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Cyphotrypa sp. Figs 3A–E, Table 2 Material. SibGIU 12/25 (sections). Description. Massive (?ramose) colony with poorly defined exozone.Autozooecia prismatic, containing straight, slightly curved diaphragms. Distance between diaphragms 0.10–0.42 mm. Autozooecial apertures polygonal, 4 spaced per 1 mm. Exilazooecia rare, small. Acanthostyles abundant, 0.02 mm in diameter, 3–5 surrounding each aperture. Autozooecial walls laminated, merged without visible autozooecial boundaries, 0.04 mm thick in exozone. Comparison. The present material is similar to C. olgae Tolokonnikova, 2010 from the lower Famennian of Gorny Altay (Tolokonnikova 2010), but differs in presence of diaphragms in each autozooecial chamber and in larger distance between them (0.10–0.42 mm vs. 0.10–0.27 mm in C. olgae), smaller exilazooecia (0.03–0.04 mm vs. 0.056 –0.098 mm in C. olgae) and acanthostyles (0.02 mm vs. 0.042 –0.056 mm in C. olgae). The present material differs from C. minuscula Volkova, 1974 (Volkova 1974) from the lower Famennian of Gorny Altay in smaller apertures (0.16–0.24 mm vs. 0.20–0.30 mm in C. minuscula) and exilazooecia (0.03–0.04 mm vs. 0.10 mm in C. minuscula), abundant acanthostyles vs. rare ones in C. minuscula. The available material is not sufficient to make a comprehensive comparison. Stratigraphical and geographical ranges. Ermaksy Formation, lower-middle Famennian (upper Devonian); Karatau, Zhankurgan outcrop, Kazakhstan., Published as part of Tolokonnikova, Zoya A. & Fedorov, Petr V., 2022, Late Devonian-early Carboniferous bryozoans from Zhankurgan (Greater Karatau, Kazakhstan) - taxonomy and palaeobiogeographical implications, pp. 252-264 in Zootaxa 5196 (2) on page 257, DOI: 10.11646/zootaxa.5196.2.6, http://zenodo.org/record/7224545, {"references":["Tolokonnikova, Z. A. (2010) New bryozoans from the Devonian of Gorny Altay (Russia). Paleontological Journal, 2 (44), 151 - 156. https: // doi. org / 10.1134 / S 0031030110020061","Volkova, K. M. (1974) Devonian bryozoans from Southeastern Altai. Moscow, Nauka, 181 pp. [in Russian]"]}

Three bryozoan species Anomalotoechus sp., Pseudobatostomella sp. and Cyphotrypa sp. are described from the upper part of the Ermaksy Formation (lower-middle Famennian) of the Zhankurgan outcrop (Greater Karatau). These descriptions are important, because it is the first finding of upper Devonian bryozoans in Southern Kazakhstan. Five more species including one new Primorella zhankurganica sp. n. were found in limestones of the lower part of Orgalysay Formation (Upper Tournaisian) in the same outcrop. The hitherto known bryozoans from the Tournaisian of Greater Karatau constitute the most important bryozoan fauna from the carbonate platform, located on southwestern passive margin of the Kazakhstan palaeocontinent. The studied fauna reveals connections between different areas of Palaeotethys and Palaeoasian oceans in late Devonian-early Carboniferous.

Pseudobatostomella sp. Figs 3H, I, Table 4 Material. SibGIU 12/27 (sections). Description. Thin branched colony 2.75–3.25 mm in diameter with 1.00– 1.25 mm wide endozones.Autozooecia long in endozones, bending at low angles in exozones. Autozooecial diaphragms complete, rare, located in the transition between endo- and exozones. Autozooecial walls laminated, 0.02 mm thick in endozone. Exilazooecia 0.10–0.15 mm wide, short, common. Comparison. The present material differs from P. majusculа Volkova, 1974 (Volkova 1974) from the lower Famennian of Gorny Altay in larger exilazooecia (0.10–0.15 mm vs. 0.06–0.10 mm in P. majusculа). The available material is not sufficient to make a comprehensive comparison. Stratigraphical and geographical ranges. Ermaksy Formation, lower-middle Famennian (upper Devonian); Karatau, Zhankurgan outcrop, Kazakhstan., Published as part of Tolokonnikova, Zoya A. & Fedorov, Petr V., 2022, Late Devonian-early Carboniferous bryozoans from Zhankurgan (Greater Karatau, Kazakhstan) - taxonomy and palaeobiogeographical implications, pp. 252-264 in Zootaxa 5196 (2) on page 259, DOI: 10.11646/zootaxa.5196.2.6, http://zenodo.org/record/7224545, {"references":["Volkova, K. M. (1974) Devonian bryozoans from Southeastern Altai. Moscow, Nauka, 181 pp. [in Russian]"]}

Anomalotoechus sp. Figs 3F, G, Table 3 Material. SibGIU 12/26 (sections). Description. Encrusting colony, 0.89–1.25 mm in thickness. The central hollow cylinder is 0.20x 0.24 mm in diameter. Autozooecia growing from 0.02 mm thick epitheca. Exozone indistinctly separated from endozone. Autozooecia having polygonal cross sections. Autozooecial apertures rounded-polygonal. Straight and inclined autozooecial diaphragms abundant; located at a distance of 0.04–0.10 mm (more often 0.08 mm). Exilazooecia rare, short. Acanthostyles having distinct cores and laminated sheaths. Autozooecial wall 0.01 mm thick in endozone and irregularly thickened in exozones. Maculae not observed. Comparison. The present material is similar to A. pervulgatus Lavrentjeva, 2001 from the lower Famennian of Transcaucasia (Lavrentjeva 2001), but differs in small acanthostyles (0.02 mm vs. 0.04–0.08 mm in A. pervulgatus). The present material differs from A. pojarkovi Modzalevskaya, 1968 from the lower Famennian of Western Talas (Southern Tien-Shan) in shape of colony (encrusting vs. branched in A. pojarkovi), smaller apertures (0.12–0.16 mm vs. 0.21–0.26 mm in A. pojarkovi), presence of diaphragms in endo- and exozone (Modzalevskaya & Orlovskij 1968). The available material is not sufficient to make a comprehensive comparison. Stratigraphical and geographical ranges. Ermaksy Formation, lower-middle Famennian (upper Devonian); Karatau, Zhankurgan outcrop, Kazakhstan., Published as part of Tolokonnikova, Zoya A. & Fedorov, Petr V., 2022, Late Devonian-early Carboniferous bryozoans from Zhankurgan (Greater Karatau, Kazakhstan) - taxonomy and palaeobiogeographical implications, pp. 252-264 in Zootaxa 5196 (2) on pages 257-258, DOI: 10.11646/zootaxa.5196.2.6, http://zenodo.org/record/7224545, {"references":["Lavrentjeva, V. D. (2001) New bryozoan species of the family Atactotoechidae from the Devonian of Transcaucasia. Paleontological Journal, 35 (2), 152 - 156.","Modzalevskaya, E. A. & Orlovskij, M. B. (1968) New species of Devonian and Silurian bryozoans from Middle Asia. New species of fossil plants and invertebrates, 1968, 47 - 54. [in Russian]"]}

Orbignyella multitabulata Coryell, 1921 (Fig. 6 E-G; Appendix 1) Orbignyella multitabulata Coryell, 1921: 284, pl. 5, figs 3-4. MATERIAL EXAMINED. — CEGH-UNC 27505 a-c, CEGH-UNC 27518 a-c. OCCURRENCE. — La Pola Formation, Upper Ordovician, Sandbian; La Pola creek section near Albardon village, San Juan Province, Argentine Precordillera, western Argentina. Pierce Limestone, Upper Ordovician (Sandbian); Tennessee, United States. DESCRIPTION Massive, subramose multilayered colonies. Secondary overgrowths common, 1-2 mm thick. Autozooecia bending gently from endozone, intersecting colony surface at right angles. Autozooecial apertures rounded to polygonal. Diaphragms in autozooecia common to abundant, straight to curved. Exilazooecia rare, polygonal in cross section, restricted to exozone. Acanthostyles common, moderately large, situated at junctions of autozooecial apertures. Autozooecial walls granular-prismatic, 0.005 -0.010 mm thick in endozone; irregularly thickened, finely laminated, displaying reverse V-structure in longitudinal section, 0.020 -0.033 mm thick in exozone. Maculae of macrozooecia 0.70-1.25 mm in diameter, spaced 1.9-2.0 mm from centre to centre. COMPARISON The present material is morphologically similar to Orbignyella multitabulata Coryell, 1921 from the Sandbian of United States. The only metric characteristic for O. multitabulata is the number of apertures per 2 mm given as 8-8.5 (Coryell 1921: 284). Interpolated, it gives apertural spacing of c. 0.23-0.25 mm what overlaps the range in the present species (0.13-0.22 mm). The present material differs from Orbignyella wetherbyi (Ulrich, 1890) from the Upper Ordovician of United States in smaller autozooecia (aperture width 0.11-0.19 mm vs 0.25-0.28 mm in O. wetherbyi)., Published as part of Ernst, Andrej & Carrera, Marcelo G., 2022, A cool-water bryozoan association from the La Pola Formation (Sandbian, Ordovician) of Argentine Precordillera, pp. 563-601 in Geodiversitas 44 (20) on page 572, DOI: 10.5252/geodiversitas2022v44a20, http://zenodo.org/record/6725916, {"references":["CORYELL H. N. 1921. - Bryozoan faunas of the Stone River Group of Central Tennessee. Proceedings of Indiana Academy of Sciences 1921: 261 - 340.","ULRICH E. O. 1890. - Palaeozoic Bryozoa: III. Report of the Geological Survey of Illinois 8: 283 - 688."]}

Dianulites rocklandensis Wilson, 1921 (Figs 5 F-G; 6A; Appendix 1) Dianulites rocklandensis Wilson, 1921: 47, pl. 2, figs 1-2. MATERIAL EXAMINED. — Single colony (three thin sections) CEGH-UNC 27503 a-c. OCCURRENCE. — La Pola Formation, Upper Ordovician, Sandbian; La Pola creek section near Albardon village, San Juan Province, Argentine Precordillera, western Argentina. Leray and Rockland formations, Upper Ordovician (Sandbian); Canada. DESCRIPTION Massive hemispherical colony, 8.5 mm thick in its central part and 20 mm wide at its base. Secondary overgrowth not observed. Exozone indistinct. Autozooecia long, prismatic, growing from epitheca. Autozooecial apertures polygonal. Diaphragms straight, rare to common in endozone, common in exozone. Autozooecial walls indistinctly granular, irregularly thickened, 0.015 -0.040 mm thick. Maculae not observed. COMPARISON The present material is similar to Dianulites rocklandensis Wilson, 1921 from the Ordovician (upper Sandbian) of Canada. This species, as described byWilson (1921), developed branched ramose colonies. However, Kang (2017) mentioned also hemispheric colonies of this species found at the type locality. Dianulites rocklandensis differs from D. microcellatus Astrova, 1945 from the Upper Ordovician of Urals in larger autozooecial apertures (0.24-0.36 mm vs 0.19-0.28 mm in D. microcellatus)., Published as part of Ernst, Andrej & Carrera, Marcelo G., 2022, A cool-water bryozoan association from the La Pola Formation (Sandbian, Ordovician) of Argentine Precordillera, pp. 563-601 in Geodiversitas 44 (20) on page 570, DOI: 10.5252/geodiversitas2022v44a20, http://zenodo.org/record/6725916, {"references":["WILSON A. E. 1921. - The range of certain Lower Ordovician faunas of the Ottawa Valley with descriptions of some new species. Canada Geological Survey Bulletin 33: 19 - 57. https: // doi. org / 10.4095 / 104998","KANG H. 2017. - Stratigraphy, Sedimentology, and Diagenesis of Ordovician Outliers, Northern Ottawa - Bonnechere Graben, Central Ontario. Master Thesis. Carleton University, Ottawa, Ontario, 284 p.","ASTROVA G. G. 1945. - Lower Silurian Trepostomata of the Kozhin River [in Russian]. Ezhegodnik Vsesoyuznogo Paleontologicheskogo Obshchestva 12: 81 - 92."]}

