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25. Tylodinus complicatus Luna-Cozar, sp. nov. (Figures 24–25, 74-75, 118, 131) Diagnosis. Length male, 2.2-2.5 mm, female, 2.2-2.5 mm. Width male, 1.0- 1.1 mm, female 1.1-1.3 mm. Body approximately 2.1x longer than wide, cuticle sometimes shiny. Head with yellow scales; vertex with dense to contiguous scales, uniformly distributed along vertex, similar in coloration and tonality to scales on head; frons moderately concave; rostrum moderately robust, moderately carinate, apical area puncticulate, eyes moderately convex. Pronotum in dorsal view weakly convex, outline with anterior constriction and without posterior constriction (Figure 10); in dorsal view with sides slightly divergent from base to midlength, then convergent to apex, constricted beyond middle; disc hollowed, punctures dense to contiguous at base and progressively more widely spaced, smaller, shallower to apex, clothed with yellow scales, very dense on lateral sides and flanks, anterior area forming thick longitudinal fringe along middle and narrow strip that diverges from middle base to flanks, with dark brown scales on middle and posterior areas of disc (Figure 14, 74). Elytra black with red coloration on base and along intervals 1-2 and intervals 1-5 on declivity (Figure 74-75), basal margin slightly sinuate (Figure 3), clothed with scattered yellow scales, dense on tubercles on intervals 1-5 at declivity base, and on intervals 1-3 along declivity, stria 10 complete; tubercle distribution as follows: I2 with a small and round tubercle on declivity base, I3 with two tubercles, the first tubercle subbasal, the second tubercle on the middle of elytral declivity, I4 with two small tubercles, first tubercle between tubercles on I3, the other tubercle on declivity base, smaller than tubercle on I2 (but in some specimens of same size), I5 with two small tubercles in same position as tubercles on I3, tubercles on I3 longer than other tubercles, surface without granules. Metasternum flat, glabrous at median section, metasternal teeth absent. Male abdomen with ventrites 3-5 in same plane as ventrites 1-2, ventrite 1 weakly concave, almost flat, glabrous at median section, ventrite 2 as long as 3-4, median section glabrous (Figure 23), female with transverse line of scales interrupted medially (Figure 24), ventrite 5 longer than ventrite 1, flat, with anterior impression. Femora armed, hind femur length reaching elytral apex, middle and hind tibiae subparallel, premucro absent. Male genitalia (Figure 118) with median lobe in lateral view moderately curved, stout; in ventral view as long as half length of apodemes, lateral sides subparallel, convex at sides beyond 0.27 of length from base, apex acute; apical process sinuate, moderately curved downward, small (0.17-0.25 as long as body of median lobe); in ventral view constricted beyond median orifice, rhomboid shape apically (Figure 98e). Female with tergite eight trapezoidal, sternite eight doliform, apex not constricted, median membranous area 2/3 as long as basal plate, spermatheca shallowly curved, ramus