Search

Quantitative reconstructions of past land cover are necessary to determine the processes involved in climate-human-land-cover interactions. We present the first temporally continuous and most spatially extensive pollen-based land-cover reconstruction for Europe over the Holocene (last 11g€¯700g€¯calg€¯yrg€¯BP). We describe how vegetation cover has been quantified from pollen records at a 11 spatial scale using the "Regional Estimates of VEgetation Abundance from Large Sites"(REVEALS) model. REVEALS calculates estimates of past regional vegetation cover in proportions or percentages. REVEALS has been applied to 1128 pollen records across Europe and part of the eastern Mediterranean-Black Sea-Caspian corridor (30-75° N, 25° W-50° E) to reconstruct the percentage cover of 31 plant taxa assigned to 12 plant functional types (PFTs) and 3 land-cover types (LCTs). A new synthesis of relative pollen productivities (RPPs) for European plant taxa was performed for this reconstruction. It includes multiple RPP values (≥2 values) for 39 taxa and single values for 15 taxa (total of 54 taxa). To illustrate this, we present distribution maps for five taxa (Calluna vulgaris, Cerealia type (t)., Picea abies, deciduous Quercus t. and evergreen Quercus t.) and three land-cover types (open land, OL; evergreen trees, ETs; and summer-green trees, STs) for eight selected time windows. The reliability of the REVEALS reconstructions and issues related to the interpretation of the results in terms of landscape openness and human-induced vegetation change are discussed. This is followed by a review of the current use of this reconstruction and its future potential utility and development. REVEALS data quality are primarily determined by pollen count data (pollen count and sample, pollen identification, and chronology) and site type and number (lake or bog, large or small, one site vs. multiple sites) used for REVEALS analysis (for each grid cell). A large number of sites with high-quality pollen count data will produce more reliable land-cover estimates with lower standard errors compared to a low number of sites with lower-quality pollen count data. The REVEALS data presented here can be downloaded from https://doi.org/10.1594/PANGAEA.937075 (Fyfe et al., 2022). © 2022 Esther Githumbi et al.

Wildfire is the most common disturbance type in boreal forests and can trigger significant changes in forest composition. Waterlogging in peatlands determines the degree of tree cover and the depth of the burning horizon associated with wildfires. However, interactions between peatland moisture, vegetation composition and flammability, and fire regime in forested peatland in Eurasia remain largely unexplored, despite their huge extent in boreal regions. To address this knowledge gap, we reconstructed the Holocene fire regime, vegetation composition, and peatland hydrology at two sites in Western Siberia near Tomsk Oblast, Russia. The palaeoecological records originate from forested peatland areas in predominantly light taiga (Pinus-Betula) with the increase in dark taiga communities (Pinus sibirica, Picea obovata, Abies sibirica) towards the east. We found that the past water level fluctuated between 8 and 30 cm below the peat surface. Wet peatland conditions promoted broadleaf trees (Betula), whereas dry peatland conditions favoured conifers and a greater forest density (dark-to-light-taiga ratio). The frequency and severity of fire increased with a declining water table that enhanced fuel dryness and flammability and at an intermediate forest density. We found that the probability of intensification in fire severity increased when the water level declined below 20 cm suggesting a tipping point in peatland hydrology at which wildfire regime intensifies. On a Holocene scale, we found two scenarios of moisture-vegetation-fire interactions. In the first, severe fires were recorded between 7.5 and 4.5 ka BP with lower water levels and an increased proportion of dark taiga and fire avoiders (Pinus sibirica at Rybanya and Abies sibirica at Ulukh Chayakh) mixed into the dominantly light taiga and fire-resister community of Pinus sylvestris. The second occurred over the last 1.5 ka and was associated with fluctuating water tables, a declining abundance of fire avoiders, and an expansion of fire invaders (Betula). These findings suggest that frequent high-severity fires can lead to compositional and structural changes in forests when trees fail to reach reproductive maturity between fire events or where extensive forest gaps limit seed dispersal. This study also shows prolonged periods of synchronous fire activity across the sites, particularly during the early to mid-Holocene, suggesting a regional imprint of centennial to millennial-scale Holocene climate variability on wildfire activity. Increasing human presence in the region of the Ulukh-Chayakh Mire near Teguldet over the last four centuries drastically enhanced ignitions compared to natural background levels. Frequent warm and dry spells predicted for the future in Siberia by climate change scenarios will enhance peatland drying and may convey a competitive advantage to conifer taxa. However, dry conditions, particularly a water table decline below the threshold of 20 cm, will probably exacerbate the frequency and severity of wildfire, disrupt conifers’ successional pathway and accelerate shifts towards more fire-adapted broadleaf tree cover. Furthermore, climate-disturbance-fire feedbacks will accelerate changes in the carbon balance of forested boreal peatlands and affect their overall future resilience to climate change.

Disturbances by fire are among the most important processes that shape forest dynamics and diversity. However, the long-term variability of fire disturbance regimes in many European forests and specifically in the mountains of the Balkan Peninsula is not well understood. Here, we present the first Holocene record of fire regimes based on macrocharcoal morphologies in combination with pollen-based reconstruction of forest dynamics and fire-related strategies of prevailing mountain forests in the Rila Mountains, Bulgaria. While biomass burning followed the main trends in climate, the frequency and intensity of fire were strongly linked to fire-related coping strategies of dominant tree taxa (resisters, avoiders or invaders). Frequent fires of low intensity between 12,000 and 9000 cal year BP were concurrent with the dominance of invaders (Betula, herbs, ferns). Intermittent occurrence of low- and high-intensity surface and crown fires with longer return intervals between 9000 and 4000 cal year BP was associated with codominance of resister (Pinus sylvestris, Pinus peuce, Pinus mugo) and avoider (Abies alba and Picea abies) forest types, whereas a lengthening of the fire return interval over the past 4000 years was linked to increased abundance of P. abies. As a rising number of fire episodes may drive land cover towards more fire-adapted plant communities and towards less intense fire events, we expect increased dominance of invaders (resprouters that rapidly reach maturity stage) as well as resisters (properties protecting from fire damage) under future warmer and drier climate. This study also shows the potential of combining charcoal morphologies with pollen records to track variability in fire intensity and plant functional attributes over long timescales that are also relevant to forest management stakeholders.

