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Agnara madagascariensis (Budde-Lund, 1885) Figs 1, 2K Metoponorthus Madagascariensis Budde-Lund, 1885: 189. Additional references Schmalfuss & Ferrara (1978); Ferrara & Taiti (1986). Material examined COLOMBIA ��� Bol��var, Cartagena de Indias, Islas del Rosario, Isla Grande ��� 5 ♂♂, 6 ♀♀; Ca��o Rat��n, Casa Comunal; 10��10���39.06��� N, 75��44���50.4��� W; 8 Sep. 2017; C.M. L��pez-Orozco, R. Borja- Arrieta, Y.M. Carpio-D��az and A. Figueroa leg.; CUDC-CRU 138 ��� 2 ♀♀; Orika; 10��10���37.2��� N, 75��44���27.59��� W; 8 Sep. 2017; A. Figueroa leg.; CUDC-CRU 139 ��� 13 ♂♂, 49 ♀♀; La Punta, Casa Sra. Miriam; 10��10���46.91��� N, 75��43���46.73��� W; 8 Sep. 2017; C.M. L��pez-Orozco, R. Borja-Arrieta and Y.M. Carpio-D��az leg.; CUDC-CRU 140. Distribution Ascension Island, Senegal, Guinea Bissau, Arabian Peninsula, Madagascar, southern China, Taiwan and Venezuela (Ashmole & Ashmole 2000; Schmidt 2001; Schmalfuss 2003). First record for Colombia., Published as part of L��pez-Orozco, Carlos Mario, Carpio-D��az, Yesenia M., Borja-Arrieta, Ricardo, Navas-S, Gabriel R., Campos-Filho, Ivanklin Soares, Taiti, Stefano, Mateos, Mariana, Olazaran, Alexandra, Caballero, Isabel C., Jotty, Karick, G��mez-Estrada, Harold & Hurtado, Luis A., 2022, A glimpse into a remarkable unknown diversity of oniscideans along the Caribbean coasts revealed on a tiny island, pp. 1-50 in European Journal of Taxonomy 793 on page 37, DOI: 10.5852/ejt.2022.793.1643, http://zenodo.org/record/6037892, {"references":["Budde-Lund G. 1885. Crustacea Isopoda terrestria per familias et genera et species descripta. Nielsen & Lydiche, Copenhagen, Denmark. https: // doi. org / 10.5962 / bhl. title. 109769","Schmalfuss H. & Ferrara F. 1978. Terrestrial isopods from West Africa, Part 2: families Tylidae, Ligiidae, Trichoniscidae, Styloniscidae, Rhyscotidae, Halophilosciidae, Philosciidae, Platyarthridae, Rhyscotidae, Trachelipidae, Porcellionidae, Armadillidiidae. Monitore zoologico italiano, nuova serie, Supplemento 11: 15 - 9. https: // doi. org / 10.1080 / 03749444.1978.10736575","Ferrara F. & Taiti S. 1986. The terrestral isopods (Oniscidea) of the Arabian Peninsula. Fauna of Saudi Arabia 7: 93 - 121.","Ashmole P. & Ashmole M. 2000. St Helena and Ascension Island: a Natural History. Anthony Nelson, Oswestry, England.","Schmidt C. 2001. Lista preliminar de los isopodos terrestres (Crustacea, Isopoda, Oniscidea) de Venezuela. Boletin de la Sociedad venezolana de Espeleologia 35: 1 - 12.","Schmalfuss H. 2003. World catalog of terrestrial isopods (Isopoda: Oniscidea). Stuttgarter Beitrage zur Naturkunde A 654: 1 - 341."]}

Family Halophilosciidae Verhoeff, 1908 Remarks The family Halophilosciidae comprises 34 species distributed in three genera, Halophiloscia Verhoeff, 1908 (9 spp.), Littorophiloscia Hatch, 1947 (22 spp.), and Stenophiloscia Verhoeff, 1908 (3 spp.) (WoRMS 2020c). In Colombia, Pearse (1915) recorded Philoscia culebrae Moore, 1901 [= Littorophiloscia c.] in coffee plantations from Hacienda Cincinati and Hacienda La Rosa, Sierra Nevada de Santa Marta (ca 1400 m a.s.l.), which is considered to be a misidentification by Van Name (1936). Littorophiloscia culebrae (Moore, 1901) is recorded from coastal areas of Angola, Brazil, Canary Islands, Caribbean islands, Florida (USA), Hawaii, Iraq, Madagascar, Socotra Archipelago and Yemen (Schmalfuss 2003; Taiti & Ferrara 2004; Araujo & Taiti 2007; Taiti & Lopez 2008; Naser et al. 2015; Lisboa et al. 2017; Campos-Filho et al. 2018). Recent expeditions to the north coast of Bol��var revealed the existence of this species in Colombia, inhabiting the coastal areas of the Bar�� Peninsula, Cartagena de Indias (C. L��pez-Orozco & Y. Carpio-D��az, pers. obs.)., Published as part of L��pez-Orozco, Carlos Mario, Carpio-D��az, Yesenia M., Borja-Arrieta, Ricardo, Navas-S, Gabriel R., Campos-Filho, Ivanklin Soares, Taiti, Stefano, Mateos, Mariana, Olazaran, Alexandra, Caballero, Isabel C., Jotty, Karick, G��mez-Estrada, Harold & Hurtado, Luis A., 2022, A glimpse into a remarkable unknown diversity of oniscideans along the Caribbean coasts revealed on a tiny island, pp. 1-50 in European Journal of Taxonomy 793 on page 29, DOI: 10.5852/ejt.2022.793.1643, http://zenodo.org/record/6037892, {"references":["Pearse A. S. 1915. An account of the Crustacea collected by the Walker Expedition to Santa Marta, Colombia. Proceedings of the United States National Museum 49: 531 - 556. https: // doi. org / 10.5479 / si. 00963801.49 - 2123.531","Van Name W. 1936. The American land and freshwater isopod Crustacea. Bulletin of the American Museum of Natural History 71: 1 - 535.","Schmalfuss H. 2003. World catalog of terrestrial isopods (Isopoda: Oniscidea). Stuttgarter Beitrage zur Naturkunde A 654: 1 - 341.","Taiti S. & Ferrara F. 2004. The terrestrial Isopoda (Crustacea: Oniscidea) of the Socotra Archipelago. Fauna of Arabia 20: 211 - 326.","Araujo P. B. & Taiti S. 2007. Terrestrial isopods (Crustacea, Oniscidea) from Rocas Atoll, northeastern Brazil. Arquivos do Museu Nacional 65: 347 - 355.","Taiti S. & Lopez H. 2008. New records and species of Halophilosciidae (Crustacea, Isopoda, Oniscidea) from the Canary Islands (Spain). In: Zimmer M., Charfi-Cheikhrouha F. & Taiti S. (eds) Proceedings of the International Symposium of Terrestrial Isopod Biology ISTIB- 07: 43 - 58. Shaker-Verlag, Aachen, Germany.","Naser M. D., Khalaf T. A. & Yasser A. G. 2015. First record of the terrestrial isopod Littorophiloscia culebrae (H. F. Moore, 1901) (Isopoda, Oniscidea) from Khor Al-Zubair, Iraq and the Persian Gulf. Crustaceana 88 (5): 611 - 615. https: // doi. org / 10.1163 / 15685403 - 00003432","Lisboa J. T., Campos-Filho I. S., Couto E. C. G. & Araujo P. B. 2017. Distribution of terrestrial isopods of the genus Littorophiloscia (Isopoda, Halophilosciidae) along the Brazilian coast. North-Western Journal of Zoology 13: e 167301.","Campos-Filho I. S., Cardoso G. M. & Aguiar J. O. 2018. Catalogue of terrestrial isopods (Crustacea, Isopoda, Oniscidea) from Brazil: an update with some considerations. Nauplius 26: e 2018038. https: // doi. org / 10.1590 / 2358 - 2936 e 2018038"]}