indistinct, nodulus slender. Comments. Male genitalia in T. complicatus, T. rufus, T. noctis and T. dominicus are very similar; however, T. complicatus is distinguished from its allies by the small size, elytra coloration and hind tibiae without external teeth. Geographic distribution. México (Chiapas). Habitat and elevation. Specimens were collected from leaf litter in cloud forest between 1800-2260 m. Derivation of specific name. Name derived from Latin complicatus = complicated, confused, intricate. Material examined. Total 64 males, 39 females. Holotype male (CMNC): México: Chiapas, Pueblo Nuevo Solistahuacán, La Yerbabuena, 2.1 km NW Pueblo Nuevo, Solistahuacan, 23.ix.1992, Coll. R. Anderson. Paratypes: México: Chiapas, Bochil, 15.1 km NW de Bochil, elev. 1930 m (17° 1'51.6"N, 92° 52'5.9"W), 24.ix.1992, coll. R. S. Anderson — 1 male, 1 female (CMNC); 17.3 km NW de Bochil, elev. 1800 m, (17° 1'53.8"N, 92° 50'49.2"W), 24.ix.1992, coll. R. S. Anderson — 1 male (CMNC); Pueblo Nuevo Solistahuacán, Reserva la Yerbabuena, 2.1 km NW de Pueblo Nuevo, Solistahuacán, elev. 2100 m (17° 11'0.2"N, 92°,53'52.4"W), 23.ix.1992, coll. R. S. Anderson — 4 males, 2 females (CMNC, ECOS); Coapilla, 5 km NNW Coapilla, elev. 1915 m, (17.18355°N 93.15222°W), 24.v.2008, coll. LLAMA (Wm-A-04-2-01) — 1 female (CMNC); 5 km NE Coapilla, elev. 1990 m (17°10'14.5"N, 93°8'12.5"W), 26.v.2008, coll. R. S. Anderson (2008-035)— 3 males, 2 females (BMNH, CMNC); 1.8 km NE Jct Rds. Tapalapa / Ocotepeque, elev. 1810 m (17.17314°N 93.14607°W), 26.v.2008, coll. R. S. Anderson — 1 male (CMNC); 2.5 km NW Jct Rds. Tapalapa / Ocotepeque, elev. 1960 m (17.17602°N 93.13293°W), 26.v.2008, coll. R. S. Anderson — 2 females (CMNC); Cerro de Tapalapa, microwave towers, elev. 2260 m (17.18786°N 93.12308°W), 26.v.2008, coll. R. S. Anderson — 6 males, 1 female (CMNC, ECOS); 6 km NE Coapilla, elev. 1990 m (17°10'33.6"N, 93° 7'58.4"W), 26.v.2008, coll. M. G. Branstetter — 1 male (CMNC); 5 km NE Coapilla, elev. 1990 m, 26.v.2008, coll. M. G. Branstetter — 1 male (CMNC); 7.3 km NE Coapilla, elev. 2200 m (17°10'59.9"N, 93°7'0.1"W), 12.vii.2007, coll. M. G. Branstetter — 1 male, 1 female (CMNC); 9 km NE Coapilla, elev. 1990, 24.v.2008, coll. LLAMA (Wm-A-04-1-01), 1 female (CMNC); ca. 10.5 km NE Coapilla, elev. 1990 (17° 9'55.1"N, 93° 8'20.4"W), 13.vii.2007, coll. R. Anderson — 3 males, 1 female (ECOS); Cerro El Calvario, near Tapalapa, elev. 2000 m (17°10'25.7"N, 93°7'52.3"W), 23.vii.2003, coll. R. Anderson — 2 males (CMNC); Coapilla (6-8 km NE), elev. 2030 m (17°10'25"N, 93°7'52"W), 26.vii.2005, coll. R. Anderson — 1 male, 1 female (CMNC); Carretera Coapilla- Ocotepec, 26.v.2006, coll. J. L. Cozar — 2 males (ECOS); El Bosque, 10 km W El Bosque, elev. 1475 m (17°2'36.6"N, 92°45'43.6"W), 15.ix.1992, coll. R. Anderson — 1 male (CMNC); Rayón, 8.9 km E Rayón, elev. 1500 m (17°10'58.1"N, 92°55'47.3"W), 19.ix.1991, coll. R. Anderson — 9 males, 8 females (BMNH, CMNC, CWOB); Tapalapa, Cerro El Calvario, ca. 12 km NE Coapilla, elev. 2000 m (17°10'28.2"N, 93°7'53.4"W), 13.vii.2007, coll. R. Anderson — 1 female (CMNC); Cerro