Boreal forests are among the ecosystems most significantly impacted by wildfires as a consequence of climate warming. A large proportion of the global boreal forest area is located in Siberia, however, its vast extent and restricted access limit datasets recording changes in wildfire activity, especially from a longer-term perspective. Such long-term records of wildfire activity are vital to understanding how fire regimes vary with changes in climate, vegetation composition and human-vegetation interaction, as well as the impacts of wildfires on boreal forests.Here, we explore how patterns in fire regime (biomass burned, fire frequency, fire type) have changed over the Holocene. We focus on the relationship between fire regime, forest density and the fire-related traits of the main tree species, and peatland hydrology. We used charcoal-morphologies based reconstructions of fire regimes, along with pollen-based assessments of vegetation composition and testate amoebae-based hydro-climate reconstructions in Pinus-Betula dominated peatlands from central-western Siberia, Tomsk Oblast, Russia.The occurrence of more severe fires (i.e., higher biomass burning per fire episode and abundant woody morphotypes) were recorded between 7500 and 5000 cal yr BP. Higher temperatures during that time, likely enhanced peatland dryness and fuel flammability creating conditions conducive to peat and forest fires. Drier peatland conditions also affected forest composition and density by favouring the expansion of a mix of light taiga and fire resisters (e.g., Pinus sylvestris, P. sibirica, Larix) with denser taiga and fire avoiders (Picea obovata and Abies sibirica) on the peatland. A shift to the lowest biomass burning and fire types affecting mostly litter and understorey vegetation, was registered between 4000 and 1500 cal yr BP. Temporally, it coincides with an increase in peatland surface moisture and a change in forest composition characterised by a decline in fire resisters, while fire avoiders remained abundant. An almost synchronous intensification in fires frequency and severity from ca. 2000 cal yr BP to the present at all sites, was concurrent with the rise to dominance of fire-invader species (Betula), as well as a more abundant biomass in the understory layer (shrubs, herbs, ferns, moss), while fire resisters and avoiders declined substantially. We found that Picea obovata to be highly vulnerable tree taxa to frequent, severe fires.This long-term perspective demonstratesthat peatland hydrology is connected to, and feedbacks on peatland and forest composition and fuel dryness and ultimately fire regime. It also shows that more frequent fires of higher severity can lead to compositional or structural changes of forests, if trees cannot reach reproductive ages prior to the next burning events. Future predicted increases in temperatures are likely to enhance peatland drying, with cascading effects on forest and peat plant composition, subsequently exacerbating wildfire activity. This study thus contributes to an understanding of disturbance regimes in boreal forests and considers their potential to adapt to new climate conditions and fire regimes.

The collection of modern spatially extensive pollen data are important for the interpretation of fossil pollen diagrams. Such datasets are readily available for percentage data but lacking for pollen accumulation rates (PAR). Filling this gap has been the motivation of the pollen monitoring network, whose contributors monitored pollen deposition in modified Tauber-traps for several years or decades across European latitudes. Here we present this monitoring dataset consisting of 351 trap locations with a total of 2742 annual samples covering the period from 1981 to 2017. This dataset shows that climate parameters correlating with latitude determine pollen productivity. A signal of regional forest cover can be detected in the data, while local tree cover seems more important. Pollen traps situated beyond 200 km of the distribution of the parent tree are still collecting occasional pollen grains of the tree in question. PAR’s of up to 30 grains cm−2yr−1 in fossil diagram should therefore be interpreted as long distance transport. Comparisons to fossil data from the same areas show comparable values. Comparisons often demonstrate that similar high values for temperate taxa in fossils sites are found further south or downhill. While modern situations comparable to high PAR values of some taxa (e.g. Corylus) may be hard to find, CO2 fertilization and land use may case high modern PAR’s that are not documented in the fossil record. The modern data is now publically available in the Neotoma Paleoecology Database and hopefully serves improving interpretations of fossil PAR data.

Information on the Holocene vegetation history of the Central Rila Mountains for the last 10,000 years was obtained by means of pollen and fossil stomata analyses of a 500 cm core retrieved from peat bog Vodniza (2113 m a.s.l.), and supported by radiocarbon chronology. The early Holocene afforestation started with an initial Betula phase with stands of Juniperus and Pinus which occupied barren soils (10,000–8300 cal. yrs. BP) while deciduous oak forests with abundant Tilia , Ulmus , Acer , and later on Corylus , spread at lower elevations. During the Holocene climatic optimum (8300–6000 cal. yrs. BP) the conifers Pinus ( Pinus sylvestris , Pinus mugo , Pinus peuce ) and Abies expanded at the expense of the birch and oak forests. The tree line was running above 2100 m comparable to the present-day situation and started to descend after ca. 3200 cal. yrs. BP mostly due to human impact. The last tree which penetrated into the coniferous belt after 3400 cal. yrs. BP and reached altitudes of 2000–2100 m, confirmed by the find of fossil stomata, was Picea abies . Its maximal distribution was achieved between 2250 and 1100 cal. yrs. BP, alongside with some enlargement of beech communities. The first indications of stock-breeding and other human activities in the Central Rila Mountains were recorded since 3400–3200 cal. yrs. BP (Late Bronze Age). The vegetation reconstruction during the Holocene followed a pattern coherent with the information from the Northern Pirin and partly from the Western Rhodopes Mountains. The palaeoecological evidence from the Rila Mountains could be compared in broad lines with the high quality data obtained from the Romanian Carpathians by the application of multi-proxy research approach.

The peat bog Vapsko-2 (42° 04ʹ 26.85″ N, 23° 31ʹ 27.81″ E; 2120 m above sea level [a.s.l.]) is located in the lower range of the subalpine belt, at the present timberline, in the southern part of t...

The Eurasian (née European) Modern Pollen Database (EMPD) was established in 2013 to provide a public database of high-quality modern pollen surface samples to help support studies of past climate, land cover, and land use using fossil pollen. The EMPD is part of, and complementary to, the European Pollen Database (EPD) which contains data on fossil pollen found in Late Quaternary sedimentary archives throughout the Eurasian region. The EPD is in turn part of the rapidly growing Neotoma database, which is now the primary home for global palaeoecological data. This paper describes version 2 of the EMPD in which the number of samples held in the database has been increased by 60% from 4826 to 8134. Much of the improvement in data coverage has come from northern Asia, and the database has consequently been renamed the Eurasian Modern Pollen Database to reflect this geographical enlargement. The EMPD can be viewed online using a dedicated map-based viewer at https://empd2.github.io and downloaded in a variety of file formats at https: //doi.pangaea.de/10.1594/PANGAEA.909130 (Chevalier et al., 2019)., Swiss National Science Foundation (SNSF) 200021_169598, University of Lausanne

The peat bog Vapsko-1 (42° 04ʹ 34.04″ N, 23° 31ʹ 25.84″ E; 2143 m above sea level) is situated in a depression in the lower part of the subalpine belt, just above the present tree-line, in the sout...