Halophiloscia trichoniscoides (Vandel, 1973) comb. nov. Figs 1, 2G, 15–17 Microphiloscia trichoniscoides Vandel, 1973: 170, figs 19–21. Material examined COLOMBIA – Bolívar, Cartagena de Indias, Islas del Rosario, Isla Grande • 3 ♂♂; Paraíso Secreto; 10°10′15.9″ N, 75°44′38.8″ W; 4 Apr. 2018; C.M. López-Orozco, R. Borja-Arrieta and K. Meza leg.; CUDC-CRU 218, 220, 221 • 1 ♂, 1 ♀ (parts in micropreparations); same locality as for preceding; 6 Sep. 2017; C.M. López-Orozco, Y.M. Carpio-Díaz and R. Borja-Arrieta leg.; CUDC-CRU 219. CUBA – Matanzas, Bahia de Conchinos • 4 ♂♂, 10 ♀♀; near Cueva de los Peces, between Playa Larga and Playa Giron; 10 Apr. 2002; S. Taiti leg.; MZUF 9607. – Pinar del Rio • 2 ♂♂, 3 ♀♀; Penisula Guanahacabibes, La Bajada; 7 Apr. 2002; S. Taiti leg.; MZUF 9608. Redescription MEASUREMENTS. Maximum body length: male 2.8 mm and female 4 mm. BODY. Color light brown (Fig. 2G), mottled with the usual pale muscle spots; posterior margins of pereonites 4–7, pleonites 3–5 and uropod branches more pigmented. Body outline as in Fig. 15A. Back smooth with numerous triangular scale-setae (Fig. 15B). One line of noduli laterales per side on the pereonites inserted at same distance from lateral margins (Fig. 15A, C). CEPHALON. Without frontal line, suprantennal line straight on median portion; eyes consisting of 10 ommatidia (Fig. 15D–E). PLEON. Narrower than pereon; epimera of pleonites 3–5 not developed. Telson with lateral sides slightly concave, distal margin broadly rounded (Fig. 15A). ANTENNULA. Composed of three articles subequal in length, distal article with two lateral aesthetascs plus apical pair (Fig. 15F). ANTENNA. When extended posteriorly reaches posterior margin of pereonite 5; flagellum as long as fifth article of peduncle, first article slightly longer than second and third, second and third articles with three and two lateral aesthetascs, respectively (Fig. 15G). MOUTH. Mandibles with molar penicil consisting of at least five branches, left mandible with 2+1 penicils (Fig. 16A), right mandible with 1+1 penicils (Fig. 16B). Maxillula (Fig. 16C) inner endite with two long penicils inserted transversely; outer endite of 5+5 teeth, inner set apically cleft. Maxilla (Fig. 16D) with bilobate apex, inner lobe rounded, bearing thick setae, outer lobe rounded, covered with thin setae plus three thick setae. Maxilliped (Fig. 16E) base bearing sparse setae, proximal article of palp with two stout setae slightly distinct in length; endite subrectangular, inner and distal margins bearing several thin setae, medial seta not surpassing distal margin, distal margin with one small hairy penicil. UROPOD. Protopod and exopod grooved on outer margin, bearing many glandular pores, exopod almost three times as long as endopod (Fig. 16F). PEREOPODS. Rather slender, pereopod 1 carpus bearing transverse antennal grooming brush and distal seta apically cleft; dactylus with ungual seta simple and surpassing outer claw, dactylar seta simple, not surpassing outer claw. Male PEREOPODS 1 AND 7. Pereopod 1 (Fig. 17A) propodus slightly flattened, carpus with transverse antennal grooming brush and distal seta hand-like. Pereopod 7 (Fig. 17B) without any sexual dimorphism. GENITAL PAPILLA. Ventral shield ovoid, papilla bifurcate with genital orifices opening at apex (Fig. 17C). PLEOPODS. Pleopod 1 (Fig. 17D) exopod triangular, inner margin almost straight, outer margin sinuous, distal portion triangular with distal margin rounded; endopod with thickest distal part with sides almost parallel, inner distal part with long pointed process concave on outer margin, prominent subquadrangular apical lobe on outer distal portion bearing thin setae. Pleopod 2 (Fig. 17E) exopod triangular, outer margin concave, bearing seven stout setae; endopod robust, distinctly longer than exopod. Exopods of pleopods 3–5 as in Fig. 17F–H. Remarks Vandel (1973) erected the genus Microphiloscia to allocate the new species M. trichoniscoides from Cueva de la Colorada, Provincia de Oriente, Sierra Maestra, Colorada del Maso, Cuba. After the examination of material belonging to this species, M. trichoniscoides shows the shape of the genital papilla and male pleopod 1 endopod typical of members of Halophiloscia (see also Taiti & Lopez 2008). Halophiloscia includes 9 halophilic species mainly distributed along the Mediterranean and Atlantic coasts of Europe and Africa (Schmalfuss 2003; Taiti & Lopez 2008; Taiti & Argano 2009, 2011). In America, only H. couchii (Kinahan, 1858) was recorded, from Argentina, Bermuda, and USA (see Schmidt 2003). The genus is mainly characterized by the runner-type habitus (sensu Schmalfuss 1984), epimera of pereonites 1–7 with one or more lines of noduli laterales, male pereopod 1 and sometimes pereopod 2 carpus and propodus enlarged and covered with several scales on frontal side, male genital papilla distally bifurcated, and male pleopod 1 endopod stout bearing a long, pointed process on the apex (see Schmidt 2003; Taiti & Lopez 2008). The species is included within the genus since it shows all the previously mentioned characters, except that the male pereopods 1 and 2 do not have the carpus and propodus enlarged. Thus, Microphiloscia is considered to be a junior synonym of Halophiloscia. Distribution Cuba (Vandel 1973). First record for Colombia.