El Calvario, ca. 14 km NE Coapilla, elev. 2150 m (17°10'57.4"N, 93° 7'5.5"W), 13/vii/2007, coll. R. Anderson — 2 males, 1 female (CMNC); Cerro El Calvario, near Tapalapa, elev. 2200 m (17°11'11.8"N, 93°7'21"W), 23.vii.2003, coll. R. Anderson — 1 male, 1 female (ECOS); Carretera Tapalapa-Coapilla a 5.8 km NE de Coapilla, elev. 2000 m (17° 10'33.22"N, 93° 9'51.9"W), 13.vii.2007, coll. J. L. Cozar — 1 male, 1 female (ECOS); Carretera Tapalapa-Coapilla a 7.3 Km NE de Coapilla, elev. 2212 m (17° 10'57.94"N, 93°7'5.29"W), 13.vii.2007, coll. J. L. Cozar — 1 male (ECOS); same locality, elev. 2190 (17°10'57.48"N, 93°7'5.4"W), 26.v.2006, coll. J. L. Cozar — 1 male (ECOS); 5 km NE Coapilla, elev. 1990 m (17.17549°N 93.13251°W), 25.v.2008, LLAMA (Wa-A-04-1-17) – 1 female (ECOS); same locality, (17.17550°N 93.13227°W), 25.v.2008, LLAMA (Wa-A-04-1-23) – 1 male, 1 female (ECOS); same locality, (17.17550°N 93.13217°W), 25.v.2008, LLAMA (Wa-A-04-1-25) – 1 female (ECOS); same locality, (17.17550°N 93.13212°W), 25.v.2008, LLAMA (Wa-A-04-1-26) – 1 female (CMNC); same locality, (17.17550°N 93.13203°W), 25.v.2008, LLAMA (Wa-A-04-1-28) – 1 male (CMNC); same locality, (17.17586°N 93.13197°W), 25.v.2008, LLAMA (Wa-A-04-1-44) – 1 male (CMNC); same locality, (17.17585°N 93.13202°W), 25.v.2008, LLAMA (Wa-A-04-1-46) – 2 males (CMNC); same locality, (17.17585°N 93.13211°W), 25.v.2008, LLAMA (Wa- A-04-1-47) – 1 female (CMNC); 5 km NNW Coapilla, elev. 1915 m (17.18300°N 93.15196°W), 25.v.2008, LLAMA (Wa-A-04-2-14) — 2 males, 1 female (CMNC, ECOS); same locality, (17.18342°N 93.15215°W), LLAMA (Wa-A-04-2-04) — 1 male (ECOS); same locality, (17.18334°N 93.15211°W), LLAMA (Wa-A-04-2- 06) — 1 female (CMNC); same locality, (17.18313°N 93.15202°W), LLAMA (Wa-A-04-2-11) — 1 male, 1 female (ECOS); same locality, (17.18305°N 93.15199°W), LLAMA (Wa-A-04-2-13) — 1 female (CMNC); same locality, (17.18296°N 93.15197°W), LLAMA (Wa-A-04-2-15) — 1 female (CMNC); same locality, (17.18292°N 93.15195°W), LLAMA (Wa-A-04-2-16) — 1 male (CMNC); same locality, (17.18287°N 93.15194°W), LLAMA (Wa-A-04-2-17) — 2 males (CMNC); same locality, (17.18292°N 93.15195°W), LLAMA (Wa-A-04-2-16) — 1 male (CMNC); same locality, (17.18255°N 93.15163°W), LLAMA (Wa-A-04-2-27) — 1 male (ECOS); same locality, (17.18227°N 93.15139°W), LLAMA (Wa-A-04-2-35) — 1 male (CMNC); same locality, (17.18248°N 93.15129°W), LLAMA (Wa-A-04-2-40) — 1 female (CMNC); same locality, (17.18277°N 93.15116°W), LLAMA (Wa-A-04-2-47) — 1 male (ECOS); same locality, (17.18281°N 93.15114°W), LLAMA (Wa-A-04-2-48) — 1 female (CMNC); same locality, (17.18289°N 93.15110°W), LLAMA (Wa-A-04-2-50) — 1 male (CMNC); Estación de Microondas, elev. 2260 m (17°11'16.4"N, 93°7'23.2"W), 26.v.2006, coll. J. L. Cozar – 1 male, 1 female (ECOS); Tenejapa, Ojo de Agua, elev. 1800 m, 21.vii.2003, coll. R. Anderson — 1 female (CMNC); Yashanal, elev. 1650 m (16°49'49"N, 92°27'9"W), 21.vii.2003, coll. R. S. Anderson — 2 males (CMNC)., Published as part of Luna-Cozar, Jesús, Anderson, Robert S., Jones, Robert W. & León-Cortés, Jorge L., 2014, A taxonomic monograph of the genus Tylodinus Champion (Coleoptera: Curculionidae: Cryptorhynchinae: Tylodina) of Chiapas, Mexico, pp. 1-63 in Zootaxa 3788 (1) on page 44, DOI: 10.11646/zootaxa.3788.1, http://zenodo.org/record/4913761

