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Supplemental material, sj-docx-1-hol-10.1177_09596836221096292 for Biodiversity and ecology of plants and arthropods on the last preserved glacier of the Apennines mountain chain (Italy) by Barbara Valle, Michele di Musciano, Mauro Gobbi, Marco Bonelli, Enzo Colonnelli, Giulio Gardini, Massimo Migliorini, Paolo Pantini, Adriano Zanetti, Emanuele Berrilli, Anna Rita Frattaroli, Davide Fugazza, Anna Invernizzi and Marco Caccianiga in The Holocene

Xantholinus audrasi was detected for the first time in South Tyrol in two different sites.

Lesteva (s.str.) fontinalis Kiesenwetter, 1850 (Figs. 9, 14���15) Lesteva fontinalis Kiesenwetter, 1850: 222; Ganglbauer 1895: 713, Luze 1903: 195, Portevin 1929: 438, Lohse 1955a: 505, 1960: 3 Lesteva (s.str.) fontinalis: Zanetti 1987: 362 Lesteva (s.str.) fontinalis fontinalis: Tronquet 2006: 33 Lesteva fontinalis gustavi Herman, 2001a: 37 (replacement name of L. fontinalis truncata Lohse, 1960: 5) syn.n. Lesteva nigra Kraatz, 1857: 934 Lesteva cavernicola Scheerpeltz, 1933: 1063 Type material examined. Lectotype (here designated) of Lesteva fontinalis Kiesenwetter, 1850 ♂ [The specimen is glued sideways on small triangular card; head, pronotum and left legs are missing]: ���Kiesenwetter��� , ���fontinalis | M[ons]. Serr[atus Cataloniae].��� , ���Sammlung | Cl. M��ller��� , ��� Holotypus | (vermutlich!) [handwritten] | Lesteva [handwritten] | fontinalis Kiesw. [handwritten] | Staatssamml.M��nchen��� , ���zweifellos die Type, | da nur dies eine Ex. | von mon serrat | St..[���illegible���]��� , ���Zool. Staatsslg. | M��nchen��� , ��� LECTOTYPE | Lesteva fontinalis | Kiesenwetter, 1850 | Shavrin A.V. des. 2021��� (ZSM). Paratype of Lesteva fontinallis truncata Lohse, 1960 ♂ [aedeagus glued under the specimen]: ��� Espagne: Estremadura | Guadalupe (scierie) | rio Guadalupejo | V���1958, G.Fagel��� , ��� Lesteva [handwritten] | fontinalis [handwritten] | ssp truncata [handwritten] | det.Dr.G.A.Lohse��� , ���Para- | type��� , ���Coll. | G. A. Lohse | MHNG-1994��� , ��� PARATYPE | Lesteva | fontinalis | truncata ssp. n. | Det Lohse 1960 ��� , ��� PARATYPE | Lesteva fontinalis | gustavi nom.n. | Det. Herman 2001��� , ���MHNG | ENTO | 00087335��� (MHNG). Paratype ♀, same labels except for ���MHNG | ENTO | 00087336��� (MHNG). Paratype ♂: ��� Espagne: Estremadura | Guadalupe | Fuente del Trincho | V���1958, G.Fagel��� , ���fontinalis [handwritten] | ssp. truncata [hadwritten] | det.Dr.G.A.Lohse��� , ���Coll. | G. A. Lohse | MHNG- 1994��� , ��� Lesteva fontinalis | infrasp. gustavi nom.n. | Det. Herman 2001��� , ���MHNG | ENTO | 00087337��� (MHNG). All paratypes with additional printed label: ��� Lesteva (str.) fontinalis Kiesenwetter, 1850 | Shavrin A.V. det. 2021���. Material examined. SPAIN: ASTURIAS: 2 ♂♂: Puerto Pajares. A. Kricheldorff leg. (NMPC); PAIS VASCO: 1 ♂: Bilbao (MHNG); CATALU��A: 1 ♂: Col d`Ares (Spain Pyren.). 31.05.1971. G. Lohse leg. (MHNG); 2 ♂♂: Barcelona, Monserrat. 22.06.2006. L. Fancello leg. (cZan); 1 ♂: Sierra de Montseny. 13.05.1981. P. Zwick leg. (MHNG); 2 ♂♂, 2 ♀♀: Seo de Urgel (Arfa), de la Coma. 05-06.1962. G. Fagel leg. (MHNG); 1 ♀: Seo de Urgel (graviers du Segre). 05-06.1962. G. Fagel leg. (MHNG); ARAG��N: 1 ♂: Ardisa (MHNG); 3 ♂♂, 2 ♀♀: Ordesa, Huesca. 31.05.1965. A. Comellini leg. (MHNG); 12 ♂♂, 2 ♀♀: Teruel, Albarracin. 1200 m a.s.l. 23.05.1993. A. Matern (cA, cW, cSh); 1 ♂, 1 ♀: Mezquita de Jarque. 18.06.1970. A. Comellini leg. (MHNG); CASTILLA Y L��ON: 5 ♂♂, 5 ♀♀: Sierra de la Demanda. H. Meybohm leg. (MHNG); 1 ♂: Santa Colomba de Somoza, Astorga. 31.05.1970. A. Comellini leg. (MHNG); MADRID: 1 ♂: Escorial (MHNG); CASTILLA LA MANCHA: 3 ♂♂: Las Torcas. 1200 m a.s.l. 23.05.1962. H. Freude leg. (MHNG, ZSM); ANDALUS��A: 3 ♂♂: Sierra de Cazorla, sources of Guadalquivir. 25.06.1975. M. Daccordi leg. (cZan); 1 ♀: Sierra de Cazorla. 23.06.1975. M. Daccordi leg. (cZan); 2 ♂♂: same locality. 27.05.2009. T. Struyve leg. (cZan); 1 ♀: Sierra de Segura, Guadalquivir, 37��50���N - 2��59���W. 2276 ft. 21.05.2002. Starke leg. (cA); 4 ♂♂, 2 ♀♀: Sierra Magina, 37��72���N - 3��46���W. 2000 m a.s.l. 03- 04.05.2014. A.V. Shavrin leg. (cSh); 1 ♂, 2 ♀♀: ��� Kiesenwetter���, ��� Jaen (Sierra)���, ��� Sammlung Cl. M��ller���, ��� fontinalis Kiesw��� (ZSM); 1 ♂: ��� Jaen���, ��� Sammlung Cl. M��ller���, ��� fontinalis Kiesw��� (ZSM); 1 ♂: Jaen (MHNG); 1 ♂: Jaen, Jaen-Valdepenas Rd., 18 km S Jaen. 1000 m a.s.l. 25.05.1967. M.E. Bacchus & D. Levey (MHNG); 1 ♀, 1 ex.: ���S.d. Jaen���, ��� Sammlung Cl. M��ller, ��� fontinalis ?��� (ZSM); 1 ♂: Cazorla. 800-2000 m a.s.l. 05.1943. G. Frey & C. Koch (MHNG); 1 ♂: ��� fontinalis Ksw type. M��� [illegible]���, ������[illegible] pr��s Granada���, ��� Andalousie��� (IRSN); 1 ♂: ��� Spanien, 1935 Sierra Nevada 2000 m, 16/6 O. Lundblad���, ��� Lesteva fontinalis Kiesw. Det. Scheepeltz 1964 ��� (SMNH); 1 ♂, 1 ♀: ��� S��d. Spanien���, ��� Andalusien��� (NMPC); LOCALITY NOT SPECIFIED: 2 ♂♂: ��� Spanien Scriba���, ��� Coll. K. Neumann��� (SF). Additional material. FRANCE: 1 ♀: ���St. Jean de Luz Basse-Pyrenees��� (NMPC); MOROCCO: 1 ♂, 1 ♀: ���Rif Ketama��� (MHNH); LOCALITY NOT SPECIFIED: 1 ♂, ���Pyr��n[ees].���, ��� fontinalis ���, ��� L. pandellei J. Bohač det. 1980��� (NMPC); 1 ♂: Pyrenees (MHNG). Remarks. Lesteva fontinalis was originally described based on an unspecified number of syntypes from ���Mons serratus [Montserrat] Cataloniae���. During the study of Lesteva from ZSM, the first author found a strongly damaged male, which undoubtedly belongs to the type series. Consequently, this specimen is designated as the lectotype. Lohse (1960) described L. fontinalis truncata as ���kleine, kurzfl��gelige und klein��ugige Rasse��� from Central Spain. Herman (2001a) replaced the preoccupied name with L. fontinalis gustavi Herman, 2001. Based on the study of the type and additional material from Spain and other countries, we did not find pronounced morphological differences between them. Body size, coloration, and the proportions of temples and elytra of L. fontinalis are variable. Thus, the new synonymy is proposed here. Habitus as in Fig. 9. Aedeagus as in Figs. 14���15. Lesteva fontinalis is known from Switzerland, France, Spain, and North Africa (Sch��lke & Smetana 2015). In North Africa it was recorded from Morocco, Algeria, and Tunisia (Fauvel 1871 [=1872], 1878, 1886, 1902, Champion 1898), but all these records require confirmation. It was recorded from Spain by Fauvel (1871a [= 1872]) and Lohse (1955 a, 1960)., Published as part of Shavrin, Alexey V. & Zanetti, Adriano, 2021, Review of the genus Lesteva Latreille, 1797 (Coleoptera: Staphylinidae: Omaliinae Anthophagini) of the Iberian Peninsula, pp. 401-427 in Zootaxa 4966 (4) on pages 408-410, DOI: 10.11646/zootaxa.4966.4.1, http://zenodo.org/record/4738667, {"references":["Kiesenwetter, E. A. H. von (1850) Funfzig Diagnosen unbeschriebener oder wenig bekannter europaischen Kafer. Entomologische Zeitung. Stettin, 11 (7), 217 - 225.","Ganglbauer, L. (1895) Die Kafer von Mitteleuropa. 