Key to species of terrestrial isopods from Isla Grande 1. Eyes present....................................................................................................................................... 2 ��� Eyes absent............................... Stenoniscus nestori L��pez-Orozco, Taiti & Campos-Filho sp. nov. 2. Animal with conglobation ability...................................................................................................... 3 ��� Animal without conglobation ability................................................................................................. 6 3. Pereon dorsally smooth; pereonite 5 epimera with ventral phylacomera.......................................... 4 ��� Pereon dorsally tuberculate; pereonite 5 epimera without phylacomera..................................................................................................................................... Ctenorillo tuberosus (Budde-Lund, 1904) 4. Phylacomera 5 paddle-shaped, extended anteriorly.......................................................................... 5 ��� Phylacomera 5 sickle-shaped.................................................. Tylos marcuzzii Giordani Soika, 1954 5. Inner lobe of schisma extending beyond posterior margin of outer lobe.............................................................................................. Tylos negroi L��pez-Orozco, Carpio-D��az & Campos-Filho sp. nov. ��� Inner lobe of schisma not extending beyond posterior margin of outer lobe................................................................................................................................................ Tylos niveus Budde-Lund, 1885 6. Flagellum of the antennae with 4 or more articles............................................................................ 7 ��� Flagellum of the antennae with 2���3 articles.................................................................................... 10 7. Flagellum of the antennae with 4 articles.......................................................................................... 8 ��� Flagellum of the antennae with more than 10 articles......... Ligia baudiniana Milne-Edwards, 1840 8. Dorsal surface almost smooth or with small tubercles...................................................................... 9 ��� Dorsal surface of pereon with 4 prominent longitudinal ribs..................................................................................................................................... Armadilloniscus caraibicus Paoletti & Stinner, 1989 9. Dorsal surface almost smooth.................................................. Armadilloniscus ninae Schultz, 1984 ��� Dorsal surface with small tubercles....................................................................................................................................................... Armadilloniscus luisi Carpio-D��az, Taiti & Campos-Filho sp. nov. 10. Flagellum of the antennae with 3 articles.........................................................................................11 ��� Flagellum of the antennae with 2 articles........................................................................................ 14 11. Genital papilla with truncate apex................................................................................................... 12 ��� Genital papilla with bifurcated apex....... Halophiloscia trichoniscoides (Vandel, 1973) comb. nov. 12. Male pleopod 1 endopod without lobe on outer margin of apical part............................................ 13 ��� Male pleopod 1 endopod with a lobe on outer margin of apical part.................................................................................................................................. Littorophiloscia tropicalis Taiti & Ferrara, 1986 13. Male pleopod 1 endopod with apex divided into three points........................................................................................................................................ Littorophiloscia amphindica Taiti & Ferrara, 1986 ��� Male pleopod 1 endopod with a long spine and 2 small distinct denticles on the inner margin of the apical part.......................................................... Littorophiloscia denticulata (Ferrara & Taiti, 1982) 14. Pleopod exopods with lungs............................................................................................................ 15 ��� Pleopod exopods without lungs....................................................................................................... 