. 15���18, (Mean = 16.25) dorsal head scales, from rostral to occiput (Hellmich Index). Nasal scales subpentagonal and in dorsolateral position. Eigth to 12 scales surrounding nasal. Frontal scale irregularly shaped, slightly longer than prefrontals. Interparietal pentagonal with a conspicuous pineal eye and surrounded by seven scales. Two parietals of irregular shape and larger than interparietals. Interparietal in contact with 6���10 (Mean = 7.30) scales. 9���14 (Mean = 12.30) circumorbitals. 6���8 (Mean = 7.15) laterally expanded supraoculars. Two rows of small scales between supraoculars and supercilliaries. 6���10 (Mean = 7.80) elongated and overlapped supercilliaries. One preocular longer than wide; one elongated subocular and one small postocular. A single row of 6���7 (Mean = 6.9) convex lorilabials. Supralabials 7���10 (Mean = 8.50) subquadrangular and elongate. Temporal scales 17���22 (Mean = 19.10) smooth and juxtaposed. Auditory meatus 1.86 times higher (2.30 mm ��� 4.00; Mean = 3.08) than wide (1.30 mm ��� 2.10 mm; Mean = 1.68 mm) surrounded by small granular scales. Mental scale pentagonal, wider than high in contact with 4���6, (Mean = 4.70) scales. Infralabials, 6���8,(Mean = 6.80) large and subquadrangular. Gular scales 30���36 (Mean = 32.80) smooth and imbricate. Y-shaped horizontal fold. Dorsal scales at midbody subtriangular, moderately imbricate and keeled, arranged in 22 longitudinal rows. Ventral scales along body midline: 79���89 (Mean = 74.15), smooth and moderately imbricate, subtriangular, of equal size or slightly larger than dorsal scales at midbody. 11���15 pigals (Mean = 12.55). 4���8 (Mean = 4.70) precloacal pores in males, absent in females. 62���72 (Mean = 66.35) scales around midbody. 68���78 (Mean = 72.70) dorsal scales of midbody from occiput to hind limbs. Suprahumerals and supraradials keeled and imbricate. Supratarsals smooth and imbricate. Infratarsals strongly imbricate. Supracarpals smooth and imbricate. Infracarpals keeled and imbricate. Supradigital lamellae smooth and imbricate. Infradigital lamellae tricarinate and imbricate. Fourth finger with 17���22 (Mean = 19.30) scales. Suprafemorals imbricate and rhomboidal. Infrafemorals smooth, imbricate and rhomboidal. Supratibials keeled and imbricate. Bulged femoral patch with 20���25 scales. Infratibials and supradigitals smooth and imbricate. Infradigitales tricarenate. Coloration. Males and females with slightly different coloration pattern. In most males, numerous light blue scales can be observed on the sides of tail and fewer on the sides of body and head. In females, these colors are absent in the scales. Dorsal color of females is more yellowish and usually white immaculate ventrally. Dorsum of head brown, with irregularly spread black spots. Nasal scale slightly lighter. A black band extends from the posterior zone of the eye to the upper lip margin. This band is bordered anteriorly and posteriorly by white spots. Also, a dark line posterior to the eye is present. Dorsal background coloration light brown. Also, has pre and postscapular spots, the latter larger than the former. Four series of black subquadrangular spots are present, two paravertebral and two lateral; both edged posteriorly by a white band. Nine to 12 black paravertebral spots are present, which are larger and more marked than the lateral spots. Paravertebral spots extend to the dorsal tail proximally to fuse, forming irregular spots less marked than those in the body. Anteriorly to the paravertebral spots, a reddish subquadrangular spot may be present. In many specimens, the vertebral region has isolated yellowish scales. Sides of body brown, slightly lighter than dorsum. Lateral region of tail and dorsal region of limbs light brown, similar to that of the body sides and slightly lighter than dorsum. Background of throat, chest and belly white or light pink. Ventral gular region is melanic or dark, reticulate anteriorlly, variegated with gray to black lines. Natural history. This is a typical psammophylus lizard of the Monte phytogeographical region (Cabrera, 1976). Vegetation in the area is typical of the Monte, with dominance of Atriplex lampa, Cercidiurn praecox, Larrea divaricata, L. n��tida and L. cuneifolia. Liolaemus goetschi seems to be most related to Larrea species. The species is difficult to observe and capture; individuals were found mainly at the time of the day when sand surface temperatures reach the highest values, between 13:00 and 15:00 hours. It is a relatively quiet species that, when chased, tends to hide and stay still or seek refuge in small caves. Most of the species studied in the group are oviparous and omnivorous (Abdala 2007); we suppose that this species shares the same characteristics. During field work, we observed some specimens feeding on ants and insect larvae. Distribution (Fig. 4). Liolaemus goetschi inhabits the province of R��o Negro. This species was found in its type locality (Laguna Playa), in Cerro Policia Town (Nori et al. 2010) and at 20 and 50 km south of the locality of Catriel. The specimens collected south of Catriel did not show morphological differences that enabled us to assume that they belong to a species different from the species of those specimens collected in Laguna Playa. All the indivuals are representatives of Liolaemus goetschi. M��ller and Hellmich (1938) mentioned the existence of Liolaemus populations south of Catriel that might correspond to the same species. This assumption is supported by the specimens studied in the present work. Phylogenetic position. The results of the phylogenetic analysis based on the morphological data matrix provided by Abdala (2007) and the incorporation of the 128 characters of Liolaemus goetschi from the type locality are presented in Figure 5. Here Liolaemus goetschi is included in the L. telsen group, which comprises: L. boulengeri L. goetschi, L. josei, L. loboi L. martorii, L. tehuelche, L. telsen and L. senguer. In our analysis, L.goetschi and L. martorii appear as sister species; this hypothesis suggests a close phylogenetic relationship between the two species. L. telsen group is part of the L. melanops group, which is included in the L. chacoensis group, all of them within the L. boulengeri group., Published as part of Nori, Javier, Abdala, Cristian Sim��n & Scrocchi, Gustavo Jos��, 2010, Liolaemus goetschi (Iguania: Liolaemidae): redescription and phylogenetic relationships within the L. boulengeri group, pp. 49-59 in Zootaxa 2440 on pages 51-56, DOI: 10.5281/zenodo.194953, {"references":["Muller, L. & Hellmich, W. (1938) Liolaemus - Arten aus dem Westlichen Argentinien (Ergebnisse der Argentinienreise von Prof. Dr. W. Goetsch, Breslau). I. Liolaemus darwini (Bell) und Liolaemus goetschi n. sp. Zoologischer Anzeiger, 123 (5 - 6), 129 - 142."]}