Homotrypa vacua McKinney, 1971 (Fig. 7 D-I; Appendix 1) Homotrypa vacua McKinney, 1971: 238-241, pl. 50, figs 2-7. — Pushkin 1987: 186. MATERIAL EXAMINED. — CEGH-UNC 27507 c, CEGH-UNC 27525 a, CEGH-UNC 27529 a, CEGH-UNC 27530 a, b, d-g, CEGH-UNC 27533 d, CEGH-UNC 27541 a, CEGH-UNC 27545 a. OCCURRENCE. — Lower Chickamauga Group, Upper Ordovician, Sandbian; Alabama, United States. La Pola Formation, Upper Ordovician, Sandbian; La Pola creek section near Albardon village, San Juan Province, Argentine Precordillera, western Argentina. DESCRIPTION Ramose colonies, branch diameter 1.45-2.38 mm. Endozone 0.60-1.45 mm wide, exozone 0.25-0.75 mm wide, distinct. Secondary overgrowths not observed. Autozooecia long in the endozone, having larger diameters than in exozone, bending gently and intersecting branch surface at low angles. Autozooecial apertures rounded-polygonal. Autozooecial diaphragms rare to absent in the endozone, concentrated mostly in transitional region between endo- and exozone, common to abundant in outer exozone.Cystiphragms occurring throughout the exozone, occupying about the half of autozooecial diameter. Mesozooecia locally 3-5 surrounding each autozooecial aperture, otherwise rare; small, short, restricted to the outermost part of exozone, containing densely spaced diaphragms. Acanthostyles common, 2-5 surrounding each autozooecial aperture, moderately large, restricted to exozone. Autozooecial walls straight, displaying granular microstructure, 0.003 -0.005 mm thick in endozone; finely laminated with indistinct medial line, 0.023 -0.055 mm thick in exozone. Indistinct maculae consisting of macrozooecia. COMPARISON Homotrypa vacua McKinney, 1971 differs from H. mundula (Ulrich, 1893) in having smaller colonies (branch diameter 1.45-2.38 mm vs 3.6-4.0 in H. mundula, measurements from Karklins [1984: 134]). Homotrypa vacua differs from H. subramosa Ulrich, 1886 in having thin branched colonies instead subramose and encrusting ones as well as in smaller autozooecial apertures (average aperture width 0.11 mm vs 0.16 mm in H. subramosa)., Published as part of Ernst, Andrej & Carrera, Marcelo G., 2022, A cool-water bryozoan association from the La Pola Formation (Sandbian, Ordovician) of Argentine Precordillera, pp. 563-601 in Geodiversitas 44 (20) on page 574, DOI: 10.5252/geodiversitas2022v44a20, http://zenodo.org/record/6725916, {"references":["MCKINNEY F. K. 1971. - Trepostomatous Ectoprocta (Bryozoa) from the lower Chickamauga Group (Middle Ordovician), Wills Valley, Alabama. Bulletins of American Paleontology 60: 195 - 333. https: // www. biodiversitylibrary. org / page / 28721254","PUSHKIN V. I. 1987. - Bryozoans of suborder Halloporina from Belorussia and East European Platform, in ROPOT V. P. & PUSHKIN V. I. (eds), Ordovician of Belarus. Nauka i Technika, Minsk: 145 - 232.","ULRICH E. O. 1893. - On Lower Silurian Bryozoa of Minnesota. The Geological and Natural History Survey of Minnesota, final report 3: 96 - 332. https: // doi. org / 10.5962 / bhl. title. 56338","KARKLINS O. L. 1984. - Trepostome and cystoporate bryozoans from the Lexington Limestone and the Clays Ferry Formation (Middle and Upper Ordovician) of Kentucky, in POJETA J. Jr. (ed.), Contributions to the Ordovician paleontology of Kentucky and nearby states. United States Geological Survey, Professional Paper 1066 I: I 1 - I 102. https: // doi. org / 10.3133 / pp 1066 I","ULRICH E. O. 1886. - Descriptions of new Silurian and Devonian fossils. Bryozoa. Contributions to American Palaeontology 1: 8 - 33. https: // doi. org / 10.5962 / bhl. title. 140012"]}

Diplotrypa sp. A (Fig. 9G; 10 A-D; Appendix 1) MATERIAL EXAMINED. — Single colony (two thin sections) CEGH-UNC 27543a, b. OCCURRENCE. — La Pola Formation, Upper Ordovician, Sandbian; La Pola creek section near Albardon village, San Juan Province, Argentine Precordillera, western Argentina. DESCRIPTION Massive colony, 6.2-6.4 mm in thickness, with indistinct endozone. Autozooecia growing from epitheca at high angles. Autozooecia in outer exozone often developing from mesozooecia. Autozooecial diaphragms common to rare, planar, curved proximally, irregularly spaced in autozooecia. Mesozooecia common, up to 3 surrounding each autozooecial aperture, bearing abundant straight diaphragms. Autozooecial walls fine fibrous microstructure, 0.005 -0.010 mm thick in endozone and 0.02-0.03 mm thick in exozone. COMPARISON Diplotrypa sp. A is similar to Diplotrypa catenulata Coryell, 1921 from the Sandbian of United States. Coryell [1921: 296] gave only the spacing of 4-4.5 apertures per 2 mm (distance from centre to centre 0.44-0.50 mm), which is more than double as large as the aperture spacing of the present material (average distance from centre to centre 0.22 mm). Size of apertures for Diplotrypa catenulata recorded by Astrova (1965: 185-186) from the Middle Ordovician of Arctic, was given as 0.42-0.75 mm (with macrozooecia of 0.9-1.0 mm width). Diplotrypa sp. A is similar to D. pusilla Astrova, 1965 from the Lower Ordovician of the Russian Arctic, but differs from it by less abundant diaphragms in autozooecia., Published as part of Ernst, Andrej & Carrera, Marcelo G., 2022, A cool-water bryozoan association from the La Pola Formation (Sandbian, Ordovician) of Argentine Precordillera, pp. 563-601 in Geodiversitas 44 (20) on page 578, DOI: 10.5252/geodiversitas2022v44a20, http://zenodo.org/record/6725916, {"references":["CORYELL H. N. 1921. - Bryozoan faunas of the Stone River Group of Central Tennessee. Proceedings of Indiana Academy of Sciences 1921: 261 - 340.","ASTROVA G. G. 1965. - Morphology, history of development and system of the Ordovician and Silurian Bryozoa. Trudy Paleontologicheskogo Instituta Akademii Nauk SSSR 106: 1 - 432 [in Russian]."]}

Genus Albardonia n. gen. urn:lsid:zoobank.org:act: CF51A226-FD7D-4583-B045-A26177B30FC0 TYPE SPECIES. — Albardonia bifoliata n. sp., by present designation. ETYMOLOGY. — The new genus is named after Albardon village near which this genus was found. DIAGNOSIS. — Ramose colony consisting of bifoliate fronds; encrusting sheets and secondary overgrowth common; autozooecia budding from mesotheca or epitheca, having angular shape of transverse section in endozone; autozooecial apertures angular with rounded corners; basal diaphragms common both in endozone and exozone, straight, or inclined; mesozooecia abundant, containing densely spaced diaphragms; aktinotostyles moderately large, abundant, originating in exozone, autozooecial walls laminated, integrated with dark median lining, showing reversal V-shaped lamination, without cingulum; maculae lacking. OCCURRENCE. — La Pola Formation, Upper Ordovician, Sandbian; Argentine Precordillera, western Argentina. COMPARISON The new genus is characterized by abundant mesozooecia and aktinotostyles. It is similar to the genus Leioclema Ulrich, 1882 in having abundant and large mesozooecia, but differs in the presence of aktinotostyles instead of acanthostyles in the latter genus., Published as part of Ernst, Andrej & Carrera, Marcelo G., 2022, A cool-water bryozoan association from the La Pola Formation (Sandbian, Ordovician) of Argentine Precordillera, pp. 563-601 in Geodiversitas 44 (20) on page 576, DOI: 10.5252/geodiversitas2022v44a20, http://zenodo.org/record/6725916, {"references":["ULRICH E. O. 1882. - American Palaeozoic Bryozoa. The Journal of the Cincinnati Society of Natural History 5: 121 - 175, 233 - 257."]}

Argentinopora robusta n. gen., n. sp. (Figs 11 F-G; 12A-F; Appendix 1) urn:lsid:zoobank.org:act: 51DCA703-84A9-4080-8A78-D507D5D6D0FC HOLOTYPE. — CEGH-UNC 27511 a-d (one colony, four thin sections). PARATYPES. — CEGH-UNC 27514 a-c, CEGH-UNC 27515 a, CEGH-UNC 27520 a, b. TYPE LOCALITY. — La Pola creek section near Albardon village, San Juan Province, Argentine Precordillera, western Argentina. TYPE HORIZON. — La Pola Formation, Upper Ordovician, Sandbian. DIAGNOSIS. — As for genus. ETYMOLOGY. — The species name refers to its thick walls and large mesozooecia and acanthostyles (from Latin “ robustus ” – robust, stable, strong). DESCRIPTION Ramose colony with multiple secondary overgrowths, 3.5- 11.0 mm in diameter, with 0.72.– 1.44 mm wide exozones. Secondary overgrowths 1.05-2.00 mm thick. Autozooecial apertures rounded-polygonal, often petaloid.Basal diaphragms abundant, straight or inclined throughout autozooecia.Cystiph - ragms common in exozones. Mesozooecia abundant, 3-7 surrounding each autozooecial aperture, originating in the early exozone, large, cystose, containing thick diaphragms. Acanthostyles large, abundant, 5-8 surrounding each autozooecial aperture, originating in endozone, having wide hyaline cores and narrow laminated sheaths, often indenting autozooecia and mesozooecia. Autozooecial walls laminated, 0.01-0.03 mm thick in endozones; thickly laminated, merged, without cingulum, 0.04-0.11 mm thick in exozones. Maculae not observed. COMPARISON As for genus.