2. Familienreihe Staphylinoidea. Theil I. Staphylinidae, Pselaphidae. Carl Gerold's Sohn, Wien, 881 pp.","Luze G. (1903) Revision der palaarktischen Arten der Staphyliniden-Gattung Lesteva Latr. Verhandlungen der Kaiserlich- Koniglichen Zoologisch-Botanischen Gesellschaft in Wien, 53, 179 - 197.","Portevin, G. (1929) Histoire naturelle des Coleopteres de France. I. Adephaga. Polyphaga: Staphylinoidea. In: Encyclopedie Entomologique. Vol. 12. P. Lechevalier, Paris, pp. i - xii + 1 - 649.","Lohse, G. A. (1955 a) Lesteva fontinalis Kiesw. und die mit ihr zusammengeworfenen Arten. Mitteilungen der Munchner Entomologischen Gesellschaft, 44 / 45, 503 - 512.","Lohse, G. A. (1960) Die Arten der Gattung Lesteva Latr. (Coleoptera Staphylinidae) der iberischen Halbinsel. Bulletin Institut Royal des Sciences Naturelles de Belgique, 36 (26), 1 - 7.","Zanetti, A. (1987) Coleoptera. Staphylinidae. Omaliinae. In: Fauna d'Italia. Vol. 25. Calderini, Bologna, pp. i - xii + 1 - 472.","Tronquet, M. (2006) Catalogue iconographique des Coleopteres des Pyrenees-orientales. Vol. 1. Staphylinidae. Revue de l`association roussillonnaise d`entomologie, 15, 1 - 79.","Herman, L. H. (2001 a) Nomenclatural changes in the Staphylinidae (Insecta: Coleoptera). Bulletin of the American Museum of Natural History, 264, 1 - 83. https: // doi. org / 10.1206 / 0003 - 0090 (2001) 264 % 3 C 0003: NCITSI % 3 E 2.0. CO; 2","Kraatz, G. (1857) Naturgeschichte der Insecten Deutschlands. Abt. 1. Coleoptera. Zweiter Band. Lief. 3 - 4 & Lief. 5 - 6. Nicolai, Berlin, pp. 377 - 768. & pp. 769 - 1080.","Scheerpeltz, O. von (1933) Staphylinidae VII (pars 129). In: Schenkling, S. (Ed.), Coleopterorum Catalogus. Volumen VI. Staphylinidae II. W. Junk, Berlin, pp. 989 - 1500.","Schulke, M. & Smetana, A. (2015) Staphylinidae. In: Lobl, I. & Lobl, D. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 2. Hydrophiloidea - Staphylinoidea. Revised and Updated Edition. Brill, Leiden, pp. 304 - 1134.","Champion, G. C. (1898) A list of Cincindelidae, Carabidae and Staphylinidae collected by Mr. J. J. Walker. R. N., F. L. S. in the region of the straits of Gibraltar. The Transactions of the Entomological Society of London, 1, 65 - 103.","Fauvel, A. (1871 a) Faune Gallo-Rhenane ou descriptions des insectes qui habitent la France, la Belgique, la Hollande, le Luxembourg, les provinces Rhenanes et la Valais avec tableaux synoptiques et planches gravees. Bulletin de la Societe Linneenne de Normandie, Series 2, 5 (1869 - 1870), 27 - 192.","Fauvel, A. (1872) Faune Gallo-Rhenane ou species des insectes qui habitent la France, la Belgique, la Hollande, le Luxembourg, la prusse Rhenane, la Nassau et la Valais avec tableaux synoptiques et planches gravees, Caen, 3 (3), 1 - 214."]}

Lesteva (s.str.) aterrima Lohse, 1967 (Figs. 1, 3–5, 8) Lesteva aterrima Lohse, 1967: 10; Zanetti 2008: 994, 996 Type material examined. Paratypes: 1 ♂: ‘ Asturies: Covadonga | (Cangas de Onis) | rio Covadonga | VI.1965 G.Fagel’ , ‘Paratypoid’ , ‘Coll. | G. A. Lohse | MHNG-1994’ , ‘ PARATYPE | Lesteva | aterrima sp. n. | Det. Lohse 1967 ’ , ‘MHNG | ENTO | 00087306’ , ‘ Lesteva (s.str.) | aterrima Lohse, 1967 | Shavrin A.V. det. 2021’ (MHNG); 1 ♀: same labels, but ‘MHNG | ENTO | 00087307’ (MHNG). Material examined. SPAIN: ASTURIAS: 10 ♂, 8 ♀♀: Puerto de La Cubilla, 1400 m a.s.l. 08.06.1991. P. Wunderle leg. (cA, cW); 3 ♂♂, 1 ♀: same data, but L. Zerche leg. (cW, MHNG); 1 ♂, 1 ♀: Oviedo, Picos de Europa, Covadonga Lago Enol. 1600 m a.s.l. 03.06.1990 (cZan); 1 ♂: Covadonga, Hutte Vega Redonga. 1300 m a.s.l. 07.06.1991. L. Zerche leg. (MHNG). Redescription. Measurements (n=25): HW: 0.68–0.75; HL: 0.47–0.52; OL: 0.24–0.25; LT: 0.07–0.08; AL (holotype): 2.07; PL: 0.75–0.78; PWMax: 0.83–0.87; PWMin: 0.60–0.65; EL: 1.58–1.62; ESL: 1.49–1.52; EW: 1.39–1.42; MTbL (holotype): 1.02; MTrL (holotype): 0.57; AW: 1.35–0.38; AedL: 0.67–0.82; FL: 2.85–2.96; TL: 3.50–4.55 (holotype: 4.45). Body dark brown, sometimes with paler apical abdominal tergites; antennes and legs yellow-brown; mouthparts and tarsi yellow. Punctation of head very dense, coarse and moderately deep, interspaces between punctures in middle about as long as diameter of one nearest puncture, finer in vertex and finer and denser around eyes; punctation of neck as that of middle portion of head; punctation of pronotum moderately regular, about as that on middle part of head, but slightly deeper, slightly sparser in mediobasal third; scutellum with indistinct, very fine and sparse punctation; punctation of elytra similar to that on pronotum, but sparser, slightly finer and denser in parascutellar portion and along suture; abdominal tergites with very fine and dense punctation. Head and neck with very dense isodiametric microsculpture, distinctly coarser in infraorbital ridges; pronotum with fine, transverse microreticulation except for mediobasal portion; scutellum with very dense, fine meshes; abdominal tergites with distinct transverse microsculpture. Habitus as in Fig. 1. Head transverse, 1.4 times as broad as long, distinctly and widely elevated in middle; middle portion separated from lateral portions by wide and deep longitudinal depressions; eyes moderately large, convex, about three times as long as temples (lateral view); distance between ocelli markedly shorter than distance between ocellus and posterior margin of eye. Antennae reaching basal third of elytra when reclined, with moderately elongate antennomeres; basal antennomere disitnctly more than twice as long as wide, 2 slightly narrower than basal antennomere, 3 slightly longer than 2, 4 slightly broader and longer than 3, 5–7 slightly longer and broader than 4, 8–10 slightly shorter than 7, apical antennomere slightly broader than 10. Pronotum evenly convex, 1.1 times as broad as long, 1.1–1.2 times as broad as head, very wide in apical third, sharply narrowed toward elongate subparallel latero-basal margins; middle portion of pronotum with indistinct, oval and moderately wide paired impressions and without or with indistinct to distinct and deep, transverse impression in mediobasal third. Elytra somewhat flattened, transverse, longer than broad, slightly less than twice as long as pronotum, significantly widened apicad. Male. Apical margins of abdominal tergite VIII slightly and sternite VIII concave. Aedeagus with very wide basal portion, significantly narrowed toward very narrow median lobe, with small rounded apex; parameres wide, exceeding apex of median lobe, widened apically, with two long apical and preapical setae; lateral sides of median lobe with several moderately long setae; internal sac without visible sclerotized structures (Fig. 3). Lateral aspect of aedeagus as in Figs. 4–5. Female. Apical margins of abdominal tergite VIII and sternite VIII rounded. Comparative notes. Regarding the general shape of the aedeagus and the presence of a deep concavity