16 15. Pleopodal lungs present in first two pairs of pleopods...... Porcellionides pruinosus (Brandt, 1833) ��� Pleopodal lungs present in all five pairs of pleopods.......................................................................................................................................................... Agnara madagascariensis (Budde-Lund, 1885) 16. Colorless body; eyes composed of two unequal ommatidia........................................................................................................................................ Trichorhina heterophthalma Lemos de Castro, 1964 ��� Pigmented body; eyes with 10 ommatidia........................................................................................................................... Trichorhina bermudezae Carpio-D��az, L��pez-Orozco & Campos-Filho, 2018, Published as part of L��pez-Orozco, Carlos Mario, Carpio-D��az, Yesenia M., Borja-Arrieta, Ricardo, Navas-S, Gabriel R., Campos-Filho, Ivanklin Soares, Taiti, Stefano, Mateos, Mariana, Olazaran, Alexandra, Caballero, Isabel C., Jotty, Karick, G��mez-Estrada, Harold & Hurtado, Luis A., 2022, A glimpse into a remarkable unknown diversity of oniscideans along the Caribbean coasts revealed on a tiny island, pp. 1-50 in European Journal of Taxonomy 793 on pages 40-41, DOI: 10.5852/ejt.2022.793.1643, http://zenodo.org/record/6037892, {"references":["Budde-Lund G. 1904. A Revision of Crustacea Isopoda Terrestria, with Additions and Illustrations. Pt 1 Eubelum, Pt 2 Spherilloninae, Pt 3 Armadillo. H. Hagerup, Copenhagen, Denmark.","Giordani Soika A. 1954. Ecologia, sistematica, biogeografia ed evoluzione del Tylos latreillei. Bollettino del Museo civico di Storia naturale di Venezia 7: 63 - 83.","Budde-Lund G. 1885. Crustacea Isopoda terrestria per familias et genera et species descripta. Nielsen & Lydiche, Copenhagen, Denmark. https: // doi. org / 10.5962 / bhl. title. 109769","Milne-Edwards M. 1840. Ordre des isopodes. In: Histoire Naturelle des Crustaces, comprenant l'Anatomie, la Physiologie et la Classficiation de ces Animaux. Tome Troisieme. 115 - 283. Librairie Encyclopedique de Roret, Paris.","Paoletti M. G. & Stinner B. R. 1989. Two new terrestrial Isopoda (Oniscidea) from coralline cays of Venezuela's Caribbean coast. Proceedings of the Entomological Society of Washington 91 (1): 71 - 80. Available from https: // biostor. org / reference / 56403 [accessed 6 Jan. 2022].","Vandel A. 1973. Les isopodes terrestres et cavernicoles de l'ile de Cuba. In: Orghidan T., Nunez A., Botosaneanu L., Decou V., Negrea S. & Vina N. (eds) Resultats des Expeditions biospeologiques cubanoroumaines a Cuba, vol. 1: 153 - 188. Editura Academiei Republicii Socialiste Romania, Bucharest.","Ferrara F. & Taiti S. 1986. The terrestral isopods (Oniscidea) of the Arabian Peninsula. Fauna of Saudi Arabia 7: 93 - 121.","Ferrara F. & Taiti S. 1982. Isopodi terrestri delle Isole Andamane. Bollettino del Museo civico di Storia naturale di Verona 8: 459 - 492.","Brandt J. F. 1833. Conspectus Monographiae Crustaceorum Oniscodorum Latreillii. Byulleten Moskovskogo Obshchestva Ispytatelei Prirody 6: 171 - 193.","Lemos de Castro A. 1964. Trichorhina heterophthalma nueva especie de isopodo terrestre cavernicola de Cuba. Poeyana 2 A: 1 - 7.","Carpio-Diaz Y. M., Lopez-Orozco C. M., Campos-Filho I. S. & Navas G. R. 2018. Terrestrial isopods (Isopoda: Oniscidea) of the Botanical Garden of Cartagena \" Guillermo Pineres \", Colombia, with the description of three new species. Arthropoda Selecta 27 (4): 301 - 318. https: // doi. org / 10.15298 / arthsel. 27.4.05"]}

Halophiloscia trichoniscoides (Vandel, 1973) comb. nov. Figs 1, 2G, 15���17 Microphiloscia trichoniscoides Vandel, 1973: 170, figs 19���21. Material examined COLOMBIA ��� Bol��var, Cartagena de Indias, Islas del Rosario, Isla Grande ��� 3 ♂♂; Para��so Secreto; 10��10���15.9��� N, 75��44���38.8��� W; 4 Apr. 2018; C.M. L��pez-Orozco, R. Borja-Arrieta and K. Meza leg.; CUDC-CRU 218, 220, 221 ��� 1 ♂, 1 ♀ (parts in micropreparations); same locality as for preceding; 6 Sep. 2017; C.M. L��pez-Orozco, Y.M. Carpio-D��az and R. Borja-Arrieta leg.; CUDC-CRU 219. CUBA ��� Matanzas, Bahia de Conchinos ��� 4 ♂♂, 10 ♀♀; near Cueva de los Peces, between Playa Larga and Playa Giron; 10 Apr. 2002; S. Taiti leg.; MZUF 9607. ��� Pinar del Rio ��� 2 ♂♂, 3 ♀♀; Penisula Guanahacabibes, La Bajada; 7 Apr. 2002; S. Taiti leg.; MZUF 9608. Redescription MEASUREMENTS. Maximum body length: male 2.8 mm and female 4 mm. BODY. Color light brown (Fig. 2G), mottled with the usual pale muscle spots; posterior margins of pereonites 4���7, pleonites 3���5 and uropod branches more pigmented. Body outline as in Fig. 15A. Back smooth with numerous triangular scale-setae (Fig. 15B). One line of noduli laterales per side on the pereonites inserted at same distance from lateral margins (Fig. 15A, C). CEPHALON. Without frontal line, suprantennal line straight on median portion; eyes consisting of 10 ommatidia (Fig. 15D���E). PLEON. Narrower than pereon; epimera of pleonites 3���5 not developed. Telson with lateral sides slightly concave, distal margin broadly rounded (Fig. 15A). ANTENNULA. Composed of three articles subequal in length, distal article with two lateral aesthetascs plus apical pair (Fig. 15F). ANTENNA. When extended posteriorly reaches posterior margin of pereonite 5; flagellum as long as fifth article of peduncle, first article slightly longer than second and third, second and third articles with three and two lateral aesthetascs, respectively (Fig. 15G). MOUTH. Mandibles with molar penicil consisting of at least five branches, left mandible with 2+1 penicils (Fig. 16A), right mandible with 1+1 penicils (Fig. 16B). Maxillula (Fig. 16C) inner endite with two long penicils inserted transversely; outer endite of 5+5 teeth, inner set apically cleft. Maxilla (Fig. 16D) with bilobate apex, inner lobe rounded, bearing thick setae, outer lobe