Genus Argentinopora n. gen. urn:lsid:zoobank.org:act: 54274BD8-98CF-4779-A8CD-D5A6CA3CC4F8 TYPE SPECIES. — Argentinopora robusta n. sp., by present designation. DIAGNOSIS. — Ramose colonies with multiple secondary overgrowths; autozooecia tubular, shape of transverse section angular in endozone and rounded-angular in exozone; basal diaphragms common both in endozone and exozone; cystiphragms common in exozone;autozooecial apertures rounded-polygonal, often petaloid; mesozooecia abundant, 3-7 surrounding each autozooecial aperture, originating in the early exozone, large, cystose, containing thick diaphragms; acanthostyles large, abundant, 5-8 surrounding each autozooecial aperture, originating in endozone, having wide hyaline cores and narrow laminated sheaths, often indenting autozooecia and mesozooecia; exozonal walls thickly laminated, merged, without cingulum; maculae not observed. ETYMOLOGY. — The new genus is named after finding it in Argentina. OCCURRENCE. — La Pola Formation, Upper Ordovician, Sandbian; Argentine Precordillera, western Argentina. COMPARISON The new genus is characterized by large acanthostyles with narrow sheaths, cystose mesozooecia and presence of cystiphragms. It shows some similarities to the Family Ralfimartitidae Gorjunova, 2005. Gorjunova (2005) introduced the term “aulozooecia” for structures characteristic for Ralfimartitidae. The “aulozooecia” represent indeed large acanthostyles filled out by a hyaline skeletal material (see also Tavener-Smith 1969). The most similar genus in the Family Ralfimartitidae is Rozhnovites Gorjunova, 2005 which differs from Argentinopora in annulated arrangement of autozooecia and mesozooecia, as well as in absence of cystiphragms., Published as part of Ernst, Andrej & Carrera, Marcelo G., 2022, A cool-water bryozoan association from the La Pola Formation (Sandbian, Ordovician) of Argentine Precordillera, pp. 563-601 in Geodiversitas 44 (20) on pages 580-582, DOI: 10.5252/geodiversitas2022v44a20, http://zenodo.org/record/6725916, {"references":["LOEBLICH A. 1942. - Bryozoa from the Ordovician Bromide Formation, Oklahoma. Journal of Paleontology 16: 413 - 436. https: // www. jstor. org / stable / 1298838","GORJUNOVA R. V. 2005. - Ralfimartitidae, a new family of Paleozoic bryozoans of the order Trepostomida. Paleontologicheskii Zhurnal 2: 51 - 63 [in Russian]."]}

Homotrypa subramosa Ulrich, 1886 (Fig. 7 A-C; Appendix 1) Homotrypa subramosa Ulrich, 1886: 81; 1893: 239-340, pl. 19, figs 21- 28. — Bassler 1911: 187-189, text-figs 99-100. — Bork & Perry 1968: 1053-1055, pl. 136, figs 1-3. — McKinney 1971: 234-237, pl. 49, fig. 4-8, pl. 50, fig. 1. — Marintsch 1998: 53-55, pl. 9, figs 1-5. Homotrypa insignis Ulrich, 1886: 82. MATERIAL EXAMINED. — CEGH-UNC 27516 a, d, CEGH-UNC 27519 a, b, CEGH-UNC 27530 a-g, CEGH-UNC 27531 b, CEGH-UNC 27536 a, CEGH-UNC 27537 a, b, CEGH-UNC 27542 a, c, d, e, CEGH-UNC 27545b. DESCRIPTION Irregularly branched, submassive or encrusting colony. Branches and fronds 5.6-9.4 mm thick, encrusting sheets 0.7-1.6 mm thick. Exozones indistinct. Autozooecia prismatic, growing for long distance along branch axis and bending gently to the colony surface. Autozooecial diaphragms common, thin, planar. Autozooecial apertures polygonal in tangential section. Cystiphragms abundant, densely spaced, constricting middle part of zooecia, about a half of their diameter. Mesozooecia few, polygonal. Autozooecial walls granular, 0.003 -0.005 mm thick in endozone; amalgamated, 0.02-0.03 mm thick in exozone. Acanthostyles few, small to moderate in size. Maculae consisting of macrozooecia, 0.80-0.95 mm in diameter. COMPARISON Homotrypa subramosa Ulrich, 1886 differs from H. callosa Ulrich, 1893 in larger autozooecial apertures (average aperture width 0.16 mm vs 0.14 mm in H. callosa; measurements for H.callosa from Marintsch, 1998). Homotrypa subramosa Ulrich, 1886 differs from H.tuberculata Ulrich, 1893 in less abundant and smaller acanthostyles and in smaller autozooecial apertures (average aperture width 0.16 mm vs 0.20 mm in H. tuberculata; measurements for H. tuberculata from Marintsch 1998)., Published as part of Ernst, Andrej & Carrera, Marcelo G., 2022, A cool-water bryozoan association from the La Pola Formation (Sandbian, Ordovician) of Argentine Precordillera, pp. 563-601 in Geodiversitas 44 (20) on pages 572-574, DOI: 10.5252/geodiversitas2022v44a20, http://zenodo.org/record/6725916, {"references":["ULRICH E. O. 1886. - Descriptions of new Silurian and Devonian fossils. Bryozoa. Contributions to American Palaeontology 1: 8 - 33. https: // doi. org / 10.5962 / bhl. title. 140012","ULRICH E. O. 1893. - On Lower Silurian Bryozoa of Minnesota. The Geological and Natural History Survey of Minnesota, final report 3: 96 - 332. https: // doi. org / 10.5962 / bhl. title. 56338","BASSLER R. S. 1911. - The early Paleozoic Bryozoa of the Baltic Provinces. Bulletin of the Smithsonian Institution, United States National Museum 77: 1 - 382. https: // hdl. handle. net / 10088 / 30445","BORK K. B. & PERRY T. G. 1968. - Bryozoa (Ectoprocta) of Champlainian age (Middle Ordovician) from northwestern Illinois and adjacent parts of Iowa and Wisconsin, Part III, Homotrypa, Orbignyella, Prasopora, Monticulipora and Cyphotrypa. Journal of Paleontology 42: 1042 - 1065. https: // www. jstor. org / stable / 1302188","MCKINNEY F. K. 1971. - Trepostomatous Ectoprocta (Bryozoa) from the lower Chickamauga Group (Middle Ordovician), Wills Valley, Alabama. Bulletins of American Paleontology 60: 195 - 333. https: // www. biodiversitylibrary. org / page / 28721254","MARINTSCH E. J. 1998. - Systematic paleontology, biostratigraphy, and paleoecology of Middle Ordovician Bryozoa (Trepostomata) from the Hermitage Formation of East-Central Tennessee. Bulletins of American Paleontology 112: 1 - 121. https: // www. biodiversitylibrary. org / page / 10673542","WILSON A. E. 1921. - The range of certain Lower Ordovician faunas of the Ottawa Valley with descriptions of some new species. Canada Geological Survey Bulletin 33: 19 - 57. https: // doi. org / 10.4095 / 104998","CORYELL H. N. 1921. - Bryozoan faunas of the Stone River Group of Central Tennessee. Proceedings of Indiana Academy of Sciences 1921: 261 - 340."]}

Genus Nicholsonella Ulrich, 1890 TYPE SPECIES. — Nicholsonella ponderosa Ulrich, 1890 by original designation,Trentonian, Sandbian, Upper Ordovician, North America. DIAGNOSIS. — Encrusting, frondose, ramose, less commonly massive colonies. Apertures rounded and irregularly petaloid. Walls structureless, very thin, irregularly thickened in different part of colonies. Diaphragms usually abundant in whole colony, more rarely – only in the exozone. Mesozooecia abundant, containing frequent diaphragms, sometimes beaded, irregularly closed by calcitic material on the colony surface. Acanthostyles usually small, abundant, short, restricted to the outermost exozone. OCCURRENCE. — Lower Ordovician to Lower Silurian of North America and Siberia. COMPARISON Nicholsonella Ulrich, 1890 and some other genera (e.g. Dianulites) are unique among trepostome bryozoans by their re-crystallized walls which suggest a diagenetically unstable aragonitic (McKinney 1971) or high Mg calcite composition (Taylor & Wilson 1999)., Published as part of Ernst, Andrej & Carrera, Marcelo G., 2022, A cool-water bryozoan association from the La Pola Formation (Sandbian, Ordovician) of Argentine Precordillera, pp. 563-601 in Geodiversitas 44 (20) on page 582, DOI: 10.5252/geodiversitas2022v44a20, http://zenodo.org/record/6725916, {"references":["ULRICH E. O. 1890. - Palaeozoic Bryozoa: III. Report of the Geological Survey of Illinois 8: 283 - 688.","MCKINNEY F. K. 1971. - Trepostomatous Ectoprocta (Bryozoa) from the lower Chickamauga Group (Middle Ordovician), Wills Valley, Alabama. Bulletins of American Paleontology 60: 195 - 333. https: // www. biodiversitylibrary. org / page / 28721254","TAYLOR P. D. & WILSON M. A. 1999. - Dianulites Eichwald, 1829: an unusual Ordovician bryozoan with a high- magnesium calcite skeleton. Journal of Paleontology 73: 38 - 48. https: // doi. org / 10.1017 / S 0022336000027529"]}

Monticulipora aff. mammulata d’Orbigny, 1850 (Fig. 6 B-D; Appendix 1) For full synonymy of the species Monticulipora mammulata d’Orbigny, 1850, see in Brown & Daly (1985: 68). MATERIAL EXAMINED. — Single colony (three thin sections) CEGH-UNC 27516 a, b, d. OCCURRENCE. — La Pola Formation, Upper Ordovician, Sandbian; La Pola creek section near Albardon village, San Juan Province, Argentine Precordillera, western Argentina. DESCRIPTION Submassive colony, multilayered, weakly differentiated into exo- and endozone. Autozooecia prismatic, bearing abundant cystiphragms.Autozooecial diaphragms common to abundant, mainly inclined or cystoid. Autozooecial apertures polygonal. Mesozooecia common, polygonal in tangential section, bearing closely spaced diaphragms, locally beaded. Zooecial walls laminated, serrated, 0.01-0.02 mm thick. Acanthostyles rare, 0.030 -0.035 mm in diameter. Maculae not observed. COMPARISON The present material is similar to Monticulipora mammulata d’Orbigny, 1850 from the Upper Ordovician of North America. However, the typical monticules consisting of abundant mesozooecia, were not found in the present material. Monticulipora parallela McKinney, 1971 from the Middle Ordovician of United States (Alabama) differs from the present material in encrusting colony and larger autozooecial apertures (average aperture width 0.20 mm vs 0.18 mm in the present material)., Published as part of Ernst, Andrej & Carrera, Marcelo G., 2022, A cool-water bryozoan association from the La Pola Formation (Sandbian, Ordovician) of Argentine Precordillera, pp. 563-601 in Geodiversitas 44 (20) on pages 570-572, DOI: 10.5252/geodiversitas2022v44a20, http://zenodo.org/record/6725916, {"references":["BROWN G. D. JR & DALY E. J. 1985. - Trepostome Bryozoa from the Dillsboro Formation (Cincinnatian Series) of Southeastern Indiana. Indiana Geological Survey Special Report 33: 1 - 95. https: // doi. org / 10.5967 / be 7 p-p 337","WILSON A. E. 1921. - The range of certain Lower Ordovician faunas of the Ottawa Valley with descriptions of some new species. Canada Geological Survey Bulletin 33: 19 - 57. https: // doi. org / 10.4095 / 104998","MCKINNEY F. K. 1971. - Trepostomatous Ectoprocta (Bryozoa) from the lower Chickamauga Group (Middle Ordovician), Wills Valley, Alabama. Bulletins of American Paleontology 60: 195 - 333. https: // www. biodiversitylibrary. org / page / 28721254"]}