25. Tylodinus complicatus Luna-Cozar, sp. nov. (Figures 24–25, 74-75, 118, 131) Diagnosis. Length male, 2.2-2.5 mm, female, 2.2-2.5 mm. Width male, 1.0- 1.1 mm, female 1.1-1.3 mm. Body approximately 2.1x longer than wide, cuticle sometimes shiny. Head with yellow scales; vertex with dense to contiguous scales, uniformly distributed along vertex, similar in coloration and tonality to scales on head; frons moderately concave; rostrum moderately robust, moderately carinate, apical area puncticulate, eyes moderately convex. Pronotum in dorsal view weakly convex, outline with anterior constriction and without posterior constriction (Figure 10); in dorsal view with sides slightly divergent from base to midlength, then convergent to apex, constricted beyond middle; disc hollowed, punctures dense to contiguous at base and progressively more widely spaced, smaller, shallower to apex, clothed with yellow scales, very dense on lateral sides and flanks, anterior area forming thick longitudinal fringe along middle and narrow strip that diverges from middle base to flanks, with dark brown scales on middle and posterior areas of disc (Figure 14, 74). Elytra black with red coloration on base and along intervals 1-2 and intervals 1-5 on declivity (Figure 74-75), basal margin slightly sinuate (Figure 3), clothed with scattered yellow scales, dense on tubercles on intervals 1-5 at declivity base, and on intervals 1-3 along declivity, stria 10 complete; tubercle distribution as follows: I2 with a small and round tubercle on declivity base, I3 with two tubercles, the first tubercle subbasal, the second tubercle on the middle of elytral declivity, I4 with two small tubercles, first tubercle between tubercles on I3, the other tubercle on declivity base, smaller than tubercle on I2 (but in some specimens of same size), I5 with two small tubercles in same position as tubercles on I3, tubercles on I3 longer than other tubercles, surface without granules. Metasternum flat, glabrous at median section, metasternal teeth absent. Male abdomen with ventrites 3-5 in same plane as ventrites 1-2, ventrite 1 weakly concave, almost flat, glabrous at median section, ventrite 2 as long as 3-4, median section glabrous (Figure 23), female with transverse line of scales interrupted medially (Figure 24), ventrite 5 longer than ventrite 1, flat, with anterior impression. Femora armed, hind femur length reaching elytral apex, middle and hind tibiae subparallel, premucro absent. Male genitalia (Figure 118) with median lobe in lateral view moderately curved, stout; in ventral view as long as half length of apodemes, lateral sides subparallel, convex at sides beyond 0.27 of length from base, apex acute; apical process sinuate, moderately curved downward, small (0.17-0.25 as long as body of median lobe); in ventral view constricted beyond median orifice, rhomboid shape apically (Figure 98e). Female with tergite