Wildfire occurrence is influenced by climate, vegetation and human activities. A key challenge for understanding fire-climate-vegetation interactions is to quantify the effect vegetation has in mediating fire regime. Here, we explore the relative importance of Holocene land cover and dominant functional forest type, and climate dynamics on biomass burned in temperate and boreo-nemoral regions of Central and Eastern Europe over the past 12 ka BP years. We used an extensive data set of Holocene pollen and sedimentary charcoal records, in combination with climate simulations and novel statistical modelling. Biomass burned was highest during the early Holocene and lowest during the mid Holocene in all three ecoregions, but diverged more markedly over the past 3–4 ka BP. Although the climate was an important driver of fire hazard during the warm and dry early Holocene, tree cover was consistently the strongest predictor of past biomass burning. In temperate forests, biomass burned was high at ~ 45 % tree cover and decreased strongly towards 60 % tree cover. In needleleaf dominated forests, biomass burned was highest at ~ 60–65 % tree cover and abruptly declined at > 65 % tree cover. Biomass burned also increased when arable lands and grasslands reached ~ 15–20 %, although this relationship was highly dynamic depending on land use intensity throughout ignition and fuel type and availability. Our observations cover the full range of Holocene climate variability and land cover changes and illustrates that percentages of land cover is a key predictor of the probability of fire occurrence over timescales of centuries to millennia. We suggest that long-term fire risk may be effectively reduced through land cover management, given that land cover has controlled fire regimes under the dynamic climates of the Holocene.

Pollen and plant macrofossil analyses were performed on a sequence 105 cm deep obtained from a peat bog (1750 m) that is located above the present timber-line in the Osogovo Mountain, Southwestern ...

The vegetation history of the Central Rila Mountains for the last 14,000 years was reconstructed by means of pollen analysis and radiocarbon chronology of a core retrieved from Lake Manastirsko-2 (2326 m). The Lateglacial landscape was dominated by open herb vegetation composed of Artemisia, Chenopodiaceae, Poaceae, and other cold-resistant herbs (14,000–11,700 cal. yr BP). Stands of Pinus, Betula, and Juniperus–Ephedra shrubland partly enlarged during the Lateglacial interstadial. Pioneer forests of Betula with Pinus and Juniperus occupied barren soils in the early Holocene (11,700–7900 cal. yr BP), while mixed oak forests with Tilia, Ulmus, Acer, and later on Corylus spread at lower elevations. A coniferous forest belt with Pinus sylvestris, Pinus peuce, and Abies developed after 7900 cal. yr BP in the conditions of milder winters, cooler summers, and increase in precipitation. The late Holocene dynamic vegetation changes were associated with the invasion of Picea abies after 3400 cal. yr BP, while Fagus communities slightly enlarged in the river valleys. Indications of human activities are visible in the pollen diagram since the ‘Late Bronze Age’ (3400–3200 cal. yr BP). The vegetation development in the study area followed a similar pattern when compared with palynological and macrofossil records from other parts of the Rila and the adjacent Northern Pirin Mountains. On a larger geographical scale, the postglacial vegetation history of the Rila Mountains displays common features with sites in the Romanian Carpathians, whereas the differences observed are result of the location of tree refugia, competing abilities, climate changes, and human activities.

The peat bog Kumata-1 (42° 35ʹ 25.54ʹʹ N, 23° 15ʹ 06.85ʹʹ E; 1770 m above sea level) is situated in the upper part of the coniferous belt on the northern slopes of Vitosha Mountain (2290 m) – a Nat...

International audience; Aim: To evaluate the biomization technique for reconstructing past vegetation in the Eastern Mediterranean-Black Sea-Caspian-Corridor using an extensive modern pollen data set and comparing reconstructions to potential vegetation and observed land cover data. Location: The region between 28-48 degrees N and 22-62 degrees E. Methods: We apply the biomization technique to 1,387 modern pollen samples, representing 1,107 entities, to reconstruct the distribution of 13 broad vegetation categories (biomes). We assess the results using estimates of potential natural vegetation from the European Vegetation Map and the Physico-Geographic Atlas of the World. We test whether anthropogenic disturbance affects reconstruction quality using land use information from the Global Land Cover data set. Results: The biomization scheme successfully predicts the broadscale patterns of vegetation across the region, including changes with elevation. The technique discriminates deserts from shrublands, the prevalence of woodlands in moister lowland sites, and the presence of temperate and mixed forests at higher elevations. Quantitative assessment of the reconstructions is less satisfactory: the biome is predicted correctly at 44% of the sites in Europe and 33% of the sites overall. The low success rate is not a reflection of anthropogenic impacts: only 33% of the samples are correctly assigned after the removal of sites in anthropogenically altered environments. Open vegetation is less successfully predicted (33%) than forest types (73%), reflecting the under-representation of herbaceous taxa in pollen assemblages and the impact of long-distance pollen transport into open environments. Samples from small basins (

A palynological study was performed on a 2 m thick sediment sequence recovered from Mire Kupena (1356 m), a former lake in the Western Rhodopes Mountains (south Bulgaria) and supported by radiocarbon dating. The record extends back to ca. 30,000 cal. yrs. BP (Middle Pleniglacial) when the study area was occupied by wooded steppe composed of Pinus sp., Pinus peuce , some Betula , Juniperus , and cold-tolerant herb vegetation dominated by Artemisia and Chenopodiaceae. In addition, the almost continuous presence of deciduous Quercus , Tilia , Corylus , Abies , Picea , Fagus , Alnus , and Carpinus betulus pollen suggests interstadial conditions and this area could be recognized as a montane refugial place. A reconstruction of the interval between ca. 24,000 and 15,000 cal. yrs. BP was not possible due to an extremely low sediment accumulation rates or, more likely, a hiatus. The lateglacial landscape was dominated by mountain-herb steppe vegetation with isolated stands of Pinus , Betula and shrubland of Juniperus . The afforestation in the Early Holocene started with broad-leaved forests composed of Quercus with C. betulus , Carpinus orientalis / Ostrya , Ulmus , Tilia and Corylus and minor amounts of Pinus , Betula and Abies . In the Late Holocene ( Fagus began to gain importance chiefly at the expense of the mixed oak forests, while after ca. 2000 cal. yrs. BP forests of Pinus sylvestris with some P . peuce quickly expanded around the mire. A comparison with other long palynological records from the mountains and lowlands of Bulgaria and in northern Greece reveals not only common trends in the vegetation development that are a reflection of the climate changes, but also site-specific features related to the location and topography of each site.