Describimos tres nuevas especies de Liolaemus del Sur de Argentina. Estas nuevas especies pertenecen al grupo de Liolaemus elongatus, grupo monofilético incluido en el subgénero Liolaemus sensu stricto. Dos de las nuevas especies habitan en el sur de la Provincia de Mendoza y la otra en el centro-oeste de la provincia de Río Negro. Las nuevas especies presentan estados de caracteres que las diferencian de todos los miembros del grupo de L. elongatus. Esos estados de carácter incluyen al patrón de coloración, número de escamas en el dorso, forma de las escamas temporales y tamaño (entre otros). Aunque con estas nuevas especies se dilucida el status taxonómico de varias poblaciones pertenecientes al grupo de L. elongatus, muchas quedan inciertas aún. We describe three new species of Liolaemus from Southern Argentina. These new species belong to the monophyletic Liolaemus elongatus group, included within the subgenus Liolaemus sensu stricto. Two of the new species inhabits in southern Mendoza Province and the other one inhabits in central west Río Negro Province. The new species exhibit character states that distinguish they from all other members of the L. elongatus group. Those character states include color pattern, number of scales in dorsum, shape of temporal scales, and size (among others). Although with those new species we clarify the taxonomic status of several populations which belongs to the L. elongatus group, many remain unknown. Fil: Abdala, Cristian Simón. Fundación Miguel Lillo. Dirección de Zoología; Argentina. Consejo Nacional de Investigaciones Científicas y Técnicas. Centro Científico Tecnológico - Tucumán. Unidad Ejecutora Lillo; Argentina Fil: Quinteros, Andres Sebastian. Universidad Nacional de Salta; Argentina. Consejo Nacional de Investigaciones Científicas y Técnicas. Centro Científico Tecnológico Conicet - Salta. Instituto de Bio y Geociencias del NOA. Universidad Nacional de Salta. Facultad de Ciencias Naturales. Museo de Ciencias Naturales. Instituto de Bio y Geociencias del NOA; Argentina Fil: Stazzonelli Sadir, Juan Carlos. Universidad Nacional de Tucumán. Facultad de Ciencias Naturales e Instituto Miguel Lillo; Argentina Fil: Scrocchi Manfrini, Gustavo Jose. Fundación Miguel Lillo. Dirección de Zoología; Argentina. Consejo Nacional de Investigaciones Científicas y Técnicas. Centro Científico Tecnológico - Tucumán. Unidad Ejecutora Lillo; Argentina

En este trabajo se presenta información sobre meses de puesta y eclosión, tamaño de la camada y medidas de huevos y neonatos de los siguientes ofidios ovíparos neotropicales: Boiruna maculata, Clelia rustica, Liophis almadensis, Liophis ceii, Oxyrhopus rhombifer, Philodryas patagoniensis, Philodryas baroni y Waglerophis merremii., Asociación Herpetológica Argentina (AHA)

El 3 de febrero de 2005 murió en San Miguel de Tucumán el Dr. Raymond Ferdinand Louis-Philippe Laurent (herpetólogo, 1917-2005)., Asociación Herpetológica Argentina (AHA)

Los casos de puestas comunales y de puestas coloniales han sido mencionados para varios grupos de Squamata (Graves y Duvall, 1995). Se define como puesta comunal a la deposición no incidental de huevos de dos o más hembras de una especie que utilizan una única cavidad (Espinoza y Lobo, 1996). En una puesta colonial, en cambio, los nidos son construidos de manera adyacente y los huevos generalmente no son depositados en la misma cavidad. Esta última estrategia de oviposición es más frecuente en aves que en lagartijas (Rand, 1967; Wiewandt, 1982; Mora, 1989; Burger, 1993; Espinoza y Lobo, 1996; Krysko et al., 2003). Entre los Teiidae sólo se conoce el caso de puesta comunal de Kentropyx calcarata (Magnusson y Lima, 1984)., Asociación Herpetológica Argentina (AHA)

Se citan varias especies de Squamata para algunos departamentos bolivianos, e inclusive algunas no citadas para Bolivia., Asociación Herpetológica Argentina