rounded, covered with thin setae plus three thick setae. Maxilliped (Fig. 16E) base bearing sparse setae, proximal article of palp with two stout setae slightly distinct in length; endite subrectangular, inner and distal margins bearing several thin setae, medial seta not surpassing distal margin, distal margin with one small hairy penicil. UROPOD. Protopod and exopod grooved on outer margin, bearing many glandular pores, exopod almost three times as long as endopod (Fig. 16F). PEREOPODS. Rather slender, pereopod 1 carpus bearing transverse antennal grooming brush and distal seta apically cleft; dactylus with ungual seta simple and surpassing outer claw, dactylar seta simple, not surpassing outer claw. Male PEREOPODS 1 AND 7. Pereopod 1 (Fig. 17A) propodus slightly flattened, carpus with transverse antennal grooming brush and distal seta hand-like. Pereopod 7 (Fig. 17B) without any sexual dimorphism. GENITAL PAPILLA. Ventral shield ovoid, papilla bifurcate with genital orifices opening at apex (Fig. 17C). PLEOPODS. Pleopod 1 (Fig. 17D) exopod triangular, inner margin almost straight, outer margin sinuous, distal portion triangular with distal margin rounded; endopod with thickest distal part with sides almost parallel, inner distal part with long pointed process concave on outer margin, prominent subquadrangular apical lobe on outer distal portion bearing thin setae. Pleopod 2 (Fig. 17E) exopod triangular, outer margin concave, bearing seven stout setae; endopod robust, distinctly longer than exopod. Exopods of pleopods 3���5 as in Fig. 17F���H. Remarks Vandel (1973) erected the genus Microphiloscia to allocate the new species M. trichoniscoides from Cueva de la Colorada, Provincia de Oriente, Sierra Maestra, Colorada del Maso, Cuba. After the examination of material belonging to this species, M. trichoniscoides shows the shape of the genital papilla and male pleopod 1 endopod typical of members of Halophiloscia (see also Taiti & Lopez 2008). Halophiloscia includes 9 halophilic species mainly distributed along the Mediterranean and Atlantic coasts of Europe and Africa (Schmalfuss 2003; Taiti & Lopez 2008; Taiti & Argano 2009, 2011). In America, only H. couchii (Kinahan, 1858) was recorded, from Argentina, Bermuda, and USA (see Schmidt 2003). The genus is mainly characterized by the runner-type habitus (sensu Schmalfuss 1984), epimera of pereonites 1���7 with one or more lines of noduli laterales, male pereopod 1 and sometimes pereopod 2 carpus and propodus enlarged and covered with several scales on frontal side, male genital papilla distally bifurcated, and male pleopod 1 endopod stout bearing a long, pointed process on the apex (see Schmidt 2003; Taiti & Lopez 2008). The species is included within the genus since it shows all the previously mentioned characters, except that the male pereopods 1 and 2 do not have the carpus and propodus enlarged. Thus, Microphiloscia is considered to be a junior synonym of Halophiloscia. Distribution Cuba (Vandel 1973). First record for Colombia., Published as part of L��pez-Orozco, Carlos Mario, Carpio-D��az, Yesenia M., Borja-Arrieta, Ricardo, Navas-S, Gabriel R., Campos-Filho, Ivanklin Soares, Taiti, Stefano, Mateos, Mariana, Olazaran, Alexandra, Caballero, Isabel C., Jotty, Karick, G��mez-Estrada, Harold & Hurtado, Luis A., 2022, A glimpse into a remarkable unknown diversity of oniscideans along the Caribbean coasts revealed on a tiny island, pp. 1-50 in European Journal of Taxonomy 793 on pages 29-32, DOI: 10.5852/ejt.2022.793.1643, http://zenodo.org/record/6037892, {"references":["Vandel A. 1973. Les isopodes terrestres et cavernicoles de l'ile de Cuba. In: Orghidan T., Nunez A., Botosaneanu L., Decou V., Negrea S. & Vina N. (eds) Resultats des Expeditions biospeologiques cubanoroumaines a Cuba, vol. 1: 153 - 188. Editura Academiei Republicii Socialiste Romania, Bucharest.","Taiti S. & Lopez H. 2008. New records and species of Halophilosciidae (Crustacea, Isopoda, Oniscidea) from the Canary Islands (Spain). In: Zimmer M., Charfi-Cheikhrouha F. & Taiti S. (eds) Proceedings of the International Symposium of Terrestrial Isopod Biology ISTIB- 07: 43 - 58. Shaker-Verlag, Aachen, Germany.","Schmalfuss H. 2003. World catalog of terrestrial isopods (Isopoda: Oniscidea). Stuttgarter Beitrage zur Naturkunde A 654: 1 - 341.","Taiti S. & Argano R. 2009. New species of terrestrial isopods from Sardinia (Isopoda: Oniscidea). Zootaxa 2318 (1): 38 - 55. https: // doi. org / 10.11646 / zootaxa. 2318.1.5","Taiti S. & Argano R. 2011. Oniscidea di Sardegna (Crustacea, Isopoda). Conservazione Habitat Invertebrati 5: 163 - 222.","Schmidt C. 2003. Contribution to the phylogenetic system of the Crinocheta (Crustacea, Isopoda). Part 2 (Oniscoidea to Armadillidiidae). Mitteilungen aus dem Zoologischen Museum in Berlin, Zoologische Reihe 79: 3 - 179. https: // doi. org / 10.1002 / mmnz. 20030790102","Schmalfuss H. 1984. Eco-morphological strategies in terrestrial isopods. Proceedings of the Zoological Society of London 53: 49 - 63."]}