The environmental changes (vegetation history, human impact and land use, influence of the Black Sea) in the area of Lake Durankulak, northeastern Bulgaria, were reconstructed and synthesized for the last ca. 8000 years. The palaeoecological information derived from various proxies (pollen, plant macrofossils, molluscs, sediments) was compared on a regional scale with the evidence from the nearby coastal lakes ShablaeEzeretz and Bolata. The Early Holocene xerothermic steppe vegetation, dominated by Chenopodiaceae, Artemisia and Poaceae species, and accompanied by stands of trees in moister habitats, was transformed after 6000 cal. BP into a forest-steppe, comprising oak woods with Carpinus betulus, Ulmus, Tilia, Acer. This vegetation pattern has been periodically modified, depleted and replaced by arable land or xerothermic herbaceous communities enriched with anthropophytes and ruderals, particularly after the intensification of human activities since 3300 cal. BP. The archaeobotanical evidence from the region has provided valuable information about the occupation phases and subsistence strategy of the local people since the Late Neolithic (5300 cal. BC/7250 cal. BP). Periods with cultivation of cereals (Triticum, Hordeum) and/or stock-breeding activity were interrupted by abandonment of the settlements and the arable land due to unfavourable environmental changes. The periodical connection/isolation of Lake Durankulak with the Black Sea and the periods of marine influence were recorded by changes in the composition of the fossil molluscan fauna and the lithology of the sediments, and chronologically confirmed by radiocarbon dates. The development of the coastal lakes throughout the largest part of the Holocene has been also considerably influenced by the fluctuations of the Black Sea level.

A pollen analysis conducted on a 600 cm core from Lake Ribno (2184 m) in the Northwestern Rila Mountains, supplemented by 13 radiocarbon dates, has revealed the basic stages in its postglacial vegetation dynamics. The lateglacial vegetation, composed of Artemisia, Chenopodiaceae and Poaceae, with stands of Pinus and Juniperus-Ephedra shrubland, dominated in the stadials and partly retreated during the Bølling/Allerød interstadial (14700–12900 cal. yrs. BP). The afforestation in the early Holocene (11600–7800 cal. yrs. BP) started with pioneer Betula forests, with groups of Pinus and Juniperus at mid-high altitudes, and Quercus forests with Tilia, Ulmus, Fraxinus, Corylus below the birch zone. A coniferous belt composed of Pinus sylvestris, P. peuce and Abies was shaped under the conditions of a more humid climate (7800–5800 cal. yrs. BP). The last trees that invaded the study area were Fagus after 4300 cal. yrs. BP and Picea abies after 3400 cal. yrs. BP. Evidence for destructive changes in the vegetation and indications of agricultural and stock-breeding activities (pollen of Triticum, Secale, Plantago lanceolata, Rumex, Juglans) was continuously recorded since the Late Bronze Age (3400–3200 cal. yrs. BP). The postglacial vegetation history in the Northwestern Rila Mountains demonstrated close similarities with that of the Northern Pirin Mountains.

Palaeoecological reconstructions from the region of southwestern Bulgaria were used for inferring the human impact on the vegetation and landscape during the last 8 millennia. They are based on data from pollen analyses of lakes and peat-bogs, plant macrofossils, archaeobotanical finds and radiocarbon dating. During the early Holocene, after 7900 cal. b.p. (5950 cal. b.c.) the climate changed to cooler summers, milder winters and higher precipitation resulting in the formation of a coniferous belt dominated by Pinus sp. and Abies alba. These favorable environmental pre-conditions had a positive influence on the Neolithisation of the Balkans after the 8200 cal. b.p. (6250 cal. b.c.) cold event, which caused drought in the Eastern Mediterranean. Direct evidence from wood charcoal records from the Neolithic settlement layers in the study area shows a slight modification of the surrounding woodlands and an increase of the light-demanding components, probably expressed through larger forest border zones and thinning out of the wood stands. The increase in the number of settlements in the valleys of southwestern Bulgaria intensified the human activity visible in the palaeobotanical record from 6950 cal. b.p. (5000 cal. b.c.) onwards. Between ca. 5700–5100 cal. b.p. (3800–3200 cal. b.c.) signs of anthropogenic influence on the vegetation are virtually absent. The intensity of human impact increased notably after 3200 cal. b.p. (1400–1250 cal. b.c., approx. Late Bronze Age), documented by a rise of pollen anthropogenic indicators. The final transformations in the natural forest cover after 2750 cal. b.p. (800 cal. b.c. onset of the Iron Age) marked the reduction of the coniferous forests dominated by Abies alba and Pinus sp. and the expansion of Fagus sylvatica and Picea abies. These vegetation changes are contemporaneous with increase of the palaeofire activities and the next peak of anthropogenic indicators. The changes in the landscape during the Roman period and the medieval period reflect regional environmental features and were forced by the diversification of anthropogenic activity.

Pollen analysis and AMS radiocarbon dating were conducted on the lateglacial sediments (175 cm in thickness) of a core obtained from the glacial Lake Ribno (2184 m) in the northwestern Rila Mountains. The vegetation dynamics at high-mid altitudes based on important changes in palynostratigraphy as a response to the lateglacial climate fluctuations for the time interval 16000-11600 cal.yrs BP is bound to an interstadial/stadial cycle, analogous with the Bolling/Allerod-Younger Dryas. The established local chronology demonstrates also good correlation with the isotopic signal from Greenland ice-cores. The reconstruction of the lateglacial vegetation shows common features with other sites from the Bulgarian mountains (Rila, Pirin, Western Rhodopes) and the glacial lakes in the southern Romanian Carpathians (Retezat Mountains).