Genus Diplotrypa Nicholson, 1879 TYPE SPECIES. — Monticulipora (Diplotrypa) petropolitana Nicholson, 1879 by original designation (non Favosites petropolitanus Pander, 1830). Sweden; Middle Ordovician. DIAGNOSIS. — Massive, variably shaped colonies, exozone poorly developed. Budding pattern interzooecial. Zooecial arrangement disordered; zooecia gradually expand distally through early ontogeny and curve outward toward colony surface; zooecia characterised by ontogenetic progression of mesozooecia expanding into autozooecia. Mesozooecial stage of early zooecial ontogeny extended; after mesozooecial stage, diaphragms widely spaced in proximal ends and closely spaced in distal ends of mesozooecia; mesozooecia occasionally fuse to form autozooecia; mesozooecia commonly isolate autozooecia. Autozooecial apertures polygonal to rounded. Autozooecial walls commonly thin throughout colony and composed of finely crystalline microlaminae. Diaphragms thin, planar, concave, convex or cystoidal, variably spaced. Acanthostyles rare. Maculae usually consisting of macrozooecia. OCCURRENCE. — Lower Ordovician to Upper Silurian; North America, Asia, and Europe. COMPARISON Diplotrypa Nicholson, 1879 differs from other genera of the Family Halloporidae by its massive colony form and thin zooecial walls., Published as part of Ernst, Andrej & Carrera, Marcelo G., 2022, A cool-water bryozoan association from the La Pola Formation (Sandbian, Ordovician) of Argentine Precordillera, pp. 563-601 in Geodiversitas 44 (20) on page 578, DOI: 10.5252/geodiversitas2022v44a20, http://zenodo.org/record/6725916, {"references":["NICHOLSON H. A. 1879. - On the Structure and Affinities of the \" Tabulate Corals \" of the Paleozoic Period, with Critical Descriptions of Illustrative Species. 15 pls, 44 text-figs. William Blackwood and Sons, Edinburgh, 342 p. https: // doi. org / 10.5962 / bhl. title. 9117","PANDER C. H. 1830. - Beitrage zur Geognosie des russischen Reiches. K. Kray, St. Petersburg, 165 p. https: // doi. org / 10.3931 / e-rara- 39166"]}

Ernst, Andrej, Carrera, Marcelo G. (2022): A cool-water bryozoan association from the La Pola Formation (Sandbian, Ordovician) of Argentine Precordillera. Geodiversitas 44 (20): 563-601, DOI: 10.5252/geodiversitas2022v44a20

Tarphophragma macrostoma (Loeblich, 1942) (Figs 10 E-G; 11A; Appendix 1) Hallopora macrostoma Loeblich, 1942: 430, pl. 62, figs 12-14. Tarphophragma macrostoma – Key 1991: 207-209, figs 6.1-6.6. MATERIAL EXAMINED. — CEGH-UNC 27520 c, d, CEGH-UNC 27539 c, d, g, CEGH-UNC 27537 d, CEGH-UNC 27538 d. OCCURRENCE. — Bromide Formation, Upper Ordovician, lower Sandbian; Oklahoma, United States. La Pola Formation, Upper Ordovician, Sandbian; La Pola creek section near Albardon village, San Juan Province, Argentine Precordillera, western Argentina. DESCRIPTION Ramose colonies, branch diameter 3.0- 3.6 mm. Exozone distinct, 0.5-0.6 mm wide, endozone 2.0- 2.4 mm wide. Autozooecia long, growing parallel to branch axis for a long distance in endozone and bending sharply in exozone, having rounded-polygonal shape in transverse section in endozone. Autozooecial apertures oval to polygonal. Autozooecial diaphragms thin, planar, widely spaced in endozone; common to abundant in exozone, planar, rarely inclined, developed as extension of wall cortex. Mesozooecia arising in endozone, polygonal in transverse section, 2-6 surrounding each autozooecial aperture. Mesozooecial diaphragms planar, densely spaced. Acanthostyles absent. Autozooecial walls indistinctly laminated, 0.005 -0.008 mm thick in endozone; displaying distinct reverse V-shaped structure with dark autozooecial boundary, with weakly developed wall cortex continued in diaphragms, 0.04-0.12 mm thick in exozone. Maculae indistinct, consisting of macrozooecia. COMPARISON Tarphophragma macrostoma (Loeblich, 1942) differs from T. ovata (McKinney, 1971) from the Middle Ordovician of United States in larger autozooecial apertures (average aperture width 0.21 mm vs 0.17 mm in T. ovata; measurements for T. ovata from McKinney 1971), as well as in less abundant and smaller mesozooecia. Tarphophragma macrostoma differs from T. multitabulata (Ulrich, 1886) from the Middle Ordovician of United States in larger autozooecial apertures (average aperture width 0.21 mm vs 0.19 mm in T. multitabulata; measurements for T. multitabulata from Karklins 1984)., Published as part of Ernst, Andrej & Carrera, Marcelo G., 2022, A cool-water bryozoan association from the La Pola Formation (Sandbian, Ordovician) of Argentine Precordillera, pp. 563-601 in Geodiversitas 44 (20) on page 580, DOI: 10.5252/geodiversitas2022v44a20, http://zenodo.org/record/6725916, {"references":["LOEBLICH A. 1942. - Bryozoa from the Ordovician Bromide Formation, Oklahoma. Journal of Paleontology 16: 413 - 436. https: // www. jstor. org / stable / 1298838","KEY M. M. JR 1991. - The halloporid trepostome bryozoans from the Ordovician Simpson Group of Oklahoma. Journal of Paleontology 65: 200 - 212. https: // doi. org / 10.1017 / S 0022336000020436","MCKINNEY F. K. 1971. - Trepostomatous Ectoprocta (Bryozoa) from the lower Chickamauga Group (Middle Ordovician), Wills Valley, Alabama. Bulletins of American Paleontology 60: 195 - 333. https: // www. biodiversitylibrary. org / page / 28721254","ULRICH E. O. 1886. - Descriptions of new Silurian and Devonian fossils. Bryozoa. Contributions to American Palaeontology 1: 8 - 33. https: // doi. org / 10.5962 / bhl. title. 140012","KARKLINS O. L. 1984. - Trepostome and cystoporate bryozoans from the Lexington Limestone and the Clays Ferry Formation (Middle and Upper Ordovician) of Kentucky, in POJETA J. Jr. (ed.), Contributions to the Ordovician paleontology of Kentucky and nearby states. United States Geological Survey, Professional Paper 1066 I: I 1 - I 102. https: // doi. org / 10.3133 / pp 1066 I"]}

Albardonia bifoliata n. gen., n. sp. (Figs 8G, H; 9 A-F; Appendix 1) urn:lsid:zoobank.org:act: 8C45367C-9280-4982-A6C2-9E757EFFD061 HOLOTYPE. — CEGH-UNC 27538 c. PARATYPES. — CEGH-UNC 27507 a, b, CEGH-UNC 27520 a, b, d, CEGH-UNC 27528 a, CEGH-UNC 27529 a, CEGH-UNC 27530 a, b, e, f, g, CEGH-UNC 27532 a, CEGH-UNC 27533 a, b, CEGH-UNC 27537 b, CEGH-UNC 27538 a, b, CEGH-UNC 27539 b, CEGH-UNC 27541 a (in total, 19 thin sections of 10 colonies). TYPE LOCALITY. — La Pola creek section near Albardon village, San Juan Province, Argentine Precordillera, western Argentina. TYPE HORIZON. — La Pola Formation, Upper Ordovician, Sandbian. DIAGNOSIS. — As for genus. ETYMOLOGY. — The species name refers to the bifoliate colony shape of the new species. DESCRIPTION Bifoliate (frondose) colony starting from encrusting sheets. Fronds 0.70-1.30 mm in thickness. Encrusting sheets 0.66- 1.31 mm thick. Secondary overgrowths occurring. Autozooecia budding from mesotheca, having angular shape of transverse section in endozone. Autozooecial apertures angular with rounded corners. Mesotheca 0.01-0.02 mm in thickness. Basal diaphragms common both in endozone and exozone, straight, or inclined. Mesozooecia abundant, 3-8 surrounding each autozooecial aperture, containing densely spaced diaphragms. Aktinotostyles moderately large, abundant, 5-7 surrounding each autozooecial aperture, originating in the outer exozone. Autozooecial walls granular, 0.010 -0.015 mm thick, irregularly undulating in endozones; laminated, integrated with dark median lining, showing reversal V-shaped lamination, without cingulum, 0.02-0.05 mm thick in exozones. Maculae not observed. COMPARISON As for genus., Published as part of Ernst, Andrej & Carrera, Marcelo G., 2022, A cool-water bryozoan association from the La Pola Formation (Sandbian, Ordovician) of Argentine Precordillera, pp. 563-601 in Geodiversitas 44 (20) on pages 576-578, DOI: 10.5252/geodiversitas2022v44a20, http://zenodo.org/record/6725916

Nicholsonella spinigera n. sp. (Figs 12G, H; 13 A-F; Appendix 1) urn:lsid:zoobank.org:act: 29ECE5F7-47D8-4DF3-8375-4BFEF20F02DF HOLOTYPE. — CEGH-UNC 27528 b, c. PARATYPES. — CEGH-UNC 27507 a, b, CEGH-UNC 27512 a, b, CEGH-UNC 27522 a, b, CEGH-UNC 27530 e, CEGH-UNC 27533 d, CEGH-UNC 27538 a, c, CEGH-UNC 27539 g. TYPE LOCALITY. — La Pola creek section near Albardon village, San Juan Province, Argentine Precordillera, western Argentina. TYPE HORIZON. — La Pola Formation, Upper Ordovician, Sandbian. DIAGNOSIS. — Ramose colonies with distinct exozones; secondary overgrowths occurring; autozooecial apertures angular with rounded corners; basal diaphragms abundant both in endozone and exozone, straight, or inclined; mesozooecia few, containing densely spaced thick diaphragms; acanthostyles large, 4-8 surrounding each autozooecial aperture, originating in endozone, having wide hyaline cores and narrow laminated sheaths; exozonal autozooecial walls thickly laminated, merged, showing reversal U-shaped lamination, without cingulum; maculae lacking. ETYMOLOGY. — The species name refers to large and abundant acanthostyles of the new species (from Latin “ spiniger ” – spinose). DESCRIPTION Branched colony with secondary overgrowths, 3.5-11.0 mm in diameter, with 0.45-1.30 mm wide exozones. Secondary overgrowths 1.05-2.00 mm thick.Autozooecial apertures angular with rounded corners. Basal diaphragms abundant both in endozone and exozone, straight, or inclined. Mesozooecia few, containing densely spaced thick diaphragms. Acanthostyles large, abundant, 4-8 surrounding each autozooecial aperture, originating in endozone, having wide hyaline cores and narrow laminated sheaths, often indenting autozooecia and mesozooecia. Autozooecial walls granular, 0.005 -0.010 mm thick, irregularly undulating in endozones; thickly laminated, merged, showing reversal U-shaped lamination, without cingulum, 0.05-0.09 mm thick in exozones. Maculae not observed. COMPARISON Nicholsonella spinigera n. sp. is similar to N. irregularis Loeblich, 1942 from the Bromide Formation (Sandbian) of Oklahoma (United States), but differs from it in more abundant acanthostyles (4-8 acanthostyles per autozooecial aperture vs 5-6 in N. irregularis) and in more abundant autozooecial diaphragms. Nicholsonella spinigera n. sp. differs from N. pulchra Ulrich, 1893 from the Upper Ordovician of United States in less abundant mesozooecia., Published as part of Ernst, Andrej & Carrera, Marcelo G., 2022, A cool-water bryozoan association from the La Pola Formation (Sandbian, Ordovician) of Argentine Precordillera, pp. 563-601 in Geodiversitas 44 (20) on pages 582-586, DOI: 10.5252/geodiversitas2022v44a20, http://zenodo.org/record/6725916, {"references":["LOEBLICH A. 1942. - Bryozoa from the Ordovician Bromide Formation, Oklahoma. Journal of Paleontology 16: 413 - 436. https: // www. jstor. org / stable / 1298838","ROSS J. R. P. 1963 a. - Chazyan (Ordovician) leptotrypellid and atactotoechid Bryozoa. Palaeontology 5: 727 - 739.","ULRICH E. O. 1893. - On Lower Silurian Bryozoa of Minnesota. The Geological and Natural History Survey of Minnesota, final report 3: 96 - 332. https: // doi. org / 10.5962 / bhl. title. 56338"]}