between the basal portion, median lobe and the narrow, curved ventral process, L. aterrima is similar to L. benicki Lohse, 1958, a species distributed in the Alps, and L. martinae Zanetti, 2008, a species described from central Italy (Zanetti 2008). Lesteva aterrima differs from these species by the smaller size of the aedeagus, a narrower apical part of the median lobe, and different features of its setation (see Fig. 4 in Zanetti (2008)). Based on the general shape of the body, all three species are similar to L. pubescens, from which they are distinguished by the morphology of the aedeagus. Distribution. Lesteva aterrima is known from several localities in Cantabrian mountain range, in north-western Spain (Fig. 8). Natural history. Specimens were collected at elevations from 1300 to 1600 m a.s.l. Some specimens were sifted from wet mosses near streams., Published as part of Shavrin, Alexey V. & Zanetti, Adriano, 2021, Review of the genus Lesteva Latreille, 1797 (Coleoptera: Staphylinidae: Omaliinae Anthophagini) of the Iberian Peninsula, pp. 401-427 in Zootaxa 4966 (4) on pages 403-406, DOI: 10.11646/zootaxa.4966.4.1, http://zenodo.org/record/4738667, {"references":["Lohse, G. A. (1967) Zwei neue westeuropaische Arten der Gattung Lesteva Latr. Entomologische Blatter fur Biologie und Systematik der Kafer, 63 (1), 10 - 12.","Zanetti, A. (2008) Description of Lesteva martinae nov. sp. from Central Italy (Coleoptera, Staphylinidae: Omaliinae). Linzer biologische Beitrage, 40 (1), 993 - 997."]}

Lesteva (s.str.) balearica Lohse, 1967 (Figs. 2, 6–7) Lesteva balearcia Lohse, 1967: 11 Type material examined. Holotype, ♂ [aedeagus glued on the same card under the specimen]: ‘Mallorca | Balearen’ , ‘ Lesteva [handwritten] | balearica n. sp. [handwritten] | det. Dr. G. A. Lohse’ , ‘Holotypus’ , ‘Coll. | G. A. Lohse | MHNG-1994’ , ‘ HOLOTYPE | Lesteva | balearica sp. n. | Det. Lohse 1967 ’ , ‘MHNG | ENTO | 00087308 , ‘ Lesteva (s.str.) | balearica Lohse, 1967 | Shavrin A.V. det. 2021’ (MHNG). Paratypes: 1 ♂ [aedeagus glued on the same card under the specimen]: same labels as the holotype except for: ‘Para- | typoid’ , ‘ PARATYPE | Lesteva | balearica sp. n. | Det. Lohse 1967 ’ , ‘MHNG | ENTO | 00087309’ (MHNG); 1 ♂, 2 ♀♀ [one specimen without head and pronotum]: ‘Mallorca | IV 65 | Benick’ , ‘balearica | Lohse | Paratyp’ , ‘Coll. | G.A. Lohse | MHNG-1994’ , ‘ PARATYPE | Lesteva | balearica sp. n. | Det. Lohse 1967 ’ , ‘MHNG | ENTO | 00087310’ , ‘MHNG | ENTO | 00087311’ , ‘MHNG | ENTO | 00087312’ (MHNG); 3 ♂♂ [dissected, preparation with the aedeagus in Canada balsam on plastic card are pinned under related specimens]: same labels as above, except for: ‘MHNG | ENTO | 00087313’ , ‘MHNG | ENTO | 00087314’ , ‘MHNG | ENTO | 00087315’ (MHNG); 1 ♂ [without right antennomeres 3-11], 1 ♀ [without left antennomeres 3-11]: same labels, but: ‘MHNG | ENTO | 00087316’ , ‘MHNG | ENTO | 00087317’ (MHNG); 1 ♂, 2 ♀♀: ‘Mallorca | IV.65 Benick’ , ‘ Lesteva | balearica Lohse | Paratyp.’ , ‘ PARATYPE | Lesteva | balearica sp. n. | Det. Lohse 1967 ’ , ‘MHNG | ENTO | 00087318’ , ‘MHNG | ENTO | 00087319’ , ‘MHNG | ENTO | 00087320’ (MHNG); 2 ♀♀ (one specimen without left antennomeres 3-11): same labels, but: ‘MHNG | ENTO | 00087321’ , ‘MHNG | ENTO | 00087322’ (MHNG); 2 ♀♀: same data as the previous except for: ‘MHNG | ENTO | 00087323’ , ‘MHNG | ENTO | 00087324’ (MHNG); 1 ♂ [dissected]: ‘Mallorca | Balearen | IV 1965 G. Benick’ , ‘ Lesteva | balearica n. sp. | det. G.A. Lohse’ , ‘Paratypus’ (cZan). All paratypes with additional printed labels: ‘ Lesteva (s.str.) | balearica Lohse, 1967 | Shavrin A.V. det. 2021’. Material examined. SPAIN: Balearic islands: 1 ♀: Mallorca, Gorg Blau, Casa Nova, 39°49‘N 02°50E. 14.05.1978. H. Malicky leg. (cA). Redescription. Measurements (n=15): HW: 0.68–0.77; HL: 0.49–0.52; OL: 0.25–0.27; LT: 0.10; AL (holotype): 2.05; PL: 0.73–0.80; PWMax: 0.81–0.87; PWMin: 0.60–0.67; EL: 1.51–1.62; ESL: 1.37–1.50; EW: 1.35–1.42; MTbL (holotype): 0.97; MTrL (holotype): 0.55; AW: 1.21–1.35; AedL: 0.65–0.85; FL: 2.82–3.06; TL: 3.50–4.60 (holotype: 4.45). Body yellow-brown to reddish-brown, usually with darker head and large, elongated, yellow to yellow-brown spot on apical half of each elytron, and paler apical abdominal segments; mouthparts, antennomeres and legs yellow to yellow-brown, usually with protibia somewhat darker. Punctation of head moderately dense and fine, sparser in middle and between ocelli, and distinctly denser around eyes; punctation of neck very sparse in middle, finer and coarser on lateral portions; punctation of pronotum about as than on middle portion of head, but slightly sparser, particularly in mediobasal third; scutellum with very fine, irregular punctation; punctation of elytra similar to that on pronotum, but sparser; abdominal tergites with regular, distinct and fine punctation, markedly finer on abdominal tergite VII. Latero-apical portions of head between supra-antennal prominces and apical margins of eyes with coarse and very dense sculpture; mediobasal portion of neck with transverse microreticulation, lateral portions of neck with moderately coarse transverse meshes; some specimens with fine, transverse microsculpture in medioapical third; abdominal tergites with distinct isodiametric sculpture. Habitus as in Fig. 2. Head 1.3–1.4 times as broad as long, distinctly elevated in middle; middle portion separated from lateral portions by very deep longitudinal depressions; eyes more than twice as long as temples; distance between ocelli slightly shorter than distance between ocellus and posterior margin of eye. Antennae reaching basal third of elytra when reclined, with moderately elongate antennomeres; basal antennomere more than three times as long as wide, 2 markedly narrower than basal antennomere, 3 slightly longer and narrower than 2, 4 slightly shorter and broader than 3, 5–8 slightly broader and longer than 4, 9 slightly broader than 8, 10 slightly shorter than 9. Pronotum slightly broader than long, 1.1 times as broad as head, from their widest anterior third sharply rounded toward elongate, subparallel latero-basal margins; dorsal surface usually without impressions, but some specimens with very indistinct, long, paired impressions in middle and/or transverse impression in mediobasal third. Elytra about as long as broad, 1.8 times as long as pronotum, slightly widened toward hind angles. Abdominal tergites IV and V with paired, transverse and moderately wide tomentose spots. Male. Apical margin of abdominal tergite VIII rounded or slightly concave. Apical margin of abdominal sternite VIII widely and deeply concave. Aedeagus with wide basal part, narrowed toward middle of median lobe, gradually widened toward apical third, with rounded apex; parameres moderately narrow, slightly widened apically, not reaching apex of median lobe, with four moderately short apical setae; internal sac wide and moderately short (Fig. 6). Lateral aspect of aedeagus as in Fig. 