eight trapezoidal, sternite eight doliform, apex not constricted, median membranous area 2/3 as long as basal plate, spermatheca shallowly curved, ramus indistinct, nodulus slender. Comments. Male genitalia in T. complicatus, T. rufus, T. noctis and T. dominicus are very similar; however, T. complicatus is distinguished from its allies by the small size, elytra coloration and hind tibiae without external teeth. Geographic distribution. México (Chiapas). Habitat and elevation. Specimens were collected from leaf litter in cloud forest between 1800-2260 m. Derivation of specific name. Name derived from Latin complicatus = complicated, confused, intricate. Material examined. Total 64 males, 39 females. Holotype male (CMNC): México: Chiapas, Pueblo Nuevo Solistahuacán, La Yerbabuena, 2.1 km NW Pueblo Nuevo, Solistahuacan, 23.ix.1992, Coll. R. Anderson. Paratypes: México: Chiapas, Bochil, 15.1 km NW de Bochil, elev. 1930 m (17° 1'51.6"N, 92° 52'5.9"W), 24.ix.1992, coll. R. S. Anderson — 1 male, 1 female (CMNC); 17.3 km NW de Bochil, elev. 1800 m, (17° 1'53.8"N, 92° 50'49.2"W), 24.ix.1992, coll. R. S. Anderson — 1 male (CMNC); Pueblo Nuevo Solistahuacán, Reserva la Yerbabuena, 2.1 km NW de Pueblo Nuevo, Solistahuacán, elev. 2100 m (17° 11'0.2"N, 92°,53'52.4"W), 23.ix.1992, coll. R. S. Anderson — 4 males, 2 females (CMNC, ECOS); Coapilla, 5 km NNW Coapilla, elev. 1915 m, (17.18355°N 93.15222°W), 24.v.2008, coll. LLAMA (Wm-A-04-2-01) — 1 female (CMNC); 5 km NE Coapilla, elev. 1990 m (17°10'14.5"N, 93°8'12.5"W), 26.v.2008, coll. R. S. Anderson (2008-035)— 3 males, 2 females (BMNH, CMNC); 1.8 km NE Jct Rds. Tapalapa / Ocotepeque, elev. 1810 m (17.17314°N 93.14607°W), 26.v.2008, coll. R. S. Anderson — 1 male (CMNC); 2.5 km NW Jct Rds. Tapalapa / Ocotepeque, elev. 1960 m (17.17602°N 93.13293°W), 26.v.2008, coll. R. S. Anderson — 2 females (CMNC); Cerro de Tapalapa, microwave towers, elev. 2260 m (17.18786°N 93.12308°W), 26.v.2008, coll. R. S. Anderson — 6 males, 1 female (CMNC, ECOS); 6 km NE Coapilla, elev. 1990 m (17°10'33.6"N, 93° 7'58.4"W), 26.v.2008, coll. M. G. Branstetter — 1 male (CMNC); 5 km NE Coapilla, elev. 1990 m, 26.v.2008, coll. M. G. Branstetter — 1 male (CMNC); 7.3 km NE Coapilla, elev. 2200 m (17°10'59.9"N, 93°7'0.1"W), 12.vii.2007, coll. M. G. Branstetter — 1 male, 1 female (CMNC); 9 km NE Coapilla, elev. 1990, 24.v.2008, coll. LLAMA (Wm-A-04-1-01), 1 female (CMNC); ca. 10.5 km NE Coapilla, elev. 1990 (17° 9'55.1"N, 93° 8'20.4"W), 13.vii.2007, coll. R. Anderson — 3 males, 1 female (ECOS); Cerro El Calvario, near Tapalapa, elev. 2000 m (17°10'25.7"N, 93°7'52.3"W), 23.vii.2003, coll. R. Anderson — 2 males (CMNC); Coapilla (6-8 km NE), elev. 2030 m (17°10'25"N, 93°7'52"W), 26.vii.2005, coll. R. Anderson — 1 male, 1 female (CMNC); Carretera Coapilla- Ocotepec, 26.v.2006, coll. J. L. Cozar — 2 males (ECOS); El Bosque, 10 km W El Bosque, elev. 1475 m (17°2'36.6"N, 92°45'43.6"W), 15.ix.1992, coll. R. Anderson — 1 male (CMNC); Rayón, 8.9 km E Rayón, elev. 1500 m (17°10'58.1"N, 92°55'47.3"W), 19.ix.1991, coll. R. Anderson — 9 males, 8 females (BMNH, CMNC, CWOB); Tapalapa, Cerro