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Climatic water availability is a major determinant of forest structure and composition, while drought events may severely impact forest dynamics. In recent decades, an increasing number of severe drought events has been reported in forests around the world. In the future, climate models project increasingly dry conditions in many temperate regions. Various tools have been applied to better understand the effects of drought on forests, such as dendrochronological analyses, climatic trend analyses and dynamic models. With these approaches, water availability is often summarized as a single scalar, termed a drought or aridity index. As droughts are complex phenomena, such indices are always associated with a loss of information. Many different such indices exist, and have been developed for various purposes. This review asks whether some of these indices are more suitable than others to quantify water availability in temperate forests. In a first step, the rationale and theoretical background of different drought indices are spelled out and compared among each other. Then, evaluations and intercomparisons of drought indices from the literature are reviewed. The implementation of drought indices in dynamic forest models is also discussed. Finally, two current research questions are identified: the role of dry air for physiological drought, and the suitability of various drought indices under climate change. It appears from this review that indices accounting for evaporative demand generally perform better than indices based on precipitation alone. When comparing sites with different edaphic conditions, indices accounting for soil moisture storage are more suitable. Nevertheless, results from intercomparisons show considerable divergence, and it is not possible to clearly favor one index. Furthermore, a differential response of tree species to different drought indices is often observed, although no clear pattern emerges from this comparison. More intercomparisons of indices, across climates and species, might provide valuable knowledge. Another key finding is that the properties of indices also depend on choices regarding, e.g., the calculation of evaporative demand, or the underlying water balance model. Reporting such methodological details could greatly increase the value of future evaluations of drought indices., iForest, 12 (1), ISSN:1971-7458

Rooting zone water storage capacity Sr is a crucial parameter for modeling hydrology, ecosystem gas exchange and vegetation dynamics. Despite its importance, this parameter is still poorly constrained and subject to high uncertainty. We tested the analytical, optimality-based model of effective rooting depth proposed by Guswa (2008, 2010) with regard to its applicability for parameterizing Sr in temperate forests. The model assumes that plants dimension their rooting systems to maximize net carbon gain. Results from this model were compared against values obtained by calibrating a local water balance model against latent heat flux and soil moisture observations from 15 eddy covariance sites. Then, the effect of optimality-based Sr estimates on the performance of local water balance predictions was assessed during model validation. The agreement between calibrated and optimality-based Sr varied greatly across climates and forest types. At a majority of cold and temperate sites, the Sr estimates were similar for both methods, and the water balance model performed equally well when parameterized with calibrated and with optimality-based Sr. At spruce-dominated sites, optimality-based Sr were much larger than calibrated values. However, this did not affect the performance of the water balance model. On the other hand, at the Mediterranean sites considered in this study, optimality-based Sr were consistently much smaller than calibrated values. The same was the case at pine-dominated sites on sandy soils. Accordingly, performance of the water balance model was much worse at these sites when optimality-based Sr were used. This rooting depth parameterization might be used in dynamic (eco)hydrological models under cold and temperate conditions, either to estimate Sr without calibration or as a model component. This could greatly increase the reliability of transient climate-impact assessment studies. On the other hand, the results from this study do not warrant the application of this model to Mediterranean climates or on very coarse soils. While the cause of these mismatches cannot be determined with certainty, it is possible that trees under these conditions follow rooting strategies that differ from the carbon budget optimization assumed by the model., Hydrology and Earth System Sciences, 22 (7), ISSN:1027-5606, ISSN:1607-7938

Projections of landscape dynamics are uncertain, partly due to uncertainties in model formulations. However, quantitative comparative analyses of forest landscape models are lacking. We conducted a systematic comparison of all forest landscape models currently applied in temperate European forests (LandClim, TreeMig, LANDIS-II, iLand). We examined the uncertainty of model projections under several future climate, disturbance, and dispersal scenarios, and quantified uncertainties by variance partitioning. While projections under past climate conditions were in good agreement with observations, uncertainty under future climate conditions was high, with between-model biomass differences of up to 200 t ha−1. Disturbances strongly influenced landscape dynamics and contributed substantially to uncertainty in model projections (~25–40% of observed variance). Overall, model differences were the main source of uncertainty, explaining at least 50% of observed variance. We advocate a more rigorous and systematic model evaluation and calibration, and a broader use of ensemble projections to quantify uncertainties in future landscape dynamics., Environmental Modelling & Software, 134, ISSN:1364-8152, ISSN:1873-6726

We present FORHYCS (FORests and HYdrology under Climate Change in Switzerland), a distributed ecohydrological model to assess the impact of climate change on water resources and forest dynamics. FORHYCS is based on the coupling of the hydrological model PREVAH and the forest landscape model TreeMig. In a coupled simulation, both original models are executed simultaneously and exchange information through shared variables. The simulated canopy structure is summarized by the leaf area index (LAI), which affects local water balance calculations. On the other hand, an annual drought index is obtained from daily simulated potential and actual transpiration. This drought index affects tree growth and mortality, as well as a species-specific tree height limitation. The effective rooting depth is simulated as a function of climate, soil, and simulated above-ground vegetation structure. Other interface variables include stomatal resistance and leaf phenology. Case study simulations with the model were performed in the Navizence catchment in the Swiss Central Alps, with a sharp elevational gradient and climatic conditions ranging from dry inner-alpine to high alpine. In a first experiment, the model was run for 500 years with different configurations. The results were compared against observations of vegetation properties from national forest inventories, remotely sensed LAI, and high-resolution canopy height maps from stereo aerial images. Two new metrics are proposed for a quantitative comparison of observed and simulated canopy structure. In a second experiment, the model was run for 130 years under climate change scenarios using both idealized temperature and precipitation change and meteorological forcing from downscaled GCM-RCM model chains. The first experiment showed that model configuration greatly influences simulated vegetation structure. In particular, simulations where height limitation was dependent on environmental stress showed a much better fit to canopy height observations. Spatial patterns of simulated LAI were more realistic than for uncoupled simulations of the forest landscape model, although some model deficiencies are still evident. Under idealized climate change scenarios, the effect of the coupling varied regionally, with the greatest effects on simulated streamflow (up to 60 mm yr−1 difference with respect to a simulation with static vegetation parameters) seen at the valley bottom and in regions currently above the treeline. This case study shows the importance of coupling hydrology and vegetation dynamics to simulate the impact of climate change on ecosystems. Nevertheless, it also highlights some challenges of ecohydrological modeling, such as the need to realistically simulate the plant response to increased CO2 concentrations and process uncertainty regarding future land cover changes., Geoscientific Model Development, 13 (2), ISSN:1991-9603, ISSN:1991-959X