Heterotrypa enodis n. sp. (Fig. 8 A-F; Appendix 1) urn:lsid:zoobank.org:act: C5482952-9218-4600-BDBE-C77CE258D9DD HOLOTYPE. — CEGH-UNC 27504 a-c (one colony, three thin sections). PARATYPES. — CEGH-UNC 27519 a, b (two thin sections of one colony), 27542 a, c-e (four thin sections of one colony). TYPE LOCALITY. — La Pola creek section near Albardon village, San Juan Province, Argentine Precordillera, western Argentina. TYPE HORIZON. — La Pola Formation, Upper Ordovician, Sandbian. ETYMOLOGY. — The species name refers to small acanthostyles of the new species (from Latin “ enodis ” – smooth). DIAGNOSIS. — Ramose colonies with distinct exozones; autozooecia with rounded-polygonal apertures; autozooecial diaphragms rare in endozone, common to abundant in exozone, developed as extension of wall cortex; mesozooecia moderately large, 1-4 surrounding each autozooecial aperture; acanthostyles small, 1-3 surrounding each autozooecial aperture; endozonal styles absent; autozooecial walls with distinct reverse V-shaped lamination with dark autozooecial border and weakly developed wall cortex continued in diaphragms; maculae consisting of macrozooecia. DESCRIPTION Ramose colonies, branch diameter 3.6 to 3.9 mm. Exozone distinct, 0.68 to 0.78 mm wide, endozone 2.24 to 2.34 mm wide. Autozooecia long, growing parallel to branch axis for a long distance in endozone, in exozone bending sharply and intersecting branch surface at angles of 66 to 74°, having rounded-polygonal shape in transverse section in endozone. Autozooecial apertures oval to polygonal. Autozooecial diaphragms thin, planar, widely spaced in endozone; common to abundant in exozone, planar, rarely inclined, developed as extension of wall cortex. Mesozooecia arising in endozone, polygonal in transverse section, few to common, 1-4 surrounding each autozooecial aperture. Mesozooecial diaphragms planar, densely spaced. Acanthostyles small, having laminated sheaths and indistinct hyaline cores, relatively abundant, 1-3 surrounding each autozooecial aperture. Endozonal styles absent. Autozooecial walls indistinctly laminated, 0.005 to 0.010 mm thick in endozone; displaying distinct reverse V-shaped structure with dark autozooecial border, with weakly developed wall cortex continued in diaphragms, 0.033 to 0.075 mm thick in exozone. Maculae consisting of macrozooecia, 1.25- 1.35 mm in diameter. COMPARISON The present species shows similarities to the genus Heterotrypa Nicholson, 1879 in the wall microstructure, abundant diaphragms arising from the wall cortex, presence of mesozooecia and acanthostyles. Heterotrypa enodis n. sp. differs from H. trentonensis (Ulrich, 1883) from the Upper Ordovician of North America in absence of endozonal styles, smaller exozonal acanthostyles (average acanthostyle diameter 0.03 mm vs 0.05 mm in H. trentonensis; measurements from Karklins 1984), and more abundant mesozooecia. Heterotrypa enodis differs from H. subtrentonensis Marintsch, 1998 from the Upper Ordovician of North America in absence of endozonal styles, less abundant exozonal acanthostyles (1-3 per aperture vs 3-5 in H. subtrentonensis), and more abundant mesozooecia., Published as part of Ernst, Andrej & Carrera, Marcelo G., 2022, A cool-water bryozoan association from the La Pola Formation (Sandbian, Ordovician) of Argentine Precordillera, pp. 563-601 in Geodiversitas 44 (20) on pages 574-576, DOI: 10.5252/geodiversitas2022v44a20, http://zenodo.org/record/6725916, {"references":["ULRICH E. O. 1886. - Descriptions of new Silurian and Devonian fossils. Bryozoa. Contributions to American Palaeontology 1: 8 - 33. https: // doi. org / 10.5962 / bhl. title. 140012","MCKINNEY F. K. 1971. - Trepostomatous Ectoprocta (Bryozoa) from the lower Chickamauga Group (Middle Ordovician), Wills Valley, Alabama. Bulletins of American Paleontology 60: 195 - 333. https: // www. biodiversitylibrary. org / page / 28721254","NICHOLSON H. A. 1879. - On the Structure and Affinities of the \" Tabulate Corals \" of the Paleozoic Period, with Critical Descriptions of Illustrative Species. 15 pls, 44 text-figs. William Blackwood and Sons, Edinburgh, 342 p. https: // doi. org / 10.5962 / bhl. title. 9117","ULRICH E. O. 1883. - American Palaeozoic Bryozoa. The Journal of the Cincinnati Society of Natural History 6: 245 - 279.","KARKLINS O. L. 1984. - Trepostome and cystoporate bryozoans from the Lexington Limestone and the Clays Ferry Formation (Middle and Upper Ordovician) of Kentucky, in POJETA J. Jr. (ed.), Contributions to the Ordovician paleontology of Kentucky and nearby states. United States Geological Survey, Professional Paper 1066 I: I 1 - I 102. https: // doi. org / 10.3133 / pp 1066 I","MARINTSCH E. J. 1998. - Systematic paleontology, biostratigraphy, and paleoecology of Middle Ordovician Bryozoa (Trepostomata) from the Hermitage Formation of East-Central Tennessee. Bulletins of American Paleontology 112: 1 - 121. https: // www. biodiversitylibrary. org / page / 10673542"]}

Genus Heterotrypa Nicholson, 1879 TYPE SPECIES. — Monticulipora frondosa d’Orbigny, 1850 by subsequent designation (Utgaard & Boardman 1965). Cincinnatian, Upper Ordovician; North America. DIAGNOSIS. — Colonies frondose, ramose or less commonly encrusting. Autozooecial walls can be extremely variable in thickness. Zooecial boundary is a conspicuous dark line in inner exozones and in a broad zone of abutting laminae or completely obscured in outer exozones. Walls generally are amalgamate in appearance. Diaphragms are generally few in endozones, but are moderately abundant in some species. In exozones, diaphragms are closely and regularly spaced, thin, planar and perpendicular to the zooecial walls. Intermonticular mesozooecia range from abundant and regularly arranged to scattered or absent. Mesozooecia develop commonly moniliform chambers at proximal ends and tend to become smaller or are terminated distally within exozones. Diaphragms in mesozooecia noticeably thicker and more closely spaced than zooecial diaphragms. Acanthostyles are at least two kinds within the genus, regular acanthostyles limited to exozone, and the endacanthostyles originating in both endozone and exozone. Endacanthostyles occur in all species. Monticules generally have a central cluster of a few mesozooecia than those in the intermonticular area. OCCURRENCE. — Middle Ordovician-?Devonian; worldwide. COMPARISON The genus Heterotrypa Nicholson, 1879 differs from the genus Leioclema Ulrich, 1882 by abundant diaphragms in autozooecia, more angular apertural shape and two kinds of acanthostyles., Published as part of Ernst, Andrej & Carrera, Marcelo G., 2022, A cool-water bryozoan association from the La Pola Formation (Sandbian, Ordovician) of Argentine Precordillera, pp. 563-601 in Geodiversitas 44 (20) on page 574, DOI: 10.5252/geodiversitas2022v44a20, http://zenodo.org/record/6725916, {"references":["NICHOLSON H. A. 1879. - On the Structure and Affinities of the \" Tabulate Corals \" of the Paleozoic Period, with Critical Descriptions of Illustrative Species. 15 pls, 44 text-figs. William Blackwood and Sons, Edinburgh, 342 p. https: // doi. org / 10.5962 / bhl. title. 9117","UTGAARD J. & BOARDMAN R. S. 1965. - Heterotrypa Nicholson, 1879, and Peronopora Nicholson, 1881 (Bryozoa, Trepostomata): proposed designation of a type species in conformity with generally accepted usage. Bulletin of Zoological Nomenclature 22: 112 - 118. https: // www. biodiversitylibrary. org / page / 12223512","ULRICH E. O. 1882. - American Palaeozoic Bryozoa. The Journal of the Cincinnati Society of Natural History 5: 121 - 175, 233 - 257."]}

Jordanopora heroensis Ross, 1963 (Fig. 11 B-E; Appendix 1) Jordanopora heroensis Ross, 1963a: 732, pl. 105, figs 1-8, pl. 106, figs 1-4, 6, 7. MATERIAL EXAMINED. — Single colony (three thin sections) CEGH-UNC 27530 e, f, g. OCCURRENCE. — Chazy Series (Darriwilian-Sandbian) of New York, United States. La Pola Formation, Upper Ordovician, Sandbian; La Pola creek section near Albardon village, San Juan Province, Argentine Precordillera, western Argentina. DESCRIPTION Colony form uncertain in the present material,possibly branched (bifoliate?) with secondary overgrowth. Exozone distinct, 0.45 mm wide. Autozooecia prismatic, autozooecial apertures rounded-polygonal. Diaphragms thin, planar, widely spaced in exozones.Autozooecial walls 0.010 -0.015 mm thick, laminated, irregularly undulating in endozones; thickly laminated, without cingulum, broadly and irregularly serrated, 0.045 -0.110 mm thick in exozones. Tubules in exozonal walls abundant, surrounding apertures in one row, hyaline, oriented parallel to autozooecial growth, 0.02-0.05 mm in diameter.Mesozooecia rare to common, occasionally 1-3 surrounding autozooecial apertures,originating in endozone, containing thick diaphragms. Styles absent. Maculae not observed in the present material. COMPARISON The present material fits to the species Jordanopora heroensis Ross, 1963 in its morphology and dimensions., Published as part of Ernst, Andrej & Carrera, Marcelo G., 2022, A cool-water bryozoan association from the La Pola Formation (Sandbian, Ordovician) of Argentine Precordillera, pp. 563-601 in Geodiversitas 44 (20) on page 580, DOI: 10.5252/geodiversitas2022v44a20, http://zenodo.org/record/6725916, {"references":["ROSS J. R. P. 1963 a. - Chazyan (Ordovician) leptotrypellid and atactotoechid Bryozoa. Palaeontology 5: 727 - 739."]}

Argentinopora robusta n. gen., n. sp. (Figs 11 F-G; 12A-F; Appendix 1) urn:lsid:zoobank.org:act: 51DCA703-84A9-4080-8A78-D507D5D6D0FC HOLOTYPE. — CEGH-UNC 27511 a-d (one colony, four thin sections). PARATYPES. — CEGH-UNC 27514 a-c, CEGH-UNC 27515 a, CEGH-UNC 27520 a, b. TYPE LOCALITY. — La Pola creek section near Albardon village, San Juan Province, Argentine Precordillera, western Argentina. TYPE HORIZON. — La Pola Formation, Upper Ordovician, Sandbian. DIAGNOSIS. — As for genus. ETYMOLOGY. — The species name refers to its thick walls and large mesozooecia and acanthostyles (from Latin “ robustus ” – robust, stable, strong). DESCRIPTION Ramose colony with multiple secondary overgrowths, 3.5- 11.0 mm in diameter, with 0.72.– 1.44 mm wide exozones. Secondary overgrowths 1.05-2.00 mm thick. Autozooecial apertures rounded-polygonal, often petaloid.Basal diaphragms abundant, straight or inclined throughout autozooecia.Cystiph - ragms common in exozones. Mesozooecia abundant, 3-7 surrounding each autozooecial aperture, originating in the early exozone, large, cystose, containing thick diaphragms. Acanthostyles large, abundant, 5-8 surrounding each autozooecial aperture, originating in endozone, having wide hyaline cores and narrow laminated sheaths, often indenting autozooecia and mesozooecia. Autozooecial walls laminated, 0.01-0.03 mm thick in endozones; thickly laminated, merged, without cingulum, 0.04-0.11 mm thick in exozones. Maculae not observed. COMPARISON As for genus., Published as part of Ernst, Andrej & Carrera, Marcelo G., 2022, A cool-water bryozoan association from the La Pola Formation (Sandbian, Ordovician) of Argentine Precordillera, pp. 563-601 in Geodiversitas 44 (20) on page 582, DOI: 10.5252/geodiversitas2022v44a20, http://zenodo.org/record/6725916