El Calvario, ca. 12 km NE Coapilla, elev. 2000 m (17°10'28.2"N, 93°7'53.4"W), 13.vii.2007, coll. R. Anderson — 1 female (CMNC); Cerro El Calvario, ca. 14 km NE Coapilla, elev. 2150 m (17°10'57.4"N, 93° 7'5.5"W), 13/vii/2007, coll. R. Anderson — 2 males, 1 female (CMNC); Cerro El Calvario, near Tapalapa, elev. 2200 m (17°11'11.8"N, 93°7'21"W), 23.vii.2003, coll. R. Anderson — 1 male, 1 female (ECOS); Carretera Tapalapa-Coapilla a 5.8 km NE de Coapilla, elev. 2000 m (17° 10'33.22"N, 93° 9'51.9"W), 13.vii.2007, coll. J. L. Cozar — 1 male, 1 female (ECOS); Carretera Tapalapa-Coapilla a 7.3 Km NE de Coapilla, elev. 2212 m (17° 10'57.94"N, 93°7'5.29"W), 13.vii.2007, coll. J. L. Cozar — 1 male (ECOS); same locality, elev. 2190 (17°10'57.48"N, 93°7'5.4"W), 26.v.2006, coll. J. L. Cozar — 1 male (ECOS); 5 km NE Coapilla, elev. 1990 m (17.17549°N 93.13251°W), 25.v.2008, LLAMA (Wa-A-04-1-17) – 1 female (ECOS); same locality, (17.17550°N 93.13227°W), 25.v.2008, LLAMA (Wa-A-04-1-23) – 1 male, 1 female (ECOS); same locality, (17.17550°N 93.13217°W), 25.v.2008, LLAMA (Wa-A-04-1-25) – 1 female (ECOS); same locality, (17.17550°N 93.13212°W), 25.v.2008, LLAMA (Wa-A-04-1-26) – 1 female (CMNC); same locality, (17.17550°N 93.13203°W), 25.v.2008, LLAMA (Wa-A-04-1-28) – 1 male (CMNC); same locality, (17.17586°N 93.13197°W), 25.v.2008, LLAMA (Wa-A-04-1-44) – 1 male (CMNC); same locality, (17.17585°N 93.13202°W), 25.v.2008, LLAMA (Wa-A-04-1-46) – 2 males (CMNC); same locality, (17.17585°N 93.13211°W), 25.v.2008, LLAMA (Wa- A-04-1-47) – 1 female (CMNC); 5 km NNW Coapilla, elev. 1915 m (17.18300°N 93.15196°W), 25.v.2008, LLAMA (Wa-A-04-2-14) — 2 males, 1 female (CMNC, ECOS); same locality, (17.18342°N 93.15215°W), LLAMA (Wa-A-04-2-04) — 1 male (ECOS); same locality, (17.18334°N 93.15211°W), LLAMA (Wa-A-04-2- 06) — 1 female (CMNC); same locality, (17.18313°N 93.15202°W), LLAMA (Wa-A-04-2-11) — 1 male, 1 female (ECOS); same locality, (17.18305°N 93.15199°W), LLAMA (Wa-A-04-2-13) — 1 female (CMNC); same locality, (17.18296°N 93.15197°W), LLAMA (Wa-A-04-2-15) — 1 female (CMNC); same locality, (17.18292°N 93.15195°W), LLAMA (Wa-A-04-2-16) — 1 male (CMNC); same locality, (17.18287°N 93.15194°W), LLAMA (Wa-A-04-2-17) — 2 males (CMNC); same locality, (17.18292°N 93.15195°W), LLAMA (Wa-A-04-2-16) — 1 male (CMNC); same locality, (17.18255°N 93.15163°W), LLAMA (Wa-A-04-2-27) — 1 male (ECOS); same locality, (17.18227°N 93.15139°W), LLAMA (Wa-A-04-2-35) — 1 male (CMNC); same locality, (17.18248°N 93.15129°W), LLAMA (Wa-A-04-2-40) — 1 female (CMNC); same locality, (17.18277°N 93.15116°W), LLAMA (Wa-A-04-2-47) — 1 male (ECOS); same locality, (17.18281°N 93.15114°W), LLAMA (Wa-A-04-2-48) — 1 female (CMNC); same locality, (17.18289°N 93.15110°W), LLAMA (Wa-A-04-2-50) — 1 male (CMNC); Estación de Microondas, elev. 2260 m (17°11'16.4"N, 93°7'23.2"W), 26.v.2006, coll. J. L. Cozar – 1 male, 1 female (ECOS); Tenejapa, Ojo de Agua, elev. 1800 m, 21.vii.2003, coll. R. Anderson — 1 female (CMNC); Yashanal, elev. 1650 m (16°49'49"N, 92°27'9"W), 21.vii.2003, coll. R. S. Anderson — 2 males (CMNC).