Mire Gyola (41° 21′ 17.12″ N, 23° 07′ 31.31″ E; 714 m above sea level (a.s.l.)) is situated on the northern slopes of the Belasitsa Mountain (2029 m) in south-western Bulgaria. The territory of the...

A 7.5 m sediment core from Lake Bolata, a small former coastal liman lake in northeastern Bulgaria, was analyzed for pollen and plant macrofossil content in attempt to trace the changes in the vegetation, human impact and the influence of the Black Sea during the last ca. 6,000 years. Lake Bolata started its existence when the rising Black Sea level reached the elevation of the bottom of the depression. By that time nearly all tree species were already present in this area with the exception of Carpinus orientalis. The comparison of the arrival time of oriental hornbeam at different sites along the Bulgarian Black Sea confirmed the reliability of the local radiocarbon chronology. For the period under study the vegetation of the region around the site can be described as a forest-steppe due to NAP values higher than 40% of the pollen sum. Forests on the slopes of the river canyon consisted of Quercus spp., Acer, Carpinus orientalis, Fraxinus ornus, Fagus and possibly Tilia. Riverine forests formed stands composed of Salix, Alnus, Ulmus, Fraxinus, Carpinus betulus and Vitis as a liana. The vegetation on the Cape Kaliakra plateau, bordering the canyon, was dominated by diverse herbs and most of these could be considered xerophytes. The oldest occupation period belongs to the Eneolithic, documented by the occurrences of Triticum-type pollen at 5570–5170 cal b.p. (3620–3220 b.c.). The next period of higher continuous Cerealia-type values corresponds to 3450–1830 cal b.p. (1500 b.c.–a.d. 120). The macrofossil record chiefly provides evidence about plant communities of aquatics and helophytes.

Pollen analysis of sediments from the glacial Lake Trilistnika (2216 m) in the Northwestern Rila Mountains (Bulgaria), supplemented by 13 radiocarbon dates, allowed the reconstruction of the palaeoenvironment and vegetation history in postglacial time. The exact time of the cirque glacier retreat is still under discussion but the lake was free of ice before 15,000 cal. BP, when sedimentation of gray silt began. The lateglacial vegetation, composed of Artemisia , Chenopodiaceae and Poaceae, with isolated stands of Pinus and Juniperus – Ephedra shrubland, dominated during the stadials and partly retreated during the Bolling/Allerod interstadial complex. The afforestation in the early Holocene (11,500–7800 cal. BP) started with the distribution of pioneer Betula forests with groups of Pinus ( P. mugo , P. sylvestris and P. peuce ) at mid-high altitudes, and Quercus forests with Tilia , Ulmus , Fraxinus , Corylus below the birch zone. The change to more humid and cooler climate ca. 7800–7500 cal. BP favored the vertical migration of Abies , P. sylvestris and P. peuce . The establishment of Fagus sylvatica took place after 5200 cal. BP, when pure or mixed beech communities were formed. The last tree which invaded the coniferous belt between 4300 and 3400 cal. BP was Picea abies . The first expansion maximum of spruce was recorded after ca. 2700 cal. BP. The vegetation development in historical times was also influenced by human interference, indicated by the continuous presence of pollen anthropogenic indicators such as Triticum , Secale , Hordeum , Plantago lanceolata , Rumex , Scleranthus , Juniperus .

Pollen analysis, supplemented by 5 14C dates, was carried out on the lateglacial section (160 cm) of a core retrieved from the glacial Lake Trilistnika (2216 m) in the northwestern Rila Mountains (Bulgaria). The reconstruction of the lateglacial vegetation history is linked for the first time to a chronological framework for the time window 13000–10000 b.p. The delimitation of an interstadial/stadial cycle, analogous with the Bolling/Allerod-Younger Dryas from Western Europe, is based on important changes in the pollen stratigraphy. Mountain-steppe vegetation composed of Artemisia, Chenopodiaceae, Poaceae and other cold-resistant herbs, with isolated stands of Pinus and Juniperus-Ephedra shrubland, dominated the landscape after the ice-retreat. Interstadial conditions were established before 12815±130 b.p. The spread of tree vegetation at lower elevation as a response to the climate improvement was confined to the time interval 12110±95–11140±75 b.p. The Younger Dryas stadial is characterised by a re-advance of the mountain-steppe vegetation. The results are compared with other sites of lateglacial age from the high mountains (Rila, northern Pirin, western Rhodopes) in southern Bulgaria.

Pollen analysis, plant macrofossil determination and radiocarbon dating were performed on profiles of Holocene age obtained from two subalpine lakes (2320-2340 in a.s.l.) in the Central Rila Mountains, Bulgaria. The palaeovegetational reconstruction dates back to the mid-Atlantic (6500-6700 cal. BP). The plant macrofossil record revealed that groups of Pinus mugo and Juniperus sibirica were distributed in the subalpine zone. The treeline was shaped by Pinus sylvestris and Pinus peuce and was higher (up to 2200-2300 in a.s.l.) compared with the present-day. A change to cooler summers and warmer winters after 6700 cal. BP, characterized by a rise in humidity and precipitation, stimulated the formation of a coniferous vegetation belt dominated by Pinus sylvestris, Pinus peuce and Abies alba. The pollen assemblages also indicated a dominance of deciduous trees (Quercetum mixtum-Corylus phase) at low altitudes until c. 6000 cal. BP. The sub-Boreal was characterized by a wider distribution of Abies alba in the lower part of the coniferous belt and a gradual enlargement of the areas occupied by Fagus sylvatica and Picea abies after 3800-3500 cal. BP. The macrofossil finds of Picea abies testify to its expansion after 2800 cal. BP when average temperatures dropped and precipitation increased. Traces of human disturbance in the subalpine area are continuously registered in the pollen and plant macrofossil records since 3770 cal. BP, indicating livestock-grazing, burning of dwarf-pine (Pinus mugo) for new pasture land and lowering of the treeline. ispartof: The Holocene: an interdisciplinary journal focusing on recent environmental change vol:15 issue:5 pages:663-671 status: published