Tylos marcuzzii Giordani Soika, 1954 Figs 1, 2A Tylos marcuzzii Giordani Soika, 1954: 79, fig. 10/1. Additional references Schultz (1974); Kensley & Schotte (1989). Material examined COLOMBIA ��� Bol��var, Cartagena de Indias, Islas del Rosario, Isla Grande ��� 1 ♀; Playa Libre; 10��10���52.4��� N, 75��43���56.1��� W; 7 Sep. 2017; C.M. L��pez-Orozco, R. Borja-Arrieta and Y.M. Carpio- D��az leg.; CUDC-CRU 136 ��� 11 ♂♂, 39 ♀♀, 1 juv.; La Punta, Hotel Majagua; 10��10���56.41��� N, 75��43���33.51��� W; 6 Oct. 2017; same collectors as for preceding; CUDC-CRU 137. Distribution Florida, Bahamas, Leeward Islands, Belize, Venezuela and Cuba (Schultz 1974; Kensley & Schotte 1989; Schmalfuss & Vergara 2000; Schmidt 2001; Schmalfuss 2003; Hurtado et al. 2014). First record for Colombia., Published as part of L��pez-Orozco, Carlos Mario, Carpio-D��az, Yesenia M., Borja-Arrieta, Ricardo, Navas-S, Gabriel R., Campos-Filho, Ivanklin Soares, Taiti, Stefano, Mateos, Mariana, Olazaran, Alexandra, Caballero, Isabel C., Jotty, Karick, G��mez-Estrada, Harold & Hurtado, Luis A., 2022, A glimpse into a remarkable unknown diversity of oniscideans along the Caribbean coasts revealed on a tiny island, pp. 1-50 in European Journal of Taxonomy 793 on page 9, DOI: 10.5852/ejt.2022.793.1643, http://zenodo.org/record/6037892, {"references":["Giordani Soika A. 1954. Ecologia, sistematica, biogeografia ed evoluzione del Tylos latreillei. Bollettino del Museo civico di Storia naturale di Venezia 7: 63 - 83.","Schultz G. A. 1974. Terrestrial isopod crustaceans (Oniscoidea) mainly from the West Indies and adjacent regions: 1. Tylos and Ligia. Studies on the fauna of Cura ҫao and other Caribbean islands 149: 162 - 173. Available from http: // www. repository. naturalis. nl / document / 549820 [accessed 7 Jan. 2022].","Kensley B. & Schotte M. 1989. Guide to the Marine Isopod Crustaceans of the Caribbean. Smithsonian Institution Press, Washington. Available from https: // www. biodiversitylibrary. org / page / 10950583 [accessed 6 Jan. 2022].","Schmalfuss H. & Vergara K. 2000. The isopod genus Tylos (Oniscidea: Tylidae) in Chile, with bibliographies of all described species of the genus. Stuttgarter Beitrage zur Naturkunde A 612: 1 - 42.","Schmidt C. 2001. Lista preliminar de los isopodos terrestres (Crustacea, Isopoda, Oniscidea) de Venezuela. Boletin de la Sociedad venezolana de Espeleologia 35: 1 - 12.","Schmalfuss H. 2003. World catalog of terrestrial isopods (Isopoda: Oniscidea). Stuttgarter Beitrage zur Naturkunde A 654: 1 - 341.","Hurtado L. A., Lee E. J., Mateos M. & Taiti S. 2014. Global diversification at the harsh sea-land interface: mitochondrial phylogeny of the supralittoral isopod genus Tylos (Tylidae, Oniscidea). PloS ONE 9 (4): e 94081. https: // doi. org / 10.1371 / journal. pone. 0094081"]}

Littorophiloscia amphindica Taiti & Ferrara, 1986 Figs 1, 2H Littorophiloscia amphindica Taiti & Ferrara, 1986: 1369, fig. 16. Material examined COLOMBIA ��� Bol��var, Cartagena de Indias, Islas del Rosario, Isla Grande ��� 5 ♂♂, 4 ♀♀; El Silencio, Laguna Caracol��; 10��10���28.4��� N, 75��44���29.96��� W; 6 Oct. 2017; Y.M. Carpio-D��az and R. Borja-Arrieta leg.; CUDC-CRU 223 ��� 1 ♂, 1 ♀; El Silencio, Laguna del Silencio; 10��10���34��� N, 75��44���26.8��� W; 25 Aug. 2017; leg. Y.M. Carpio-D��az, C.M. L��pez-Orozco and R. Borja-Arrieta; CUDC- CRU 224 ��� 1 ♂, 4 ♀♀; Laguna La Charca; 10��10���47.8��� N, 75��44���0.4��� W; 7 Sep. 2017; same collectors as for preceding; CUDC-CRU 225 ��� 2 ♂♂, 3 ♀♀; Hotel La Pola; 10��10���19.4��� N, 75��44���48.9��� W; 24 Nov. 2017; same collectors as for preceding; CUDC-CRU 226 ��� 1 ♂, 1 ♀; Ca��o Rat��n; 10��10���42.6��� N, 75��44���52.6��� W; 24 Nov. 2017; C.M. L��pez-Orozco leg.; CUDC-CRU 227 ��� 1 ♂, 1 ♀; 10��10���33.4��� N, 75��44���52.4��� W; 24 Nov. 2017; C.M. L��pez-Orozco, Y.M. Carpio-D��az and R. Borja-Arrieta leg.; CUDC- CRU 228 ��� 1 ♂, 3 ♀♀; same collection data as for preceding; 10��10���44.91��� N, 75��44���53.01��� W; CUDC- CRU 229 ��� 1 ♂, 1 ♀; La Punta, Laguna Encantada; 10��10���48.3��� N, 75��43���38.9��� W; 25 Aug. 2017; C.M. L��pez-Orozco, Y.M. Carpio-D��az and R. Borja-Arrieta leg.; CUDC-CRU 230 ��� 5 ♂♂, 6 ♀♀; same collection data as for preceding; 10��10���48.1��� N, 75��43���39.1��� W; CUDC-CRU 231 ��� 9 ♀♀; same collection data as for preceding; 10��10���48.1��� N, 75��43���37.5��� W; CUDC-CRU 232 ��� 1 ♀; same collection data as for preceding; 10��10���48.4��� N, 75��43���38.4��� W; CUDC-CRU 233 ��� 1 ♀; same collection data as for preceding; 10��10���48��� N, 75��43���39.3��� W; CUDC-CRU 234 ��� 1 ♂, 5 ♀♀; same collection data as for preceding; 10��10���48.1��� N, 75��43���37.5��� W; 7 Sep. 2017; CUDC-CRU 235 ��� 3 ♂♂, 1 ♀; Para��so Secreto; 10��10���15.9���N, 75��44���38.8��� W; 24 Nov.2017; C.M. L��pez-Orozco leg.; CUDC-CRU236 ��� 13 ♂♂, 13 ♀♀, 8 juvs; same locality and collector as for preceding, 6 Sep. 2017; CUDC-CRU 237 ��� 14 ♂♂, 32 ♀♀, 3 juvs; same locality as for preceding; 4Apr. 2018; C.M. L��pez-Orozco, R. Borja-Arrieta and K. Meza leg.; CUDC-CRU 238. Distribution Tanzania (Zanz��bar), Maldives, Comoro Islands, Indonesia (Bali) and Taiwan (Schmalfuss 2003; Taiti 2014). First record for Colombia., Published as part of L��pez-Orozco, Carlos Mario, Carpio-D��az, Yesenia M., Borja-Arrieta, Ricardo, Navas-S, Gabriel R., Campos-Filho, Ivanklin Soares, Taiti, Stefano, Mateos, Mariana, Olazaran, Alexandra, Caballero, Isabel C., Jotty, Karick, G��mez-Estrada, Harold & Hurtado, Luis A., 2022, A glimpse into a remarkable unknown diversity of oniscideans along the Caribbean coasts revealed on a tiny island, pp. 1-50 in European Journal of Taxonomy 793 on page 34, DOI: 10.5852/ejt.2022.793.1643, http://zenodo.org/record/6037892, {"references":["Ferrara F. & Taiti S. 1986. The terrestral isopods (Oniscidea) of the Arabian Peninsula. Fauna of Saudi Arabia 7: 93 - 121.","Schmalfuss H. 2003. World catalog of terrestrial isopods (Isopoda: Oniscidea). Stuttgarter Beitrage zur Naturkunde A 654: 1 - 341.","Taiti S. 2014. The terrestrial Isopoda (Crustacea, Oniscidea) of the Maldives. Tropical Zoology 27 (1): 9 - 33. https: // doi. org / 10.1080 / 03946975.2014.894397"]}