Genus Homotrypa Ulrich, 1882 TYPE SPECIES. — Homotrypa curvata Ulrich, 1882 by original designation. Cincinnatian, Upper Ordovician; North America. DIAGNOSIS. — Ramose and frondose colonies, often flattened, sometimes encrusting and irregularly massive in initial stages. Autozooecia with polygonal, rounded or oval apertures. Walls slightly thickened in exozone, integrate, diagonally and longitudinally laminated. Cystiphragms only in exozone, diaphragms commonly in exozone. Mesozooecia from rare to abundant, sometimes clustering in maculae. Acanthostyles abundant, commonly small. OCCURRENCE. — Middle Ordovician to Lower Silurian; North America, Europe, Australia, Siberia. COMPARISON The genus Homotrypa Ulrich, 1882 differs from the genus Monticulipora d’Orbigny, 1850 by its branched erect colony instead of the encrusting or massive one in Monticulipora. Furthermore, cystiphragms in Homotrypa are usually concentrated in the inner exozone, whereas cystiphragms in Monticulipora occur throughout autozooecial chambers., Published as part of Ernst, Andrej & Carrera, Marcelo G., 2022, A cool-water bryozoan association from the La Pola Formation (Sandbian, Ordovician) of Argentine Precordillera, pp. 563-601 in Geodiversitas 44 (20) on page 572, DOI: 10.5252/geodiversitas2022v44a20, http://zenodo.org/record/6725916, {"references":["ULRICH E. O. 1882. - American Palaeozoic Bryozoa. The Journal of the Cincinnati Society of Natural History 5: 121 - 175, 233 - 257."]}

Heterotrypa enodis n. sp. (Fig. 8 A-F; Appendix 1) urn:lsid:zoobank.org:act: C5482952-9218-4600-BDBE-C77CE258D9DD HOLOTYPE. — CEGH-UNC 27504 a-c (one colony, three thin sections). PARATYPES. — CEGH-UNC 27519 a, b (two thin sections of one colony), 27542 a, c-e (four thin sections of one colony). TYPE LOCALITY. — La Pola creek section near Albardon village, San Juan Province, Argentine Precordillera, western Argentina. TYPE HORIZON. — La Pola Formation, Upper Ordovician, Sandbian. ETYMOLOGY. — The species name refers to small acanthostyles of the new species (from Latin “ enodis ” – smooth). DIAGNOSIS. — Ramose colonies with distinct exozones; autozooecia with rounded-polygonal apertures; autozooecial diaphragms rare in endozone, common to abundant in exozone, developed as extension of wall cortex; mesozooecia moderately large, 1-4 surrounding each autozooecial aperture; acanthostyles small, 1-3 surrounding each autozooecial aperture; endozonal styles absent; autozooecial walls with distinct reverse V-shaped lamination with dark autozooecial border and weakly developed wall cortex continued in diaphragms; maculae consisting of macrozooecia. DESCRIPTION Ramose colonies, branch diameter 3.6 to 3.9 mm. Exozone distinct, 0.68 to 0.78 mm wide, endozone 2.24 to 2.34 mm wide. Autozooecia long, growing parallel to branch axis for a long distance in endozone, in exozone bending sharply and intersecting branch surface at angles of 66 to 74°, having rounded-polygonal shape in transverse section in endozone. Autozooecial apertures oval to polygonal. Autozooecial diaphragms thin, planar, widely spaced in endozone; common to abundant in exozone, planar, rarely inclined, developed as extension of wall cortex. Mesozooecia arising in endozone, polygonal in transverse section, few to common, 1-4 surrounding each autozooecial aperture. Mesozooecial diaphragms planar, densely spaced. Acanthostyles small, having laminated sheaths and indistinct hyaline cores, relatively abundant, 1-3 surrounding each autozooecial aperture. Endozonal styles absent. Autozooecial walls indistinctly laminated, 0.005 to 0.010 mm thick in endozone; displaying distinct reverse V-shaped structure with dark autozooecial border, with weakly developed wall cortex continued in diaphragms, 0.033 to 0.075 mm thick in exozone. Maculae consisting of macrozooecia, 1.25- 1.35 mm in diameter. COMPARISON The present species shows similarities to the genus Heterotrypa Nicholson, 1879 in the wall microstructure, abundant diaphragms arising from the wall cortex, presence of mesozooecia and acanthostyles. Heterotrypa enodis n. sp. differs from H. trentonensis (Ulrich, 1883) from the Upper Ordovician of North America in absence of endozonal styles, smaller exozonal acanthostyles (average acanthostyle diameter 0.03 mm vs 0.05 mm in H. trentonensis; measurements from Karklins 1984), and more abundant mesozooecia. Heterotrypa enodis differs from H. subtrentonensis Marintsch, 1998 from the Upper Ordovician of North America in absence of endozonal styles, less abundant exozonal acanthostyles (1-3 per aperture vs 3-5 in H. subtrentonensis), and more abundant mesozooecia.

Genus Jordanopora Ross, 1963 TYPE SPECIES. — Jordanopora heroensis Ross, 1963 by original designation. Ordovician, Llanvirn-Llandeilo; United States. DIAGNOSIS. — Colonies ramose or encrusting, with distinct exozones. Autozooecia prismatic, autozooecial apertures rounded-polygonal. Diaphragms generally lacking in endozones; thin, planar, closely to widely spaced or lacking in exozones. Autozooecial walls thin, laminated, irregularly undulating in endozones; thickly laminated, without cingulum, broadly and irregularly serrated in exozones. Tubules in exozonal walls abundant, thick, hyaline, generally oriented parallel to autozooecial growth. Mesozooecia rare to common, originating in endozone, containing thick diaphragms. Styles absent. Maculae low to flush, consisting of macrozooecia surround cluster of mesozooecia and massive extrazooidal skeleton near centres. OCCURRENCE. — Chazy Series (Darriwilian-Sandbian) of New York, United States, and Sandbian of Argentina. COMPARISON Jordanopora Ross, 1963 differs from other genera of the Family Trematoporidae in absence of acanthostyles and presence of tubules in zooecial walls., Published as part of Ernst, Andrej & Carrera, Marcelo G., 2022, A cool-water bryozoan association from the La Pola Formation (Sandbian, Ordovician) of Argentine Precordillera, pp. 563-601 in Geodiversitas 44 (20) on page 580, DOI: 10.5252/geodiversitas2022v44a20, http://zenodo.org/record/6725916, {"references":["ROSS J. R. P. 1963 a. - Chazyan (Ordovician) leptotrypellid and atactotoechid Bryozoa. Palaeontology 5: 727 - 739."]}

A bryozoan fauna containing 16 species is described from a set of small mud mounds occurring in the Lebanza Formation (Lower Devonian) of the Arauz area, NW Spain. One genus with one species is new: the cystoporate Petalosis clarus n. gen., n. sp. Eleven new species are described: three cystoporates Altshedata hispanica n. sp., A. gracilis n. sp. and Fistulipora arauzensis n. sp.; four trepostomes: Neotrematopora tenuis n. sp., Leioclema arauzensis n. sp., Eridotrypella validaeformis n. sp. and Leptotrypella armata n. sp.; two rhabdomesine cryptostomes Orthopora spinosa n. sp. and Vidronovella elegantula n. sp.; and two fenestrates Rectifenestella exiliformis n. sp. and Tectulipora tuberculata n. sp. Two species were identified at genus and family level respectively: Eridopora sp. and Rhomboporidae sp. indet. Two identified species show palaeobiogeographic relations to the Lower Devonian of Altai, Mongolia, and Tajikistan respectively: Leioclema multiacanthoporum Astrova in Astrova & Yaroshinskay...

Bryozoans are among the most abundant fossil groups in the cool water limestones that characterize the uppermost Katian (Upper Ordovician) of Southwestern Europe and North Africa. The Cystoid Limestone Formation, the representative unit of those widespread rocks in the Iberian Chains (NE Spain), yields one of the most diversified bryozoan faunas of the whole area. Herein the first systematic description of the trepostomate family Monticuliporidae is undertaken, this is one of the most diverse families within the associations collected. Four species of the genera Monticulipora and Prasopora are described, including the new species Monticulipora cystiphragmata n. sp. and Prasopora spjeldnaesi n. sp., and an indeterminate Monticuliporidae. Monticulipora and Prasopora have shown to be very similar genera, with several species that are difficult to distinguish between them. Their diagnoses include common features such as the wall microstructure, density of mesozoecia, and number of acanthostyles. Herein a new generic diagnosis of Monticulipora is proposed, putting special emphasis on the shape and distribution of cystiphragms. The species Monticulipora kolaluensis is identified for the first time out of Siberia. Although the record of these new taxa can indicate a certain isolation of the Iberian Chains region within the North Gondwana platform, the occurrence of Prasopora carnica shows a relation to the neighbouring Carnic Alps (Italy and Austria).

Genus Dyscritella Girty, 1911 TYPE SPECIES. — Dyscritella robusta Girty, 1911 by original designation. Lower Carboniferous; Arkansas, USA. OCCURRENCE. — Devonian to Triassic; worldwide. DIAGNOSIS. — Dendroid and encrusting colony with abundant acanthostyles and exilazooecia. Autozooecia parallel to longitudinal direction of the colony in endozone; gradually bending outward in exozone. Diaphragms in autozooecia lacking or very rare; lacking in exilazooecia. Exilazooecia circular to angular in cross section and separated from the autozooecia and from each other by thick walls. Two sizes of acanthostyles may be present. Zooecial walls thin in endozone, rapidly thickening in the exozone (modified after Ernst & Gorgij, 2013). COMPARISON Dyscritella Girty, 1911 generally lacks diaphragms which are commonly developed in the similar genus Dyscritellina Morozova in Dunaeva & Morozova, 1967., Published as part of Ernst, Andrej, Wyse Jackson, Patrick N. & Aretz, Markus, 2015, Bryozoan fauna from the Mississippian (Visean) of Roque Redonde (Montagne Noire, southern France), pp. 151-213 in Geodiversitas 37 (2) on pages 170-174, DOI: 10.5252/g2015n2a2, http://zenodo.org/record/4534998, {"references":["GIRTY G. H. 1911. - New genera and species of Carboniferous fossils from the Fayettville Shale of Arkansas. Annals of the New York Academy of Sciences 20: 189 - 238.","ERNST A. & GORGIJ M. N. 2013. - Lower Permian bryozoan faunas from Kalmard area, central Iran. Neues Jahrbuch fur Geologie und Palaontologie, Abhandlngen 268 (3): 275 - 324.","PHILLIPS J. 1836. - Illustrations of the Geology of Yorkshire. Pt. 2. The Mountain Limestone District. John Murray, London, xx + 253 p.","DELVOLVE J. J. & MCKINNEY F. K. 1983. - A Carboniferous bryozoan faunule from the Pyrenees. Senckenbergiana lethaia 64: 315 - 335.","NICHOLSON H. A. 1881. - On the Structure and Affinities of the Genus Monticulipora and its Subgenera, with Critical Descriptions of Illustrative Species. Willam Blackwood & Sons, Edinburgh & London, 240 p.","DUNAEVA N. N. & MOROZOVA I. P. 1967. - Osobennosti razvitiya i sistematicheskoe polozhenie nekotorykh pozdnepaleozojskikh trepostomat. Paleontologicheskiy Zhurnal 1967 (4): 86 - 94 [in Russian; English translation: Evolution and systematic position 9.5]."]}

A bryozoan fauna from the Mississippian (Visean) of Roque Redonde (Montagne Noire, southern France), contains 38 species (12 cystoporates, 2 trepostomes, 6 cryptostomes [rhabdomesines], and 18 fenestrates). Of them 9 are new: Fistulipora tolokonnikovae n. sp., Dybowskiella rotunda n. sp., Dybowskiella piriforme n. sp., Eridopora suarezi n. sp., Volgia deftera n. sp., Cystodictya gallensis n. sp., Megacanthopora enodata n. sp., Fabifenestella macrofenestrata n. sp., and Baculopora redondensis n. sp. Beside them, one new genus with a new species is established: Gorjunopora gallica n. gen., n. sp. Furthermore, 23 species characteristic of the Mississippian of Eurasia and North America were identified. The bryozoan species show various palaeobiogeographic connections. The closest connections are to the Mississippian (Visean) of the British Isles and the Russian Platform. Moreover, some connections to the Mississippian of Germany, Ukraine, Spain, Kuznets Basin, North America and Kazakhstan can be trac...