Site details Mire Kupena is a former lake (41◦ 59′ 07.5′′ N, 24◦ 19′ 05.1′′ E; 1356 m above sea level (a.s.l.)) located on the territory of the Biosphere Reserve Kupena, Western Rhodopes Mountains, Bulgaria. The reserve is located between 600 and 1400 m a.s.l. on a north-facing slope and its vegetation is rather diverse. In the lower parts, the most common plant communities are those of Quercus dalechampii Ten. mixed in some places with Carpinus betulus L., Fagus sylvatica L., Pinus nigra Arn. and Pinus sylvestris L. At higher altitudes, plant communities of F. sylvatica dominate, followed by those of Pinus sylvestris with some Abies alba Mill. and Picea abies (L.) Karst. The mire is formed in a depression on an area of 6 ha, surrounded by an almost pure forest of Pinus sylvestris with an admixture of Picea abies, A. alba, F. sylvatica, Betula pendula Roth., Sorbus aucuparia L., Q. dalechampii and an undergrowth of Juniperus communis L., Vaccinium myrtillus L. and V. vitis-idaea L. The mire vegetation is represented by sparse moss cover (Sphagnum spp.) and includes various species of Carex, Juncus, Potentilla, Ranunculus, Galium. The basin is fed by rainfall and water from melting snow. During the last two decades most of the mire surface on the hummocks has been overgrown by pine trees (Huttunen et al., 1992; Bozilova et al., 2011). The climate in the area is montane with a mean annual temperature of 5–10 ◦S and a mean annual precipitation of 600–800 mm with a maximum in May–June. The basic soil types are brown forest (60%), humic-carbonate and cinnamomic-forest.

We present the first area-average time series reconstructions of warmest month, coldest month and mean annual surface air temperatures across Europe during the last 12,000 years. These series are based on quantitative pollen climate reconstructions from over 500 pollen sites assimilated using an innovative four-dimensional gridding procedure. This approach combines three-dimensional spatial gridding with a fourth dimension represented by time, allowing data from irregular time series to be ‘focussed’ onto a regular time step. We provide six regional reconstructed temperature time series as well as summary time series for the whole of Europe. The results suggest major spatial and seasonal differences in Holocene temperature trends within a remarkably balanced regional and annual energy budget. The traditional mid-Holocene thermal maximum is observed only over Northern Europe and principally during the summer. This warming was balanced by a mid-Holocene cooling over Southern Europe, whilst Central Europe occupied an intermediary position. Changes in annual mean temperatures for Europe as a whole suggest an almost linear increase in thermal budget up to 7800 BP, followed by stable conditions for the remainder of the Holocene. This early Holocene warming and later equilibrium has been mainly modulated by increasing winter temperatures in the west, which have continued to rise at a progressively decreasing rate up to the present day.

Pollen analysis and radiocarbon dating were performed on a sequence 150 cm deep from a peat bog (Mozgovitsa, 1800 m above sea-level) in the Northwestern Pirin Mountains, Bulgaria. The palaeovegetational reconstruction focused on the main stages of forest development starting from approximately 8700 years BP. The vegetational response to the amelioration of the climate in the Early Holocene until 7500 cal BP resulted in the spread of birch forests (Betula pendula) that occupied large areas at high altitudes forming the upper tree-line together with pine (Pinus sylvestris, Pinus peuce, Pinus mugo), juniper (Juniperus), alder (Alnus), and willow (Salix). Below them were mixed deciduous oak forests with some Ulmus, Tilia, Corylus, Carpinus, and Acer. The continuous Early Holocene pollen record of mesophyllous demanding trees (Abies alba, Fagus sylvatica and Picea abies), though in low frequencies, suggests that groups of these trees survived in environmentally favourable habitats with sufficient moisture such as deep mountain valleys. The most important change in the vegetation cover started ca. 7500 cal BP when birch forests declined and coniferous vegetation dominated by Pinus sylvestris, Pinus peuce and Abies alba invaded the slopes. The spread of the conifers and the expansion of their areas presume an increase in humidity and precipitation, and the development of soils with humic horizons. The last trees to establish after ca. 4000 cal BP in the study area were Picea abies and Fagus sylvatica. Their expansion in the Northwestern Pirin Mountains started at the Subboreal/Subatlantic transition ca. 3000 cal BP when average temperatures dropped and precipitation increased. Deforestation in historic times resulted in the appearance of new pasture land and the lowering of the upper tree-line.

Pollen analysis was performed on a 343 cm long core recovered from Lake Ribno Banderishko (2190 m) on Northern Pirin Mountain (southwestern Bulgaria), supplemented by six '4C AMS dates. The lower part of the sequence (343-290 cm) was deposited during the Lateglacial when mountain-steppe vegetation with Artemnisia, Poaceae, Chenopodiaceae and other herbs, and with stands of Pinus and Juniperus, dominated at high altitudes. The identification of pollen of mesophilous trees suggests that local populations may have survived in refuges below 1000 m where environmental conditions were favourable for their growth. The reforestation dynamics in early-Holocene times (11200-8400 cal. BP) started with Betula forests at high altitudes and deciduous forests with Quercus, Ulmus, Tilia and Fraxinus excelsior at lower altitudes. The time of the maximal spread of Corylus at c. 8000 cal. BP coincided with the establishment of Abies alba in the forests, and the beginning of the formation of a coniferous belt, dominated by pines (Pinus sylvestris, P. peuce, P. nigra and P. heldreichii). The late immigrants Fagus sylvatica and Picea abies invaded the mountain slopes at c. 4400 cal. BP and 3300 cal. BP, respectively. Palynological traces of anthropogenic activity and landscape management began after c. 2200 cal. BP.