7. Female. Apical margin of abdominal tergite VIII straight or rounded. Apical margin of abdominal sternite VIII rounded. Comparative notes. Based on the size and the general shape of the body and antennomeres, length of temples and features of the punctation of the elytra, L. (s.str.) balearica is similar to European L. (s.str.) pubescens, from which it can be distinguished by the paler coloration of the body and the morphology of the aedeagus, without pubescence on the median lobe. Regarding the general shape of the aedeagus, it is somewhat similar to L. (s.str.) fontinalis and the Corsican L. (s.str.) praeses Fauvel, 1900 (Fig. 98e in Zanetti (1987)), from which it differs by the paler coloration, longer temples and the shape of the apical part of the median lobe. Distribution. Lesteva balearica is an endemic species of Mallorca (Majorca), the largest of the Balearic islands. Natural history. The detailed ecological data are unknown., Published as part of Shavrin, Alexey V. & Zanetti, Adriano, 2021, Review of the genus Lesteva Latreille, 1797 (Coleoptera: Staphylinidae: Omaliinae Anthophagini) of the Iberian Peninsula, pp. 401-427 in Zootaxa 4966 (4) on pages 406-407, DOI: 10.11646/zootaxa.4966.4.1, http://zenodo.org/record/4738667, {"references":["Lohse, G. A. (1967) Zwei neue westeuropaische Arten der Gattung Lesteva Latr. Entomologische Blatter fur Biologie und Systematik der Kafer, 63 (1), 10 - 12.","Zanetti, A. (1987) Coleoptera. Staphylinidae. Omaliinae. In: Fauna d'Italia. Vol. 25. Calderini, Bologna, pp. i - xii + 1 - 472."]}

Lesteva (s.str.) osellai Zanetti & Shavrin, sp.n. (Figs. 8, 29–30, 32–37) Type material examined. Holotype, ♂: ‘ Portugal | Estrela m 1950 |10.vii.[19]79 | Osella’, ‘ HOLOTYPE | Lesteva (s.str.) | osellai sp.n. | Zanetti A. & Shavrin A.V. des. 2020’ (MCSNV). Paratypes: 23 ♂♂ (three specimens dissected): ‘Portugal | Estrela m 1950 | 10.vii.[19]79 Osella’ (2 ♂♂: cZan, 21 ♂♂: MCSNV); 1 ♂ (dissected): ‘Portugal | Estrela 1950 | m 10-VII-79 | Osella’ , ‘ Lesteva [handwritten] | prope [handwritten] | lusitana [handwritten] | det. A. Zanetti 197’ , ‘ Lesteva | lusitanica Lohse | Det. Lohse’ , ‘Coll. | G.A. Lohse | MHNG-1994’ (MHNG); 1 ♂ (dissected; right antennomeres 10-11 missing): ‘Portugal | Estrela 1950m | 10.VII.1979 | Osella’ (cSh); 1 ♂ (aedeagus mounted), 31 ♀♀ (two specimens without head and pronotum): ‘Portugal | Estrela 1950 | m 10.VII.[19]79 | Osella’ (1 ♀: cZan, 30 ♀♀: MCSNV); 2 ♂♂, 1 ♀: ‘Espagne | Picos de | Europa’ , ‘Covadonga | Lago Enol | 1600 m. 3.VI.90’ (1 ♂, 1 ♀: cZan; 1 ♂: cSh); 1 ♀ (pronotum and mediobasal portion of left elytron damaged): ‘Espagne | Picos de Europa | (Oviedo)’ , ‘Covadonga | Lago Enol | 1200 m 2.VI.[19]90’ (cZ); 1 ♀: ‘E Cord.Cantabrica |Pto. de Cubilla | 1400m Spritzmoos | 08.06.[19]91 [P.] Wunderle’ , ‘ Lesteva [handwritten] | det.P.Wunderle’ , ‘?coiffaiti’ (cW); 1 ♂ (aedeagus glued on the same card under the specimen): ‘Pto. de Pajares, | Spain. | G. C. C.’ , ‘coiffaiti [handwritten] | det.Dr.G.A.Lohse 197’ , ‘Coll. | G.A. Lohse | MHNG-1994’ (MHNG); 1 ♀ (without left antennomeres 3-11 and right antenna, without apical part of the abdomen): ‘ SPAIN.Leon, | Pto. de Tarna, | 17km.N.Riano, | 1490m, 1.vii.1965 ’ , ‘Headwaters Rio Nalon.’ , ‘Among stones and | grit: fast stream.’ , ‘J.Balfour-Browne. | B.M. 1965-461.’ , ‘coiffaiti [handwritten] | det.Dr.G.A.Lohse 19’ , ‘Coll. | G.A. Lohse | MHNG-1994’ (MHNG). All paratypes with additional red printed labels: ‘ PARATYPE | Lesteva (s.str.) | osellai sp.n. | Zanetti A. & Shavrin A.V. des. 2020’. Description. Measurements (n=50): HW: 0.67–0.80; HL: 0.52–0.60; OL: 0.18–0.25; LT: 0.10–0.12; AL (mean): 2.38; PL: 0.72–0.85; PWMax: 0.82–0.97; PWMin: 0.60–0.70; EL: 1.50–1.80; ESL: 1.35–1.60; EW: 1.27–1.57; MTbL: 1.02; MTrL: 0.47; AW: 1.22–1.57; AedL: 0.77–1.02; FL: 3.15–3.35; TL: 4.10–5.05 (holotype: 4.10). Body brown, with paler elytra and apical abdominal tergites in some specimens; antennae and legs yellow-brown; mouthparts and tarsi yellow. Punctation of head dense, moderately coarse and deep, sparser in middle, and slightly finer around eyes (some paratypes from Spain with oval impunctate portion between ocelli); punctation of neck as that of head, with impunctate longitudinal band in middle; punctation of pronotum usually coarser and deeper than that of head, sometimes deeper and coarser in medioapical third, finer and sparser in middle and mediobasal third (some paratypes from Estrela with indistinct impunctate longitudinal band); scutellum with several fine punctures; punctation of elytra similar to that on pronotum, but sparser, slightly finer and denser in parascutellar portion and finer and sparser along suture; abdominal tergites with fine, moderately dense, and sometimes indistinct punctation. Body glossy, mostly without microsculpture; basal portion of neck with coarse, dense, isodiametric sculpture, basal portion of scutellum with fine transverse meshes, abdominal tergites with distinct transverse microsculpture on tergites IV–VI and finer, isodiametric microreticulation on tergites VII–VIII. Forebody covered with dense and moderately long semierect setation; abdomen with fine and dense setae. Habitus as in Figs. 29–30. Head 1.2–1.3 times as broad as long, distinctly elevated in middle; middle portion separated from lateral portions by deep longitudinal depression, markedly deeper between middle length of eyes; eyes large and convex, about twice as long as temples (lateral view), gradually narrowing toward neck; ocelli large, distance between ocelli subequal to distance between ocellus and posterior margin of eye. Antennae moderately short, reaching basal portion of elytra when reclined, with slightly thickened and moderately short antennomeres 4–10; basal antennomere about twice as long as wide, 2 about half as wide as basal antennomere, 3 slightly longer than 2, 4 slightly broader and longer than 3, 5–7 slightly longer than 4, 8–10 slightly shorter and broader than 7, apical antennomere about 1.3 times as long as 10, in apical third sharply narrowed toward apex. Pronotum convex, slightly broader than long and head, widest slightly before middle, from their widely rounded lateral margins gradually narrowed toward subparallel latero-basal margins, with obtuse hind angles; mediobasal third of some paratypes with distinct, transverse, moderately deep impression. Elytra distinctly convex, slightly longer than broad, twice as long as pronotum, slightly widened toward hind angles; apical margins widely rounded; mediobasal surface of elytra in some paratypes with indistinct depression. Metatarsus more than twice as long as metatibia, with basal metatarsomere about as long as combined length of tarsomeres 2–3 and about as long as apical tarsomere. Abdomen slightly narrower or about as broad as elytra, with paired tomentose spots in the middle of abdominal tergites IV and V, more transverse in tergite IV, smaller and round in tergite V, with distinct narrow