Tabulipora howsii (Nicholson, 1881) (Fig. 16 D-H; Appendix) For synonymy see Wyse Jackson (1996: 151, 152). MATERIAL. — SMF 21.849- SMF 21.854,TCD.60339, 60340, 60343. OCCURRENCE. — Carboniferous, Mississippian (Visean); Britain, Ireland, Germany (Velbert Anticline, Rhenish Massif (Wyse Jackson & Weber, 2005)), Roque Redonde (Montagne Noire, southern France). DESCRIPTION Encrusting to submassive colony, 2.7 mm thick. Autozooecia prismatic,having polygonal shape in transverse section,containing rare diaphragms and abundant ring septa. Ring septa concentrated in exozones, occupying about a half of the autozooecial chamber space. Acanthostyles moderately large, concentrated predominantly in maculae, having narrow hyaline cores and wide laminated sheaths. Exilazooecia rare, small, polygonal in transverse section, occurring predominantly in maculae. Endozonal walls granular, 0.015 -0.020 mm thick; exozonal walls monilae-shaped thickened, laminated, serrated, with distinct autozooecial boundaries, 0.05-0.07 mm thick. Maculae consisting of larger autozooecia present, 2.2-2.4 mm in diameter. COMPARISON Tabulipora howsii (Nicholson, 1881) differs from T. urii (Fleming, 1828) in larger autozooecial apertures (average aperture width 0.28 mm vs 0.19 mm in T. urii). Tabulipora howsii differs from T. stragula Karklins, 1986 from the Mississippian of Utah, USA in smaller autozooecial apertures (average aperture width 0.28 mm vs 0.39 mm in T. stragula)., Published as part of Ernst, Andrej, Wyse Jackson, Patrick N. & Aretz, Markus, 2015, Bryozoan fauna from the Mississippian (Visean) of Roque Redonde (Montagne Noire, southern France), pp. 151-213 in Geodiversitas 37 (2) on page 170, DOI: 10.5252/g2015n2a2, http://zenodo.org/record/4534998, {"references":["NICHOLSON H. A. 1881. - On the Structure and Affinities of the Genus Monticulipora and its Subgenera, with Critical Descriptions of Illustrative Species. Willam Blackwood & Sons, Edinburgh & London, 240 p.","WYSE JACKSON P. N. 1996. - Bryozoa from the Lower Carboniferous (Visean) of County Fermanagh, Ireland. Bulletin of the Natural History Museum (Geology) 52: 119 - 171.","WYSE JACKSON P. N. & WEBER H. M. 2005. - A bryozoan fauna from the Carboniferous (Mississippian, Late Visean) of the Velbert Anticline, Germany, in MOYANO G. H. I., CANCINO J. M. & WYSE JACKSON P. N. (eds), Bryozoan Studies 2004: Proceedings of the 13 th International Bryozoology Association conference, Conception, Chile. A. A. Balkema: Lisse, Abingdon, Exton (PA), Tokyo: 375 - 381.","FLEMING J. 1828. - A History of British Animals, Exhibiting their Descriptive Characters and Systematic Arrangement of the Genera and Species of Quadrupeds, Birds, Reptiles, Fishes, Mollusca, and Radiata of the United Kingdom. Bell & Bradfute, Edinburgh, 565 p.","KARKLINS O. L. 1986. - Chesterian (Late Mississippian) bryozoans from the Upper Chainman Shale and the lowermost Ely Limestone of western Utah. Paleontological Society Memoir 17: 1 - 48."]}

Genus Tabulipora Young, 1883 TYPE SPECIES. — Cellepora urii Fleming, 1828 by monotypy. Carboniferous; Scotland. DIAGNOSIS. — Ramose, encrusting, cylindrical or massive colonies. Autozooecia with basal diaphragms and ring septa. Autozooecial walls irregularly thickening with development of monilae. Exilazooecia rare. Acanthostyles of two sizes: small microacanthostyles and large macroacanthostyles. OCCURRENCE. — Carboniferous-Permian; worldwide. COMPARISON The genus Tabulipora Young, 1883 differs from the genera Stenopora Lonsdale, 1844 and Stenodiscus Crockford, 1945 by the development of ring septa., Published as part of Ernst, Andrej, Wyse Jackson, Patrick N. & Aretz, Markus, 2015, Bryozoan fauna from the Mississippian (Visean) of Roque Redonde (Montagne Noire, southern France), pp. 151-213 in Geodiversitas 37 (2) on page 170, DOI: 10.5252/g2015n2a2, http://zenodo.org/record/4534998, {"references":["YOUNG J. 1883. - On Ure's \" Millepore \". Tabulipora (Cellepora) Urii, Flem. Annals and Magazine of Natural History 12 (5): 154 - 158.","FLEMING J. 1828. - A History of British Animals, Exhibiting their Descriptive Characters and Systematic Arrangement of the Genera and Species of Quadrupeds, Birds, Reptiles, Fishes, Mollusca, and Radiata of the United Kingdom. Bell & Bradfute, Edinburgh, 565 p.","LONSDALE W. 1844. - Description of six species of corals from the Palaeozoic formation of Van Diemen's Land, in Geological observations on the Volcanic Islands visited during the voyage of H. M. S. \" Beagle \" (C. Darwin). Smith, Elder & Co, London: 161 - 169.","CROCKFORD J. 1945. - Stenoporids from the Permian of New South Wales and Tasmania. Proceedings of the New South Wales Linnean Society 70: 9 - 24."]}

Genus Leioclema Ulrich, 1882 TYPE SPECIES. — Callopora punctata Hall, 1858, by subsequent designation (Ulrich 1882). Lower Carboniferous; Iowa (USA). OCCURRENCE. — Lower Silurian to Upper Carboniferous; worldwide. DIAGNOSIS. — Encrusting,branched, less commonly massive colonies.Autozooecia with polygonal to rounded-polygonal, sometimes petaloid apertures. Autozooecial diaphragms rare. Mesozooecia abundant, with abundant diaphragms, often beaded. Acanthostyles abundant, commonly large. Autozooecial walls thin in endozone; laminated, regularly thickened in exozones (modified from Astrova 1978). COMPARISON Leioclema Ulrich, 1882 differs from Heterotrypa Nicholson, 1879 in having rare autozooecial diaphragms and abundant acanthostyles and mesozooecia, and from Stigmatella Ulrich& Bassler, 1904 in having abundant mesozooecia., Published as part of Ernst, Andrej, Fernández, Luis Pedro, Fernández-Martínez, Esperanza & Vera, Carmen, 2012, Description of a bryozoan fauna from mud mounds of the Lebanza Formation (Lower Devonian) in the Arauz area (Pisuerga-Carrión Province, Cantabrian Zone, NW Spain), pp. 693-738 in Geodiversitas 34 (4) on page 710, DOI: 10.5252/g2012n4a1, http://zenodo.org/record/5380890, {"references":["ULRICH E. O. 1882. - American Paleozoic Bryozoa. The Journal of the Cincinnati Society of Natural History 5: 121 - 175, 233 - 257."]}

Genus Prasopora Nicholson & Etherigde, 1877 TYPE SPECIES. — Prasopora grayae Nicholson & Etheridge, 1877; Upper Ordovician of Scotland., Published as part of Jiménez-Sánchez, Andrea, 2010, New Monticuliporidae (Bryozoa, Trepostomata) from the Cystoid Limestone Formation (Upper Ordovician) of the Iberian Chains (NE Spain), pp. 177-199 in Geodiversitas 32 (2) on page 188, DOI: 10.5252/g2010n2a1, http://zenodo.org/record/5375809, {"references":["NICHOLSON H. A. & ETHERIDGE R. 1877. - On Prasopora grayae, a new genus and species of Silurian corals. Annals and Magazine of Natural History series 4 (20): 388 - 392."]}

Genus Prasopora Nicholson & Etherigde, 1877 TYPE SPECIES. — Prasopora grayae Nicholson & Etheridge, 1877; Upper Ordovician of Scotland.

Genus Dyscritella Girty, 1911 TYPE SPECIES. — Dyscritella robusta Girty, 1911. Lower Carboniferous, Arkansas (USA)., Published as part of Ernst, Andrej, Senowbari-Daryan, Baba & Hamedani, Ali, 2006, Middle Permian Bryozoa from the Lakaftari area, northeast of Esfahan (central Iran), pp. 543-590 in Geodiversitas 28 (4) on page 550, DOI: 10.5281/zenodo.4665450