For most of the last glacial period, which ended about 10 ka BP 2 , the temperate forest species were restricted to small areas (termed refugia) with a milder climate, situated mostly along the Mediterranean borderlands and around the Black Sea. Species only started to expand from these glacial period refugia with the large-scale shifts in the global climate in the late-glacial (15–10 ka BP) and the beginning of the Holocene period (10 ka BP to present). Fossil pollen data from sites across Europe have been used to reconstruct the location of refugia of the deciduous oak species, and the spread from these refugia into their current ranges. Three areas of southern Europe have been identified as refugia for deciduous Quercus: southern Iberian peninsula, southern Italian peninsula and the southern Balkan peninsula. The spread of Quercus took place in two steps. First, in the late-glacial interstadial (13–11 ka BP) Quercus spread to the central European mountains from these refugia. Second, with the stabilisation of a climate favourable to deciduous trees species in the Holocene, oak spread into northern Europe, rapidly into the north–west, and more slowly into the centre and east, due to physical barriers. The earlier distribution changes are strongly correlated with the shifts in climate, whereas the later changes are most strongly controlled by competition between species, landscape topography and other edaphic factors. By approximately 6 ka BP, the deciduous oak had reached its maximum extension in Europe. Two types of refugia have been identified from the observed range expansion: primary, full glacial refugia; and secondary, temporary refugia, which supported populations of the oak during the short, climatically unfavourable late-glacial stadial.

ispartof: Grana: an International Journal of Palynology vol:50 issue:2 pages:162-164 status: published

Records of past climate variability and associated vegetation response exist in various regions throughout Central and Eastern Europe (CEE). To date, there has been no coherent synthesis of the existing palaeo-records. During an INTIMATE meeting (Cluj Napoca, Romania) focused on identifying CEE paleo-records, it was decided to address this gap by presenting the palaeo-community with a compilation of high-quality climatic and vegetation records for the past 60-8 kyrs. The compilation should also serve as a reference point for the use in the modelling community working towards the INTIMATE project goals, and in data-model inter-comparison studies. This paper is therefore a compilation of up to date, best available quantitative and semi-quantitative records of past climate and biotic response from CEE covering this period. It first presents the proxy and archive used. Speleothems and loess mainly provide the evidences available for the 60-20 ka interval, whereas pollen records provide the main source of information for the Lateglacial and Holocene. It then examines the temporal and spatial patterns of climate variability inferred from different proxies, the temporal and spatial magnitude of the vegetation responses inferred from pollen records and highlights differences and similarities between proxies and sub-regions and the possible mechanisms behind this variability. Finally, it identifies weakness in the proxies and archives and their geographical distribution. This exercise also provides an opportunity to reflect on the status of research in the area and to identify future critical areas and subjects of research. ispartof: Quaternary Science Reviews vol:106 pages:206-224 status: published

Modern pollen deposition is monitored for the Central Rila Mountains, Southwestern Bulgaria, by means of pollen traps, moss samples and lake surface sediments. Results are presented for three sampling sites from different vegetation belts along an altitudinal transect (1000–1900 m) for a seven-year period of observations (1993–1999). Both the annual influx and percentage values of the main arboreal and non-arboreal taxa are presented and discussed in relation to pollen production and dispersal distance. The greatest proportion of the pollen being deposited originates from Pinus diploxylon -type ( P. sylvestris and P. mugo ) irrespective of its abundance in the local vegetation. The influx values of the endemic Balkan species Pinus peuce ( Pinus haploxylon -type pollen) are several times lower than those of P. diploxylon -type. Abies alba and Picea abies have a representation in the deposited pollen assemblage which is comparable with the distribution and abundance of those species in the surrounding vegetation. Although the monitoring series is too short to confirm it, the cyclical flowering of Fagus sylvatica seems to be reflected in the trap results. The amount of pollen of low altitude species, which is deposited in the sub-alpine zone is very high. The data obtained from the different depositional media are compared. That the moss samples reflect pollen deposition from more than one year is evidenced by the absence of the distinct peak years seen in the trap results. In the light of these results the interpretation of previously analysed fossil pollen assemblages from the same study area may need to be revised.

A palynological investigation was conducted on two cores with Holocene sediments collected from the northeastern littoral part of the border Lake Doirani in northern Greece. The radiocarbon dates indicated that the analyzed sediments accumulated during the last 5000 yrs. The pollen-stratigraphic record revealed the environmental changes in the catchment area, starting from a natural undisturbed landscape to one modified by increasing anthropogenic influences. The tree vegetation dominated by Quercus woods in the lowlands and byPinus, Abies, and Fagus at higher altitudes, lasted for the period 2900 - 830 cal. B.C. Subsequently it was replaced by xerothermic herb and tree vegetation as a result of intensive human activity - and farming and stock-breeding. The accumulation of sediments with more sand and gravel in historical time was the result of increased erosion.

Palynological investigation and radiocarbon dating of a 6-m core from lake Durankulak, north-eastern Bulgaria, enables vegetation development and human occupation from ca. 5500–5300 cal. B.C. onwards to be traced. Steppe vegetation that included with groves of deciduous trees asQuercus, Ulmus, Carpinus belulus andCorylus changed to a forest-steppe after 4000 cal. B.C. The archaeopalynological record indicates three distinct phases of human activity as follows: (1) 5300–4200 cal. B.C. (late Neolithic and Eneolithic) during which farming, that included a substantial arable component, was pursued, (2) 3500–3000 cal. B.C. (transition to early Bronze Age) when stock rearing appears to have dominated, and (3) after 1300 cal. B.C. (late Bronze Age) when arable farming again assumed importance. The palynological data correlate well with the rich archaeological record for human settlement that is available for the region from late Neolithic times onwards.

Comparisons of climate model hindcasts with independent proxy data are essential for assessing model performance in non-analogue situations. However, standardized palaeoclimate data sets for assessing the spatial pattern of past climatic change across continents are lacking for some of the most dynamic episodes of Earth's recent past. Here we present a new chironomid-based palaeotemperature dataset designed to assess climate model hindcasts of regional summer temperature change in Europe during the late-glacial and early Holocene. Latitudinal and longitudinal patterns of inferred temperature change are in excellent agreement with simulations by the ECHAM-4 model, implying that atmospheric general circulation models like ECHAM-4 can successfully predict regionally diverging temperature trends in Europe, even when conditions differ significantly from present. However, ECHAM-4 infers larger amplitudes of change and higher temperatures during warm phases than our palaeotemperature estimates, suggesting that this and similar models may overestimate past and potentially also future summer temperature changes in Europe.

Davis, B.A.S. et al., Modern pollen samples provide an invaluable research tool for helping to interpret the quaternary fossil pollen record, allowing investigation of the relationship between pollen as the proxy and the environmental parameters such as vegetation, land-use, and climate that the pollen proxy represents. The European Modern Pollen Database (EMPD) is a new initiative within the European Pollen Database (EPD) to establish a publicly accessible repository of modern (surface sample) pollen data. This new database will complement the EPD, which at present holds only fossil sedimentary pollen data. The EMPD is freely available online to the scientific community and currently has information on almost 5,000 pollen samples from throughout the Euro-Siberian and Mediterranean regions, contributed by over 40 individuals and research groups. Here we describe how the EMPD was constructed, the various tables and their fields, problems and errors, quality controls, and continuing efforts to improve the available data.