palisade fringe on apical margin of abdominal tergite VII. Male (Fig. 29). Apical margins of abdominal tergite VIII slightly and sternite VIII deeply concave. Aedeagus with wide basal portion, gradually narrowed toward moderately narrow, long median lobe, in apical third sharply narrowed toward subacute apex; parameres narrow, slightly reaching apex of median lobe, with four short apical setae; internal sac moderately wide and long (Figs. 32, 34, 36). Lateral aspect of aedeagus as in Figs. 33, 35, 37. Female (Fig. 30). Apical margins of abdominal tergite VIII and sternite VIII rounded. Comparative notes. Based on the general body shape, coloration, robust antennomeres and moderately sparse and large punctation of the pronotum, L. osellai sp.n. is similar to L. lusitana, L. longoelytrata , and L. monticola. Regarding body size, the shape of the apical margins of the elytra, the length of the apical metatarsomere (not longer than basal metatarsomere), this species is similar to L. lusitana and L. monticola. It differs from them by the shorter antennomeres 8–10, a narrower median lobe, and shorter parameres. Distribution. The species is known from several localities in Serra da Estrela, Portugal, and the Cantabrian mountains, Spain (Fig. 8). Natural history. Specimens were collected at elevations from 1200 to 1950 m a.s.l. Detailed ecological data are unknown. Etymology. Patronymic, the species is named to honour our colleague, G. Osella (Verona, Italy), collector of the type material., Published as part of Shavrin, Alexey V. & Zanetti, Adriano, 2021, Review of the genus Lesteva Latreille, 1797 (Coleoptera: Staphylinidae: Omaliinae Anthophagini) of the Iberian Peninsula, pp. 401-427 in Zootaxa 4966 (4) on pages 419-420, DOI: 10.11646/zootaxa.4966.4.1, http://zenodo.org/record/4738667

Lesteva (s.str.) monticola Kiesenwetter, 1847 Lesteva monticola Kiesenwetter, 1847: 77; Luze 1903: 190, Portevin 1929: 437, Palm 1948: 120, Lohse 1956: 56, 1960: 2, 1964: 60, Tóth 1982: 100 Lesteva (s.str.) monticola: Zanetti 1987: 355, 2012: 69, Tronquet 2006: 33 Lesteva bicolor var. collina Haliday, 1855: 11 Lesteva sharpi Rye, 1866: 124 Lesteva nivicola Fauvel, 1871a: 130 [= 1872: 104] Lesteva pandellei Fauvel, 1871b: 18 Lesteva (Lestevella) pandellei: Jeannel & Jarrige 1949: 317 Lesteva karnoschizkyi Scheerpeltz, 1937: 192 Material examined. SPAIN: CANTABRIA: 1 ♂: Ruberdiago. 11.06.1990. D.A. Lott leg. (BMNH); ASTURIAS: 4 spec: De Pajares (BMNH); MADRID: 1 ♂: Escorial (IRSN); 1 ♀: same locality. 30.07.1879. L. Bleuse leg. (NMPC); 2 ♀♀: Parque Nacional de Guadarrama, San Ildefonso - Majada Hambrienta. 21.06- 08.07.2016. J.L. Lencina & J.C. Otero González leg. (cSh); ARAGÓN: 1 ♀: Estrella (IRSN); ANDALUSÍA: 1 ♂: Granada (IRSN); 1 ex.: ‘Andalucia’ (ZIN); LOCALITY NOT SPECIFIED: 1 ♀: ‘Spanien’, ‘ L. pandellei J. Bohač det. 80’ (NMPC); 1 ♀: ‘Pyrén.’, ‘ fontinalis ’, ‘ L. pandellei J. Bohač det. 80’ (NMPC). Remarks. Lesteva monticola is widespread in Europe and also known from Turkey (Schülke & Smetana 2015). It was recorded from Spain as L. pandellei (Fauvel 1876, Quedenfeldt 1884, Wagner 1927). The record from Algeria (Fauvel 1902) requires confirmation., Published as part of Shavrin, Alexey V. & Zanetti, Adriano, 2021, Review of the genus Lesteva Latreille, 1797 (Coleoptera: Staphylinidae: Omaliinae Anthophagini) of the Iberian Peninsula, pp. 401-427 in Zootaxa 4966 (4) on page 417, DOI: 10.11646/zootaxa.4966.4.1, http://zenodo.org/record/4738667, {"references":["Kiesenwetter, E. A. H. von (1847) [New species]. In: Kiesenwetter, H. von & Markel, F., Eine entomologische Excursion im Riesengebirge. Entomologische Zeitung. Stettin, 8 (3), pp. 73 - 87.","Luze G. (1903) Revision der palaarktischen Arten der Staphyliniden-Gattung Lesteva Latr. Verhandlungen der Kaiserlich- Koniglichen Zoologisch-Botanischen Gesellschaft in Wien, 53, 179 - 197.","Portevin, G. (1929) Histoire naturelle des Coleopteres de France. I. Adephaga. Polyphaga: Staphylinoidea. In: Encyclopedie Entomologique. Vol. 12. P. Lechevalier, Paris, pp. i - xii + 1 - 649.","Palm, T. (1948) Svensk Insekfauna. 9. Skalbaggar. Coleoptera. Kortvingar: Fam. Staphylinidae. Unterfam. Micropeplinae, Phloeocharinae, Olisthaerinae, Proteininae, Omaliinae. Almquist and Wiksells, Uppsala, 133 pp.","Lohse, G. A. (1956) Die mir aus Deutschland bekannt gewordenen Arten der Gattung Lesteva Latr. Entomologische Blatter fur Biologie und Systematik der Kafer, 51 (1 / 2), 51 - 60. [1955]","Lohse, G. A. (1960) Die Arten der Gattung Lesteva Latr. (Coleoptera Staphylinidae) der iberischen Halbinsel. Bulletin Institut Royal des Sciences Naturelles de Belgique, 36 (26), 1 - 7.","Lohse, G. A. (1964) Staphylinidae I. (Micropeplinae bis Tachyporinae). In: Freude, H., Harde, K. & Lohse, G. (Eds.), Die Kafer Mitteleuropas. Band 4. Goecke & Evers, Krefeld, pp. 1 - 264.","Toth, L. (1982) Magyarorszag Allatvilaga - Fauna Hungariae, Holyvak II. Staphylinidae II VII. Akademiai Kiado, Budapest, 110 pp.","Zanetti, A. (1987) Coleoptera. Staphylinidae. Omaliinae. In: Fauna d'Italia. Vol. 25. Calderini, Bologna, pp. i - xii + 1 - 472.","Zanetti, A. (2012) Unterfamilie Omaliinae. In: Assing, V. & Schulke, M. (Eds.), Freude-Harde-Lohse-Klausnitzer - Die Kafer Mitteleuropas. Band 4. Staphylinidae I. Zweite neubearbeitete Auflage. Spektrum Akademischer Verlag, Heidelberg, pp. 49 - 117.","Tronquet, M. (2006) Catalogue iconographique des Coleopteres des Pyrenees-orientales. Vol. 1. Staphylinidae. Revue de l`association roussillonnaise d`entomologie, 15, 1 - 79.","Haliday, A. H. (1855) Entomological remarks. Enumeration of the Irish species of Trichopterygidae. The Natural History Review, 2, 116 - 124, pl. III.","Rye, E. C. (1866) Descriptions of new species, & c., of Brachyelytra. The Entomologist's Monthly Magazine, 3, 121 - 125.","Fauvel, A. (1871 a) Faune Gallo-Rhenane ou descriptions des insectes qui habitent la France, la Belgique, la Hollande, le Luxembourg, les provinces Rhenanes et la Valais avec tableaux synoptiques et planches gravees. Bulletin de la Societe Linneenne de Normandie, Series 2, 5 (1869 - 1870), 27 - 192.","Fauvel, A. (1872) Faune Gallo-Rhenane ou species des insectes qui habitent la France, la Belgique, la Hollande, le Luxembourg, la prusse Rhenane, la Nassau et la Valais avec tableaux synoptiques et planches gravees, Caen, 3 (3), 1 - 214.","Fauvel, A. (1871 b) [No title]. Bulletin de la Societe Linneenne de Normandie, Series 2, 5 (1869 - 1870), 17 - 21.","Jeannel, R. & Jarrige, J. (1949) Biospeologica. LXVIII. Coleopteres staphylinides (Premiere Serie). Archives de Zoologie Experimentale et Generale, 86, 255 - 392.","Scheerpeltz, O. von (1937) Wissenschaftliche Ergebnisse einer von Herrn Hofrat F. Schubert, seinem Sohne Herrn cand. Phil. F. Schubert und Herrn Prof. Ing. K. Mandl im Sommer 1935 (1936) nach Bulgarien unternommenen Studienreise. Coleoptera: I. Staphylinidae. Mitteilungen aus den Koniglichen Naturwissenschaftlichen Instituten, 10, 185 - 246.","Schulke, M. & Smetana, A. (2015) Staphylinidae. In: Lobl, I. & Lobl, D. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 2. Hydrophiloidea - Staphylinoidea. Revised and Updated Edition. Brill, Leiden, pp. 304 - 1134.","Fauvel, A. (1876) Faune Gallo-Rhenane ou descriptions des insectes qui habitent la France, la Belgique, la Hollande, le Luxembourg, les provinces Rhenanes et la Valais avec tableaux synoptiques et planches gravees. Bulletin de la Societe Linneenne de Normandie, Series 2, 10, 26 - 267.","Quedenfeldt, G. (1884) Beitrage zur Kenntniss der Staphylinen-Fauna von Sud-Spanien, Portugal und Marokko. Berliner Entomologische Zeitschrift, 28, 351 - 379. https: // doi. org / 10.1002 / mmnd. 18840280212","Wagner, H. (1927) Eine Sammelreise nach Zentral-Spanien. (I. Fortsetzung). Coleopterologisches Centralblatt, 2 (1), 103 - 120.","Fauvel, A. (1902) Catalogue des Staphylinides de la Barbarie de la Basse-Egypte et des Iles Acores, Maderes, Salvages et Canaries. Revue d'Entomologie, 21, 45 - 189."]}