Armadilloniscus luisi Carpio-D��az, Taiti & Campos-Filho sp. nov. urn:lsid:zoobank.org:act: 88600E5C-BFF9-49BA-976C-8853E33A6E58 Figs 1, 2E, 10���12 Etymology The new species is named after Mr Luis Felipe Arrieta Z����iga, for his interest in nature and support to Ricardo Borja-Arrieta as a biologist. Material examined Holotype COLOMBIA ��� Bol��var, Cartagena de Indias, Islas del Rosario, Isla Grande ��� ♂; Laguna del Silencio, El Silencio; 10��10���35.4��� N, 75��44���26��� W; 24 Aug. 2017; C.M. L��pez-Orozco, Y.M. Carpio-D��az and R. Borja-Arrieta leg.; CUDC-CRU 193. Paratypes COLOMBIA ��� Bol��var, Cartagena de Indias, Islas del Rosario, Isla Grande ��� 1 ♂, 1 ♀ (parts in micropreparations); same collection data as for holotype; CUDC-CRU 194 ��� 10 ♂♂, 5 ♀♀; same collection data as for holotype; CUDC-CRU 195 ��� 1 ♀; 10��10���32.9��� N, 75��44���27.3��� W; same date and collectors as for holotype; CUDC-CRU 206 ��� 1 ♀; 10��10���33.5��� N, 75��44���26.9��� W; same date and collectors as for holotype; CUDC-CRU 207 ��� 1 ♂; 10��10���35��� N, 75��44���26.4��� W; same date and collectors as for holotype; CUDC-CRU 208 ��� 1 ♂; 10��10���34.8��� N, 75��44���26.6��� W; same date and collectors as for holotype; CUDC-CRU 209 ��� 6 ♂♂, 12 ♀♀; same collection data as for holotype; 10��10���35��� N, 75��44���16.4��� W; CUDC-CRU 210 ��� 14 ♂♂, 10 ♀♀; 10��10���34��� N, 75��44���26.8��� W; 25 Aug. 2017; same locality and collectors as for holotype; CUDC-CRU 211 ��� 3 ♂♂; La Punta, Laguna Encantada; 10��10���48.4��� N, 75��43���38.4��� W; 25 Aug. 2017; C.M. L��pez-Orozco, Y.M. Carpio-D��az and R. Borja- Arrieta leg.; CUDC-CRU 196 ��� 1 ♂; same locality as for preceding; 10��10���47.72��� N, 75��43���40.09��� W; same date and collectors as for holotype; CUDC-CRU 197 ��� 1 ♂, 1 ♀; same locality as for preceding; 10��10���48��� N, 75��43���39.3��� W; same date and collectors as for holotype; CUDC-CRU 198 ��� 1 ♀; same locality as for preceding; 10��10���47.7��� N, 75��43���39.93��� W; same date and collectors as for holotype; CUDC-CRU 199 ��� 2 ♂♂, 3 ♀♀; same locality as for preceding; 10��10���48.3��� N, 75��43���38.9��� W; same date and collectors as for holotype; CUDC-CRU 200 ��� 3 ♀♀; same locality as for preceding; 10��10���48.1��� N, 75��43���39.1��� W; same date and collectors as for holotype; CUDC-CRU 201 ��� 2 ♀♀; same locality as for preceding; 10��10���48.1��� N, 75��43���37.5��� W; same date and collectors as for holotype; CUDC-CRU 202 ��� 4 ♂♂, 4 ♀♀; same locality as for preceding; 10��10���47.72��� N, 75��43���40.09��� W; same date and collectors as for holotype; CUDC-CRU 203 ��� 2 ♂♂, 2 ♀♀; same locality as for preceding; 7 Sep. 2017; same date and collectors as for holotype; CUDC-CRU 204 ��� 3 ♂♂, 4 ♀♀; Laguna La Charca; 10��10���47.8��� N, 75��44���0.4��� W; 7 Sep. 2017; C.M. L��pez-Orozco, Y.M. Carpio-D��az and R. Borja-Arrieta leg.; CUDC- CRU 205. Description MEASUREMENTS. Maximum length: male 2.5 mm; female 3.1 mm. BODY. Color brown with usual pale muscle spots; antennae brown with pale flagellum (Fig. 2E). Body elongated and elliptical-shaped (Fig. 10A, H). Dorsal surface bearing elongated scale-setae (Fig. 10B). Dorsum covered with broad tubercles (Fig. 10A, C���E, H): cephalon bearing seven tubercles in two rows, anterior row with four and posterior row with three; ♂ pereonites 1���3 with one row of 10 tubercles, 4���7 with one row of eight tubercles; ♀ pereonites 1, 5 and 6 with one row of 10 tubercles, 2���4 and 7 with one row of eight tubercles; ♂ pleonites 1���5 and telson with two paramedian tubercles, pleonite 2 with two broad tubercles at sides, pleonite 3 with two small additional tubercles; ♀ pleonites 1���5 and telson with two paramedian tubercles, pleonites 2 and 3 with two additional small tubercles at sides. CEPHALON. Lateral lobes well developed and directed outwards, median lobe triangular, slightly surpassing distal margin of lateral lobes, frontal and suprantennal lines absent; eyes consisting of four ommatidia (Fig. 10C���D). PLEON. Outline continuous with that of pereonite 7, epimera of pleonites 3���5 rectangular (Fig. 10A, E, H). Telson (Fig. 10E) triangular, lateral margins almost straight, distal part truncate and not surpassing distal margin of pleonite 5 epimera. ANTENNULA. Composed of two articles, second article bearing a few lateral setae, distal margin bearing one flagellar seta and two aesthetascs (Fig. 10F). ANTENNA. When extended posteriorly slightly surpassing pereonite 1, flagellum of four articles (Fig. 10G). MOUTH. Mandibles with molar penicil of several plumose