Annual pollen-accumulation rates (PAR) of Fagus (beech) obtained within the framework of the Pollen Monitoring Programme (PMP) were analyzed in pollen traps along a N–S transect from the Baltic Sea to the Black Sea in different European vegetation units. The study regions are situated in the lowlands of northern Poland, the uplands of SE Poland, the Czech Krkonoše Mts, the Czech Šumava Mts, the Swiss Jura Mts, the Swiss Alps, the Bulgarian Rila Mts and the Bulgarian Strandzha Mts. Most time series are 10 or 11 years long, some are 5–16 years long. Inter-annual fluctuations in Fagus PAR were analyzed and compared with seed mast years. Years with high Fagus PAR and others with low Fagus PAR occurred most frequently in parallel within each region and often in two neighbouring regions. 2006 was exceptional as it had a very high Fagus sylvatica pollen deposition in all study regions and it was also a mast year. In Bulgaria, the trend in the 5 years of Fagus orientalis PAR in the Strandzha Mts differed from that of F. sylvatica PAR in the Rila Mts. Aiming at establishing the relationship between average Fagus PAR and tree cover, differences in Fagus PAR (averaged per pollen trap) were related in each region to the proportion of beech trees in the vegetation within 2 km of the pollen traps, the distance to the nearest pollinating Fagus tree, regional or local presence of beech forests, the degree of landscape openness, and the size of forest opening in which a trap is situated. Average Fagus PAR was found to track the regional abundance of beech trees in the vegetation, not the distance of the nearest Fagus tree. Regional occurrence of beech-dominated forests was reflected by a Fagus PAR of ca. 1,400 grains cm−2 year−1, local abundance very close to pollen traps by ca. 2,400 grains, small patches of forest with admixture of Fagus by ca. 170–220 grains, and scarcity or absence of Fagus by ca. 40 grains or less. peerReviewed

This paper compares pollen spectra derived from modified Tauber traps and moss samples from a selection of woodland types from Bulgaria, the Czech Republic, Georgia, Greece, Poland, Switzerland and Wales. The study examines the representation of individual taxa in the two sampling media and aims to ascertain the duration of pollen deposition captured by a moss. The latter aim was pursued through the calculation of dissimilarity indexes to assess how many years of pollen deposited in a pollen trap yield percentage values that are most similar to those obtained from the moss. The results are broadly scattered; the majority of moss samples being most similar to several years of pollen deposition in the adjacent trap. For a selection of samples, a comparison of the pollen accumulation rate in pollen traps with the pollen concentration in the moss per unit surface indicates that the entrapment and/or preservation of individual pollen types in the moss differ from that in the pollen trap. A comparison of the proportion of different taxa in the moss with the pollen spectrum of 2 years of pollen deposition in the trap also revealed large differences. There is a tendency for bisaccate grains such as Pinus and Picea to have a higher representation in moss than in traps but there is considerable regional variation. The results indicate that pollen proportions from moss samples often represent the pollen deposition of one area over several years. However, bisaccate pollen grains tend to be over-represented in moss samples compared to both pollen traps and, potentially, lake sediments. peerReviewed

Pollen stratigraphies are the most spatially extensive data available for the reconstruction of past land-cover change. Detailed knowledge of past land-cover is becoming increasingly important to evaluate the present trends in, and drivers of, vegetation composition. The European Pollen Database (EPD) was established in the late 1980s and developed in the early 1990s to provide a structure for archiving, exchanging, and analysing Quaternary pollen data from Europe. It provides a forum for scientists to meet and engage in collaborative investigations or data analysis. In May 2007 several EPD support groups were developed to assist in the task of maintaining and updating the database. The mapping and data accuracy work group (MADCAP) aims to produce an atlas of past plant distributions as detected by pollen analyses in Europe, in order to meet the growing need for this data from palaeoecologists and the wider scientific community. Due to data handling problems in the past, a significant number of EPD datasets have errors. The initial task of the work group, therefore, was a systematic review of pollen sequences, in order to identify and correct errors. The EPD currently (January 2009) archives 1,032 pollen sequences, of which 668 have age-depth models that allow chronological comparison. Many errors have been identified and corrected, or flagged for users, most notably errors in the pollen count data. The application of spatial analyses to pollen data is related to the number of data points that are available for analysis. We therefore take this opportunity to encourage the submission of pollen analytical results to the EPD or other relevant pollen databases. Only in this way will the scientific community be able to gain a better understanding of past vegetation dynamics. peerReviewed

The peat-bog Begbunar (42°09¢ N, 22° 33¢ E; 1750 m a.s.l.) is located in the central treeless zone of the Osogovo mountains, which are situated at the border between south-western Bulgaria and the north-eastern former Yugoslav Republic of Macedonia. This bog was formed near a freshwater spring on a north-western slope, which continues into a steep, deep ravine where isolated stands of beech (Fagus sylvatica L.) exist. The modern mountain vegetation consists of several vegetation belts: the oak-hornbeam belt (up to 1000 m), the compact beech belt (1000 – 1900 m) with fragments of conifers (Pinus nigra J. F. Arnold and Abies alba Mill.) and the subalpine belt, which is occupied by plant communities of Juniperus sibirica Burgsd., Vaccinium myrtillus L., Bruckenthalia spiculifolia (Salisb.) Rchb., Chamaecytisus absinthioides Janka (Kuzm.), Nardus stricta L., and scattered groups of Pinus sylvestris L., etc. The present-day vegetation composition has been strongly influenced by the long-lasting anthropogenic impact, including ore-mining industry and erosion caused by deforestation (Velcev & Tonkov, 1986; Tonkov, 2003). Geologically, the massif is composed mainly of Palaeozoic metamorphic and intrusive rocks, while geomorphological evidence suggests that small valley glaciers existed in the highest parts during Quaternary glaciations. The climate above 1000 m is typical montane with a mean annual precipitation of 700–900 mm. The basic soil types are cinnamomic-forest, brown forest and mountainous-meadow (Velcev et al., 1994).