Lesteva (Lestevina) sicula heeri Fauvel, 1871 Lesteva heeri Fauvel, 1871a: 132 [= 1872: 106]; Portevin 1929: 438, Palm 1948: 119, Lohse 1956: 52 Lesteva sicula: Ganglbauer 1895: 714, Reitter 1909: 184, Luze 1903: 185 Lesteva sicula heeri: Lohse 1960: 2, 1964: 59 Lesteva (Lestevina) heeri: Bordoni 1973: 234 Lesteva (Lestevina) sicula heeri: Tóth 1982: 98, Zanetti 1987: 342, 2012: 69, Tronquet 2006: 33 Material examined. PORTUGAL: ALGARVE: 1 &female;: Serra Monchique E Monchique, 37°19’N 8°31’W. 530 m a.s.l. 10.04.2002. H. Meybohm leg. (cA); ESTREMADURA : 4 &male;&male;, 1 &female;: Lisboa, Serra do Sintra. 24.05.1992. P. Wunderle leg. (cW); SPAIN: PAIS VASCO: 1 &female;: Irun. 10.02.2005. A. Anichtchenko leg. (cSh); 1 &female;: same data. 17- 19.03.2005 (cSh); 1 &male;, 3 &female;&female;: same data. 09.04.2005 (cSh); 1 &male;, 2 &female;&female;: same data. 05.2006 (cSh); 1 &female;: same data. 15.06.2006 (cSh); ARAGÓN : 2 &male;&male;: Barbastro, Ariño de Cregenzán. 09.04.1995. D.A. Lott leg. (BMNH); 1 &female;: Villanueva de Sigena. 09.04.1995. D.A. Lott leg. (BMNH); 1 &male;: Guadalavier. 19.05.1994. D.A. Lott leg. (BMNH); 1 &male;: Estrella (BMNH); CASTILLA Y LÉON : 2 &male;&male;: Sierra de Gredos, 10 km SSW Burgohondo, 40°21’15N 04°49’ 10W. 1250 m a.s.l. 25.03.2007. V. Assing leg. (cA); 2 &male;&male;, 1 &female;: Jarandilla, Caseres. 16.06.1969. A. Comellini leg. (MHNG); 1 &male;: Sierra de la Demanda. 2.8.1971. (MHNG); EXTREMADURA : 2 &male;&male;: Jarandilla. 29.06.1969. A. Comellini leg. (MHNG); CASTILLA LA MANCHA : 1 &male;: Daimiel. 14.08.1969. A. Comellini leg. (MHNG); CATALUñA : 1 &male;: Santa Perpètua de Gaiá. 08.04.1995. D.A. Lott leg. (BMNH); VALENCIA : 1 &male;: El Toro. 950 m a.s.l. 07.04.1995. D.A. Lott leg. (BMNH); ANDALUSÍA : 1 &male;: Sierra Magina, 37.72°N - 3.46°W. 2000 m a.s.l. 03- 04.05.2014. A.V. Shavrin leg. (cSch); 3 &male;&male;, 2 &female;&female;: Sierra Nevada, Capileira, Rio Nante, 36.99°N - 3.35°W. 1550 m a.s.l. 08.05.2014. A.V. Shavrin leg. (cSh); 3 &male;&male;, 4 &female;&female;: Capileira. 1600 m a.s.l. 22.03.1994. V. Assing leg. (cA); 1 &female;: Sierra Nevada, Puerto de la Ragua, 37.09°N - 3.02°W. 2000 m a.s.l. 08- 09.05.2014. A.V. Shavrin leg. (cSch); 3 &male;&male;, 3 &female;&female;: Sierra de Bermeja, Jubrique. 500 m a.s.l. 26.03.1994. V. Assing leg. (cA); 1 &male;, 1 &female;: Sierra Nevada, Guejar Sierra. 1200 m a.s.l. 28.09.1993. P. Wunderle leg. (cW); 1 &female;: San Pedro, Ronda. 14- 21.03.1990. Mattern leg. (cW); 4 &male;&male;: Alpujarra Capileira. 1900 m a.s.l. 22.03.1994. P. Wunderle leg. (cW); 2 &male;&male;, 1 &female;: Málaga, Jubrique. 500 m a.s.l. 26.03.1994. P. Wunderle leg. (cW); 1 &male;, 2 &female;&female;: Sierra de Cazorla. 1400 m a.s.l. 06.10.1993. P. Wunderle leg. (cW); 1 &male;: Esteperona (MHNG); 1 &male;, 1 &female;: Castille Puerto de Bejar. 800-1400 m a.s.l. 05-06.1957. G. Fagel leg. (MHNG); 1 &male;: Puerto de la Ragua, Granada E. 1200 m a.s.l. 17.07.1971. A. Comellini leg. < Lesteva fontinalis infrasp. gustavi Det. Herman 2001> (MHNG). Additional material. FRANCE: 1 &male;: ‘ Htes Pyrénées’ (IRSN); MOROCCO: 1 &male;, 1 &female;: Grand Atlas Mts., Oukaimeden env., 31°13’09.5’’N - 0.7°49’29.7’’W. 2200-2300 m a.s.l. 16- 17.06.2010. A. Anichtchenko leg. (cSh); 2 &male;&male;: Alrene, 55 km S Marrakesh. 5000 ft. 26.iv.1961. P.N. Lawrence leg. (BMNH); LOCALITY NOT SPECIFIED: 1 &male;: ‘ Fauvel’, ‘ Pyrenäen’ (SF); 1 &male;, 1 &female;: ‘ Pyren. ’, ‘ Fauvel’, ‘ heeri ’ (SMNH). Remarks. Based on a study of material from several collections, L. sicula heeri was recorded by Lohse (1960) from the Pyrenees and central Spain. Horion (1963) recorded it from southern Spain (“Sierra Nevada, Kiesenwetter leg. Ca. 1850, 6 Ex. Z.S.M.”). Some studied specimens from southern Spain and Morocco have a paler body, welldeveloped hind wings and relatively wide elytra as in L. sicula sicula Erichson, 1840, a subspecies distributed in Italy and Algeria. Lohse (1960) also noted this morphological similarity for specimens from Guadalupe, South Spain. This is probably morphological variability (from apterous to macropterous forms) of the species at its southern border of the distribution. The aedeagi of both subspecies have similar internal and external structures. A genetic analysis would be required for a clarification of the status and possible synonymy of both taxa. The species is here recorded from Morocco for the first time., Published as part of Shavrin, Alexey V. & Zanetti, Adriano, 2021, Review of the genus Lesteva Latreille, 1797 (Coleoptera: Staphylinidae: Omaliinae Anthophagini) of the Iberian Peninsula, pp. 401-427 in Zootaxa 4966 (4) on page 403, DOI: 10.11646/zootaxa.4966.4.1, http://zenodo.org/record/4738667, {"references":["Fauvel, A. (1871 a) Faune Gallo-Rhenane ou descriptions des insectes qui habitent la France, la Belgique, la Hollande, le Luxembourg, les provinces Rhenanes et la Valais avec tableaux synoptiques et planches gravees. Bulletin de la Societe Linneenne de Normandie, Series 2, 5 (1869 - 1870), 27 - 192.","Fauvel, A. (1872) Faune Gallo-Rhenane ou species des insectes qui habitent la France, la Belgique, la Hollande, le Luxembourg, la prusse Rhenane, la Nassau et la Valais avec tableaux synoptiques et planches gravees, Caen, 3 (3), 1 - 214.","Portevin, G. (1929) Histoire naturelle des Coleopteres de France. I. Adephaga. Polyphaga: Staphylinoidea. In: Encyclopedie Entomologique. Vol. 12. P. Lechevalier, Paris, pp. i - xii + 1 - 649.","Palm, T. (1948) Svensk Insekfauna. 9. Skalbaggar. Coleoptera. Kortvingar: Fam. Staphylinidae. Unterfam. Micropeplinae, Phloeocharinae, Olisthaerinae, Proteininae, Omaliinae. Almquist and Wiksells, Uppsala, 133 pp.","Lohse, G. A. (1956) Die mir aus Deutschland bekannt gewordenen Arten der Gattung Lesteva Latr. Entomologische Blatter fur Biologie und Systematik der Kafer, 51 (1 / 2), 51 - 60. [1955]","Ganglbauer, L. (1895) Die Kafer von Mitteleuropa. 