setae, left mandible with 2+1 penicils (Fig. 11A), right with 1+1 penicils (Fig. 11B). Maxillula (Fig. 11C) inner endite with two penicils; outer endite of 4+8 teeth, five of them pectinated, and one slender stalk between outer and inner set, outer distal margin bearing fringe of long thin setae. Maxilla (Fig. 11D) with lobes fused, inner middle portion with five long setae, distal margin rounded, bearing several thin and thick setae. Maxilliped (Fig. 11E) palp bearing two tufts of setae on first article, distal articles of palp fused, bearing several thin and thick setae; endite rectangular, inner margin bearing fringe of long thin setae plus four long thick setae, distal margin bilobed, bearing broad rounded penicil with long thick setae. UROPOD. Protopod enlarged with distal margin rounded, exopod not surpassing protopod, endopod longer than exopod (Fig. 11F). PEREOPODS. Pereopods 1���7 stout, merus to propodus bearing sparse setae on sternal margin, carpus 1 with strong distal seta bearing long sensilla, dactylus with very long ungual seta, dactylar seta elongated, apically cleft and plumose. Male PEREOPODS 1 AND 7. Without any sexual modifications (Fig. 12A���B). GENITAL PAPILLA. Ventral shield triangular, papilla slightly cleft at apex, bearing small setae (Fig. 12C). PLEOPODS. Pleopod 1 (Fig. 12D) exopod ovoid; endopod three times as long as exopod, apical portion bent outwards. Pleopod 2 (Fig. 12E) exopod subtriangular; endopod with distal article flagelliform. Exopods of pleopods 3���5 as in Fig. 12F���H. Remarks In the Caribbean region, four species of Armadilloniscus have been recorded: A. caraibicus from Venezuela, A. ellipticus (Harger, 1878) from the Atlantic coasts of USA and Bermuda, A. ninae Schultz, 1984 from Belize, and A. steptus Schotte & Heard, 1991 from Turks and Caicos Islands (British West Indies) (Schultz 1984; Paoletti & Stinner 1989; Taiti & Ferrara 1989; Schotte & Heard 1991). For a definition of the genus see Arcangeli (1957), Vandel (1962), Taiti & Ferrara (1989), and Schmidt (2002). Armadilloniscus luisi Carpio-D��az, Taiti & Campos-Filho sp. nov. is readily distinguished from all the other species of the genus in the number and arrangement of the dorsal tubercles on the cephalon, pereon and pleon. Moreover, it can be distinguished from the other four Caribbean species in having two broad tubercles on pleonite 2 (vs small in A. caraibicus and A. steptus, absent in A. ninae), the cephalon with the median lobe triangular (vs rounded in A. caraibicus) and lateral lobes subquadrangular (vs subtriangular in A. ninae and A. steptus), eyes composed of four ommatidia (vs 4���5 in A. caraibicus, 6 in A. ellipticus and 5 in A. ninae), telson with distal margin truncate (vs rounded in A. caraibicus and A. ellipticus, triangular in A. ninae), and ♂ pleopod 1 exopod ovoid (vs subtriangular in A. caraibicus, A. ellipticus and A. ninae, almost rounded in A. steptus) (Paoletti & Stinner 1989; Schultz 1984; Schotte & Heard 1991; Garthwaite et al. 1992; Schmidt 2002). Distribution Presently known only from Isla Grande, Cartagena de Indias, Colombia., Published as part of L��pez-Orozco, Carlos Mario, Carpio-D��az, Yesenia M., Borja-Arrieta, Ricardo, Navas-S, Gabriel R., Campos-Filho, Ivanklin Soares, Taiti, Stefano, Mateos, Mariana, Olazaran, Alexandra, Caballero, Isabel C., Jotty, Karick, G��mez-Estrada, Harold & Hurtado, Luis A., 2022, A glimpse into a remarkable unknown diversity of oniscideans along the Caribbean coasts revealed on a tiny island, pp. 1-50 in European Journal of Taxonomy 793 on pages 21-26, DOI: 10.5852/ejt.2022.793.1643, http://zenodo.org/record/6037892, {"references":["Schotte M. & Heard R. W. 1991. Studies on the Crustacea of the Turks and Caicos Islands, British West Indies. II. Armadilloniscus steptus, n. sp. (Isopoda: Oniscidea: Scyphacidae) from Pine Cay. Gulf Research Reports 8 (3): 247 - 250. https: // doi. org / 10.18785 / grr. 0803.04","Paoletti M. G. & Stinner B. R. 1989. Two new terrestrial Isopoda (Oniscidea) from coralline cays of Venezuela's Caribbean coast. Proceedings of the Entomological Society of Washington 91 (1): 71 - 80. Available from https: // biostor. org / reference / 56403 [accessed 6 Jan. 2022].","Taiti S. & Ferrara F. 1989. New species and records of Armadilloniscus Uljanin 1875 (Crustacea Isopoda Oniscidea) from the coasts of the Indian and Pacific oceans. Tropical Zoology 2: 59 - 88. https: // doi. org / 10.1080 / 03946975.1989.10539427","Arcangeli A. 1957. Il genere Armadilloniscus Ulj. e gli Scyphacidae (crostacei isopodi terrestri). Atti della reale Accademia delle Scienze di Torino 91: 399 - 428.","Vandel A. 1962. Isopodes terrestres (Deuxieme Partie). Faune de France 66: 417 - 931.","Schmidt C. 2002. Contribution to the phylogenetic system of the Crinocheta (Crustacea, Isopoda). Part 1 (Olibrinidae to Scyphacidae s. str.). Mitteilungen aus dem Museum fur Naturkunde in Berlin, Zoologische Reihe 78: 275 - 352. https: // doi. org / 10.1002 / mmnz. 20020780207","Garthwaite R., Lawson R. & Taiti S. 1992. Morphological and genetical relationships among four species of Armadilloniscus Uljanin, 1875 (Isopoda: Oniscidea: Scyphacidae). Journal of Natural History 26: 327 - 338. https: // doi. org / 10.1080 / 00222939200770171"]}