Ernst, Andrej, Senowbari-Daryan, Baba, Hamedani, Ali (2006): Middle Permian Bryozoa from the Lakaftari area, northeast of Esfahan (central Iran). Geodiversitas 28 (4): 543-590, DOI: http://doi.org/10.5281/zenodo.4665450, {"references":["BASSLER R. S. 1929. - The Permian Bryozoa of Timor. Palaontologie von Timor 16 (28): 37-90, 23 pls.","CROCKFORD J. 1944a. - Bryozoa from Wandagee and Nooncanbah Series of Western Australia. Journal of the Royal Society of Western Australia 28: 165-185.","CROCKFORD J. 1944b. - Bryozoa from the Permian of Western Australia. Part I. Cyclostomata and Cryptostomata from the north-west basin and Kimberly district. Proceedings of the Linnean Society of N. S. Wales 69 (3-4): 139-175.","ENGEL B. A. 1975. - A new ?bryozoan from the Carboniferous of Eastern Australia. Palaeontology 18 (3): 571-605.","FANTININ SESTINI F. N. 1965. - The geology of the upper Djadjerud and Lar valleys (north-Iran). II, Palaeontology. Bryozoans, brachiopods and molluscs from Ruteh limestone (Permian). Revista Italiana di Paleontologia 71 (1): 13-108.","GORJUNOVA R. V. 1975. - [Permian Bryozoans of the Pamir]. Transactions of the Paleontological Institute of the Academy of Sciences 148: 1-127 (in Russian).","GORJUNOVA R. V. & MOROZOVA I. P. 1979. - [Late Paleozoic Bryozoans of Mongolia]. Trudy Sovmestnoi Sovetsko-Mongolskoi Paleontologischeskoi Ekspeditzii 9: 1-134 (in Russian).","HAGEMAN S. J. 1991a. - Approaches to systematic and evolutionary studies of perplexing groups: an example using fenestrate Bryozoa. Journal of Paleontology 65 (4): 630-647.","HAGEMAN S. J. 1991b. - Discrete morphotaxa from a Mississippian fenestrate faunule: presence and implications, in BIGEY F. P. (ed.), Bryozoaires actuels et fossiles, Bryozoa living and fossil. Bulletin de la Societe des Sciences naturelles de l'Ouest de la France, Memoire HS 1: 147-150.","KIESSLING W., FLUGEL E. & GOLONKA J. 2003. - Patterns of Phanerozoic carbonate platform sedimentation. Lethaia 36 (3): 195-226.","KISELEVA A. B. 1982. - Late Permian Bryozoans of the South Primorye. Nauka, Moscow, 128 p.","LEVEN E. 1998. - Permian fusulinid assemblages and stratigraphy of the Transcaucasia. Rivista Italiana di Paleontologia e Stratigrafia 104 (3): 299-328.","MEEK F. B. 1872. - Report on the paleontology of Eastern Nebraska, with some remarks on the Carboniferous rocks of that district. Hayden's Final Report of the U.S. Geological Survey of Nebraska and Portions of Adjacent Territories. Government Printing Office, Washington D.C.. Part II. Paleontology: 81-264; Bryozoa: 141-158.","MOROZOVA I. P. 1965. - [Bryozoa], in RUZHENTSEV V. E. & SARYCHEVA T. G. (eds), [The evolution and changes in the marine organisms at the Paleozoic-Mesozoic boundary]. Trudy Paleontologicheskogo Instituta 108: 183-197 (in Russian).","MOROZOVA I. P. 1970a. - [Late Permian Bryozoa]. Trudy Paleontologicheskogo Instituta 122: 1-347 (in Russian).","MOROZOVA I. P. 1970b. - [Some new Late Permian bryozoans from southeastern Mongolia], in ASTROVA G. G. & CHUDINOVA I. I. (eds), [New Species of Paleozoic Bryozoans and Corals]. Nauka, Moscow: 69-75 (in Russian).","MOROZOVA I. P. 1974. - [Revision of the genus Fenestella]. Paleontologicheskii Zhurnal 2: 54-77 (in Russian).","MOROZOVA I. P. 1981. - [Late Paleozoic bryozoans of the Northwest of the USSR]. Trudy Paleontologicheskogo Instituta 188: 1-119 (in Russian).","MOROZOVA I. P. 2001. - [Bryozoans of the order Fenestrida]. Trudy Paleontologicheskogo Instituta 277: 1-176 (in Russian).","MOROZOVA I. P. & LISITSYN D. V. 1996. - [Revision of the genus Polypora M'Coy, 1844]. Paleontologicheskii Zhurnal 4: 38-47 (in Russian).","NIKIFOROVA A. I. 1933. - [Upper Paleozoic Bryozoans of Dshulfa Region]. Trudy Vsesoyusnogo Geologo-razvedochnogo obyedineniya 364: 1-44 (in Russian).","RIGBY J. K., SENOWBARI- DARYAN B. & HAMEDANI A. 2005. - First reported occurrence of wewokellid sponges (Calcarea, Heteractinida) from the Permian of central Iran. Facies 51: 516-521.","ROSS C. A. & ROSS J. R. P. 1987. - Biostratigrafic zonation of Late Paleozoic depositional sequences, in ROSS C. A. & HAMAN D. (eds), Timing depositional history of eustatic sequences constraints on seismic stratigraphy. Cushman Foundation for Foraminiferal Research, Special Publication 24: 151-168.","ROSS J. R. P. 1978. - Biogeography of Permian Ectoproct Bryozoa. Palaeontology 21 (2): 341-356.","ROSS J. R. P. & ROSS C. A. 1996. - Bryozoan evolution and dispersal and Paleozoic sea-level fluctuations, in GORDON D. P., SMITH A. M. & GRANT- MACKIE J. A. (eds), Bryozoans in Space and Time (Proceedings of the 10th International Bryozology Association Conference, Wellington, New Zealand, 1995). NIWA, Wellington: 243-258.","SAKAGAMI S. 1961. - Japanese Permian Bryozoa. Paleontological Society of Japan, Special Papers 7: 1-58.","SAKAGAMI S. 1964. - Bryozoa of Akiyoshi; part 1, Permian Bryozoa from the Shigeyasu quarry. Bulletin Akioyoshi-dai Science Museum 3: 1-24.","SAKAGAMI S. 1966. - The cryptostomatous Bryozoa from Ko Muk, Peninsular Thailand; contribution to the geology and palaeontology of southeast Asia, in KOBAYASHI T. & TORIYAMA R. (eds), Geology and palaeontology of Southeast Asia. Japanese Journal of Geology and Geography 37 (2-4): 157-168.","SAKAGAMI S. 1968a. - Permian Bryozoa from Khao Ta Mong Rai, Peninsular Thailand. Contributions to the geology and palaeontology of Southeast Asia. 42. Geology and Palaeontology of Southeast Asia 5: 47-67.","SAKAGAMI S. 1968b. - Permian Bryozoa from Khao Phrik, near Rat Buri, Thailand. Contributions to the geology and palaeontology of Southeast Asia. 43. Geology and Palaeontology of Southeast Asia 4: 45-66.","SAKAGAMI S. 1968c. - Permian Bryozoa from Khao Chong Krachok, Peninsular Thailand. Contributions to the geology and palaeontology of Southeast Asia. 44. Geology and Palaeontology of Southeast Asia 4: 67-81.","SAKAGAMI S. 1970. - Addition to the Permian Bryozoa from Ko Muk, Peninsular Thailand. Contributions to the geology and palaeontology of Southeast Asia. 53. Geology and Palaeontology of Southeast Asia 8: 43-68.","SAKAGAMI S. 1971. - On the Palaeozoic Bryozoa collected by Dr. C.K. Burton from Chumphon, Peninsular Thailand. Contributions to the geology and palaeontology of Southeast Asia. 92. Geology and Palaeontology of Southeast Asia 9: 135-146.","SAKAGAMI S. 1976. - On the Permian Bryozoa from the northern part of Sainbeyli, Central Turkey. Transactions and Proceedings of the Palaeontological Society of Japan, New Series 103: 398-405.","SAKAGAMI S. 1980. - Permian Ectoprocta (Bryozoa) from the Abadeh Region, Central Iran. Transactions and Proceedings of the Palaeontological Society of Japan, New Series 118: 269-289.","SAKAGAMI S. 1985. - Paleogeographic distribution of Permian and Triassic Ectoprocta (Bryozoa), in NA- KAZAWA K. & DICKINS J. M. (eds), The Tethys, her Paleogeography and Paleobiogeography from Paleozoic to Mesozoic. Tokai University Press, Tokai: 171-183.","SAKAGAMI S. 1995. - Upper Paleozoic bryozoans from the Lake Titicaca region, Bolivia. Part 1. Introductory remarks, stratigraphy and systematic paleontology. Part 2. Systematic paleontology. Transactions and Proceedings of the Palaeontological Society of Japan, New Series 180: 226-281.","SAKAGAMI S. 1999. - Permian bryozoans from some localities in the Khao Hin Kling Area near Phetchbun, north-central Thailand. Bulletin of the Kitakyushi Museum of Natural History 18: 77-103.","SAKAGAMI S. 2000. - Middle Permian Bryozoa from Irian Jaya, Indonesia. Bulletin of the National Science Museum. Series C: Geology & Paleontology 26 (3-4): 139-168.","SENOWBARI- DARYAN B. & HAMEDANI A. 2002. - First report of the occurrence of Amblysiphonella (thalamid sponge) in Permian of Iran and description of A. iranica n. sp. from central Iran. Revue de Paleobiologie 21 (2): 795-801.","SNYDER E. M. 1991a. - Revised taxonomic approach to acanthocladiid Bryozoa, in BIGEY F. P. (ed.), Bryozoaires actuels et fossiles, Bryozoa living and fossil. Bulletin de la Societe des Sciences naturelles de l'Ouest de la France, Memoire HS 1: 431-445.","SNYDER E. M. 1991b. - Revised taxonomic procedures and paleoecological applications for some North American Mississippian Fenestellidae and Polyporidae (Bryozoa). Palaeontographica Americana 57: 1-275.","STOCKLIN J. 1968. - Structural history and tectonics of Iran. A Review. Bulletin of American Association of Petroleum Geologists 52 (7): 1229-1258.","TERMIER H. & TERMIER G. 1971. - Bryozoaires du Paleozoique superieur de l'Afghanistan. Documents des Laboratoires de Geologie de la Faculte des Sciences de Lyon 47: 1-52.","VINE G. R. 1883. - Fourth report of the committee appointed for the purpose of reporting on fossil Polyzoa. British Association for the Advancement in Science Report of 53rd Meeting, 1883: 161-209.","WAAGEN W. & PICHL I. 1885. - Salt Range fossils. Productus -limestone fossils. Memoirs of the Geological Survey of India, Paleontologica Indica Series 13, 1 (5): 771-966.","WAAGEN W. & WENTZEL I. 1886. - Salt Range fossils. Productus -limestone fossils: Coelenterata. Memoirs of the Geological Survey of India, Paleontologica Indica Series 13, 1 (6): 835-924.","WYSE JACKSON P. N. 1993. - Case 2847. Ascopora Trautschold, 1876 (Bryozoa, Cryptostomata): proposed designation of Ceriopora nodosa Fischer von Waldheim, 1837 as the type species. Bulletin of Zoological Nomenclature 50: 13-15.","XIA F. 1986. - Carboniferous and Early Permian bryozoans from Xainza, Northern Xizang. Bulletin of Nanjing Institute of Geology and Palaeontology, Academia Sinica 10: 201-254.","XIA F. 1991. - [Early-Middle Permian bryozoans from Rutog region, Xizan (Tibet)], in SUN D., XU J. et al. (eds), [Papers for Scientific Co-Expedition of the Nanjing Institute of Geology and Palaeontology, Academia Sinica and Regional Geological Survey Team, Geological Bureau of Xizang Stratigraphy and Palaeontology of Permian, Jurassic and Cretaceous of the Rutog Region, Xizan (Tibet)]. Nanjing University Press, Nanjing: 165-210 (in Chinese, with English abstract).","YANAGIDA J. (& A RESEARCH GROUP 1988). - Biostratigraphic Study of Paleozoic and Mesozoic Groups in Central and Northern Thailand. An Interim Report. Department of Geology, Faculty of Sciences, Kyushu University, Kyushu, 46 p.","YANG J. & LU L. 1962. - [Paleozoic bryozoans of Gilianshan], in [Geology of Gilianshan]. Science Press, Beijing 4: 1-114 (in Chinese).","YANG J. & LU L. 1980. - [Upper Permian bryozoans from eastern Sichuan and western Guizhou]. Gushengwu Xuebao [= Acta Palaeontologica Sinica] 19 (6): 461-474 (in Chinese, with English abstract).","YANG J., LU L. & XIA F. 1981. - [Upper Paleozoic bryozoans from Xizang], in [Palaeontology of Xizang]. Science Press, Beijing: 81-100 (in Chinese).","YOUNG J. & YOUNG J. 1875. - New species of Glauconome from the Carboniferous limestone strata of the West of Scotland. Proceedings of the Natural History Society of Glasgow 2: 325-335.","ZAKHAROV Y. D., BORISKINA N. G. & POPOV A. M. 2001. - [The Reconstruction of Late Paleozoic and Mesozoic Marine Environments from Isotopic Data (Evidence from North Eurasia)]. Dalnauka, Vladivostok, 112 p. (in Russian)."]}

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