2. Familienreihe Staphylinoidea. Theil I. Staphylinidae, Pselaphidae. Carl Gerold's Sohn, Wien, 881 pp.","Reitter, E. (1909) Fauna Germanica. Die Kafer des Deutschen Reiches. Nach der analytischen Methode bearbeitet. II Band. Schriften des Deutschen Lehrervereins fur Naturkunde 24. K. G. Lutz, Stuttgart, 392 pp., pls. 41 - 80. https: // doi. org / 10.5962 / bhl. title. 9570","Luze G. (1903) Revision der palaarktischen Arten der Staphyliniden-Gattung Lesteva Latr. Verhandlungen der Kaiserlich- Koniglichen Zoologisch-Botanischen Gesellschaft in Wien, 53, 179 - 197.","Lohse, G. A. (1960) Die Arten der Gattung Lesteva Latr. (Coleoptera Staphylinidae) der iberischen Halbinsel. Bulletin Institut Royal des Sciences Naturelles de Belgique, 36 (26), 1 - 7.","Lohse, G. A. (1964) Staphylinidae I. (Micropeplinae bis Tachyporinae). In: Freude, H., Harde, K. & Lohse, G. (Eds.), Die Kafer Mitteleuropas. Band 4. Goecke & Evers, Krefeld, pp. 1 - 264.","Bordoni, A. (1973) Lesteva (Lestevina nov.) sbordonii n. sp. della Campania (Col. Staphylinidae). Redia, 54, 229 - 234.","Toth, L. (1982) Magyarorszag Allatvilaga - Fauna Hungariae, Holyvak II. Staphylinidae II VII. Akademiai Kiado, Budapest, 110 pp.","Zanetti, A. (1987) Coleoptera. Staphylinidae. Omaliinae. In: Fauna d'Italia. Vol. 25. Calderini, Bologna, pp. i - xii + 1 - 472.","Zanetti, A. (2012) Unterfamilie Omaliinae. In: Assing, V. & Schulke, M. (Eds.), Freude-Harde-Lohse-Klausnitzer - Die Kafer Mitteleuropas. Band 4. Staphylinidae I. Zweite neubearbeitete Auflage. Spektrum Akademischer Verlag, Heidelberg, pp. 49 - 117.","Tronquet, M. (2006) Catalogue iconographique des Coleopteres des Pyrenees-orientales. Vol. 1. Staphylinidae. Revue de l`association roussillonnaise d`entomologie, 15, 1 - 79.","Horion, A. D. (1963) Faunistik der Mitteleuropischen Kafer. Band IX: Staphylinidae. 1 Teil. Micropeplinae bis Euaesthetinae. s. n., Uberlingen-Bodensee, 412 pp.","Erichson, W. F. (1840) Genera et species Staphylinorum insectorum coleopterorum familiae. F. H. Morin, Berlin, 954 pp. https: // doi. org / 10.5962 / bhl. title. 59644"]}

Lesteva (s.str.) lusitana Lohse, 1955 (Figs. 8, 11, 18–19) Lesteva lusitana Lohse, 1955b: 513; Lohse 1960: 3 Type material examined. Holotype, &male; [apical part of the aedeagus slightly damaged; aedeagus contained in a small glass vial with a cork pinned under the card with the beetle]: ‘Covilhã | 20.5.[19]12’ , ‘Samml. | K. Daniel’ , ‘Typus [handwritten in blue] | Lesteva | lusitana n.sp. [handwritten in blue] | G. A. Lohse det.’ , ‘ Holotypus [printed] &male; | Lesteva | lusitana Lohse | 1955 | Staatssamml.München [printed]’ , ‘Zool. Staatsslg. | München’ (ZSM). Cotype, 1 &male;: ‘Covilhã | 20.5.[19]12’ , ‘Sammlg. | K. Daniel’ , [red full label], ‘cotypus’ , ‘ Lesteva | lusitana n.sp. [handwritten]’ , ‘G.A. Lohse det.’ , ‘Coll. | G.A. Lohse | MHNG-1994’ , ‘COTYPE | Lesteva | lusitana sp. n. | Det. Lohse 1955’ , ‘MHNG | ENTO | 00087329’ , ‘ Lesteva (s.str.) | lusitana Lohse, 1955 | Shavrin A.V. det. 2021’ (MHNG). Material examined. PORTUGAL: ALTO TÂMEGA: 3 &male;&male;, 2 &female;&female;: Serra do Gêres. 800 m a.s.l. 26.05.1992. P. Wunderle leg. (cW); CASTELO BRANCO: 2 &male;&male;: Sierra de Estrela. H. Franz leg. (NHMW); 1 &male;, 1 &female;: Estrela. 1950 m a.s.l. 10.07.1979. G. Osella leg. (cZan); SPAIN: GALICIA: 1 &male;: Lugo, Valle de Florenza. H. Franz leg. (NHMW); 1 &male;: Sierra de Queixa. 1300-1780 m a.s.l. 27- 28.07.1992. M. Tronquet leg. (cZan); CASTILLA Y LEÓN: 1 &male;: Cordillera Cantabrica, NE Puerto de Ventana, 43°04’N - 6°00’W. 1585 m a.s.l. 13.06.2000. D.W. Wrase leg. (cSch); CANTABRIA: 1 &female;: Reinosa, Campoo Cebernigua, 15 km NW Reinosa. 06.06.1991. P. Wunderle leg. (cW); ASTURIAS: 1 &male;: Picos de Europa, Covadonga, Lago Enol. 1200 m a.s.l. 02.06.1990. (cZan); 1 &female;: same data, Cornion. 2000 m a.s.l. 04.07.1994. T. Assmann leg. (cA). Redescription. Measurements (n=15): HW: 0.67–0.70; HL: 0.45–0.50; OL: 0.20–0.25; LT: 0.08–0.10; AL (holotype): 1.69; PL: 0.62–0.70; PWMax: 0.72–0.80; PWMin: 0.60–0.66; EL: 1.50–1.63; ESL: 1.35–1.55; EW: 1.35–1.40; MTbL (holotype): 0.92; MTrL (holotype): 0.50; AW: 1.18–1.38; AedL: 0.82–0.86; FL: 2.62–3.18; TL: 3.20–4.00 (holotype: 3.90). Body brown to reddish-brown, sometimes with slightly paler elytra; antennae and legs yellow-brown; mouthparts and tarsi yellow to yellow-brown. Punctation of head moderately large, sparser in middle and denser between longitudinal depressions and eyes; punctation of pronotum dense, slightly deeper than that of head. Habitus as in Fig. 11. Head 1.4 times as broad as long, with distinctly elevated middle portion, indistinct transverse impression at level of middle length of eyes and separated from lateral portions by very deep longitudinal depressions; eyes more than twice as long as temples. Antennae reaching one third of elytra when reclined, with slightly thickened antennomeres 7–10; antennomeres 2–7 approximately equal in length, 8–9 slightly shorter than 7, 10 about as long as and slightly broader than 9. Pronotum slightly broader than long, about as wide as head, widest in anterior third portion, sharply narrowed toward subparallel latero-basal margins, with obtuse or subacute hind angles; mediobasal third of pronotum with distinct transverse impression. Elytra about as broad as long, more than twice as long as pronotum. Metatarsus less than twice as long as metatibia, with basal metatarsomere slightly shorter than combined length of tarsomeres 2–3. Abdomen with paired transverse tomentose spots in the middle of abdominal tergite V. Male. Apical margin of abdominal tergite VIII rounded. Apical margin of abdominal sternite VIII widely concaved.Aedeagus with very wide basal portion, strongly narrowed toward acute apex; parameres narrow, widened apically, distinctly exceeding apex of median lobe, with four long apical and preapical setae; internal sac moderately short and narrow (Fig. 18). Lateral aspect of aedeagus as in Fig. 19. Female. Apical margins of abdominal tergites and sternites VIII rounded. Comparative notes. Regarding the coloration, large and deep punctation of the pronotum, the shape of the apical margins of the elytra, and the length of the apical metarsomere, L. lusitana is similar to L. monticola and L. osellai sp.n. Regarding the shape of antennomeres it is more similar to L. monticola, from which it is distinguished by smaller body size and broader pronotum. From both these species, it differs by the shape of narrow median lobe, significantly longer parameres, and the different structure of the internal sac. Distribution. The species is known from several localities in central Portugal and the north-western provinces of Spain (Fig. 8). Natural history. Specimens were collected at elevations from 450 to 1585 m a.s.l. Detailed ecological data are unknown. Remarks. The species is here recorded from Spain for the first time., Published as part of Shavrin, Alexey V. & Zanetti, Adriano, 2021, Review of the genus Lesteva Latreille, 1797 (Coleoptera: Staphylinidae: Omaliinae Anthophagini) of the Iberian Peninsula, pp. 401-427 in Zootaxa 4966 (4) on pages 412-414, DOI: 10.11646/zootaxa.4966.4.1, http://zenodo.org/record/4738667, {"references":["Lohse, G. A. (1955 b) Zwei neue Lesteven aus dem Material der Zoologischen Sammlung des Bayrischen Staates in Munchen (Col., Staph.). Mitteilungen der Munchner Entomologischen Gesellschaft, 44 / 45, 512 - 515.","Lohse, G. A. (1960) Die Arten der Gattung Lesteva Latr. (Coleoptera Staphylinidae) der iberischen Halbinsel. Bulletin Institut Royal des Sciences Naturelles de Belgique, 36 (26), 1 - 7."]}

Lesteva (s.str.) pubescens Mannerheim, 1830 (Figs. 31, 38–46) Lesteva pubescens Mannerheim, 1830; Luze 1903: 196, Johansen 1914: 589, Portevin 1929: 437, Palm 1948: 120, Lohse 1956: 55, 1964: 60, Tóth 1982: 100 Lesteva (Lestevella) pubescens: Jeannel & Jarrige 1949: 315 Lesteva (s.str.) pubescens: Zanetti 1987: 367, 2012: 72, Tronquet 2006: 33 Lesteva pubescens var. subaptera Mulsant and Rey, 1880: 69 Lesteva pubescens reyi Jarrige, 1968 Lesteva fageli Lohse, 1960: 6 syn.n. Type material examined. Paratypes of Lesteva fageli Lohse, 1960: 1 &female;: ‘ Espagne: Estremadura | Guadalupe | (canal scierie) | V—1958, G. Fagel’ , ‘fageli sp. n. [handwritten] | det. Dr. G. A. Lohse’ , ‘Para- | type’ , ‘Coll. | G.A. Lohse | MHNG-1994’ , ‘ PARATYPE | Lesteva | fageli sp. n. | Det. Lohse 1960 ’ , ‘MHNG | ENTO | 00087326’ (MHNG); 1 &male; [dissected]: same labels, but: ‘MHNG | ENTO | 00087327’ (MHNG); 1 &male;: same labels, but: ‘MHNG | ENTO | 00087328’ (MHNG). All paratypes with additional printed label: ‘ Lesteva (str.) pubescens Mannerheim, 1830 | Shavrin A.V. det. 2021’. Material examined. PORTUGAL: CASTELO BRANCO: 3 &male;&male;: Serra de Estrela. H. Franz leg. (NHMW); 1 &male;: Estrela (MHNG); 3 &female;&female;: same data. H. Franz leg. (MHNG); SPAIN: GALICIA: 1 &male;, 1 &female;: Pontevedra Galice. 18- 30.07.1959. H. Coiffait leg. (MHNH); 1 &male;: Lugo, Linares. 1000 m a.s.l. 27.05.1996. W. Starke leg. (cA); ASTURIAS: 1 &male; [identified by G.A. Lohse as fageli], 2 &female;&female;: ‘ Hispania la Granja’ (BMNH); SORIA: 2 &male;&male;: Sierra de Urbión, forest W of Santa Inés. 1700 m a.s.l. 23.07.1996. D. Wrase & J.P. Zaballos leg. (cSch); CANTABRIA: 1 &female;: Espinama, 27.07.1972 (MHNG); 1 &male;, 4 &female;&female;: same locality, Valle de Salvoron. 30.07.1972. (MHNG); 3 &male;&male;: Poblagion (BMNH); CASTILLA Y LEÓN: 1 &female;: Sierra de Gredos. 16.07.1959. H. Coiffait leg. (MHNH); 1 &male;: Sierra de Gredos, Novacepeda. 01.06.1984. (cZan); 3 &female;&female;: Puerto de Béjar. 800-1400 m a.s.l. 05-06.1957. G. Fagel leg. (MHNG); ARAGÓN: 1 &male;: Teruel (IRSN); CATALUñA: 1 &male;: Col de Aras (Span. Pyren.). 31.05.1971. G. Lohse leg. (MHNG); 1 &male;, 4 &female;&female;: Barcelona (MHNG); 1 &male;: Sierra de Montseny, Tordera River, 5 km E San-Bernat. 02- 12.5.1981. P. Zwick leg. (MHNG); 4 &male;, 6 &female;&female;: Montseny (MHNG); MADRID: 1 &female;: Escorial (NMPC); 1 &male;: same locality. 30.07.1879. L. Bleuse leg. (NMPC); 1 &male;: ‘ Guadarrama | Eppelsh. ’, ‘ L. pandellei J. Boha&ccaron; det. 1980’ (NMPC); 2 &male;&male;, 2 &female;&female;: Sistema Central Mts., Pinilla de Valle, 40°91’N -3°82’W. 28.04.2014. A.V. Shavrin & A. Anichtchenko leg. (cSh); 2 &female;&female;: Puerto de la Morcuera. 11.10.2004. de la Rosa leg. (cSh); 1 &male;: Sierra de Estrela, Manteigas. 1000-1100 m a.s.l. H. Franz leg. (MHNH); CASTILLA LA MANCHA: 2 &male;&male;: Sierra de Alcaraz, 15 km NNE Riópar, 38°34’01N 02°20’03W 1120 m a.s.l. 10.04.2003. P. Wunderle leg. (cW); VALENCIA: 1 ex.: ‘ Buñol Valencia Moróder’ (MHNH); ANDALUSÍA: 5 &male;&male;, 7 &female;&female;: Despenaperros, Arroyo de los Arcos, 38°38‘N - 3°51‘W. 01- 02.05.2014. A.V. Shavrin leg. (cSh); 1 &male;: ‘ Pnte. Las Herrerias S. Cazorla (Jaén) Mateau-Cobos coll.’ (MHNH); 1 &female;: ‘ Fte. Bermejo. Jaén’, ‘ruela S. Cazorla Mateau-Cobos coll.’ (MHNH); 1 &male;: Sierra Nevada, Capileira. 1600 m a.s.l. 22.03.1994. V. Assing leg. (cSh); 9 &male;&male;, 4 &female;&female;: same data, but P. Wunderle leg. (cW, cSh); 1&male;: Mt. Mulhacen. 2000 m. a.s.l. 21.05.1967. M.E. Bacchus & B. Levey leg. (MHNG). LOCALITY NOT SPECIFIED: 1 &male;: ‘ Miguelon’ (MHNH). Additional material. ANDORRA: 1 &male;: Puerto de Envalira. 2200-2300 m a.s.l. 30.05.1994. M. Schülke & Grünberg leg. (cSch); FRANCE: 1 &male;: ‘ Gavarnie 25.5.[19]31’ (MHNH); 2 &male;&male;: ‘ Gavarnie, H. Pyr. 12-31, vii. 14. G.C.C. ’ (BMNH); 2 &male;&male;: ‘ Lourdes’ (IRSN); 8 &male;&male;, 7 &female;&female;: Por d`Boucharo. 30.06.1976. (MHNG); 14 &male;&male;, 6 &female;&female;: Ht. Canigou. 02.07.1976. (MHNG); 4 &male;&male;, 3 &female;&female;: Sahorre d. Vernet. 04.07.1976. (MHNG); 1&female;: Ariège, Auzat. 19.07.1972 (MHNG); 1 ex.: ‘ Pyr. Or. La Vall. H. Coiffait 24.3.[19]48’ (MHNH); 1 &male;, 1 &female;, 1 ex.: Banyuls-sur-Mer (IRSN, MHNH); 1 &male;: Gard Fressac. 28.10.1982. I. Löbl leg. (MHNG); 8 &male;&male;, 2 &female;&female;: ‘ Col de Aras Pyren. or.’ (MHNG). Remarks. Lesteva fageli was originally described based on several type specimens from Cantabria (“Puente del Cerezo”) and central Spain (“Rio Guadalupejo”). Lohse compared it with L. fontinalis and L. pubescens. He distinguished it from the latter by the darker body and smaller aedeagus (“0,64 mm gegenüber 0,75 mm bei pubescens ”). During the study of types of L. fageli, we found that these specimens are morphologically conspecific with L. pubescens, a variable species regarding coloration, body size, and the proportions of the forebody and aedeagus (Figs. 38–46). Thus, we synonymize L. fageli with L. pubescens. Regarding the presence of stout pubescence on the median lobe, this species is similar to L. pourtoyi Jarrige, 1972, a species originally described from “Cirque de Mahourat (Pyr[énées].-Atl[antiues].)”, but the aedeagus of this species is evidently narrower (Fig. 5 in Jarrige (1972)) and is furnished with longer pubescence. According to Tronquet (2014), L. pourtoyi is also known from Spain (“ pourtoyi Jarrige, 1972: 158. - Pyr. occ. et cent. Esp. (chaîne Cantabrique”)), but this record needs confirmation. Habitus of L. pubescens as in Fig. 31; aedeagus as in Figs. 38–46. Lesteva pubescens is widely distributed in Europe (Schülke & Smetana 2015) and had already been reported from Spain (Lohse 1960). It is here recorded from Portugal for the first time., Published as part of Shavrin, Alexey V. & Zanetti, Adriano, 2021, Review of the genus Lesteva Latreille, 1797 (Coleoptera: Staphylinidae: Omaliinae Anthophagini) of the Iberian Peninsula, pp. 401-427 in Zootaxa 4966 (4) on pages 420-421, DOI: 10.11646/zootaxa.4966.4.1, http://zenodo.org/record/4738667, {"references":["Mannerheim, C. G. von (1830) Precis d'un nouvel arrangement de la famille des brachelytres de l'ordre des insectes coleopteres. s. n., St. Petersbourg, 87 pp","Luze G. (1903) Revision der palaarktischen Arten der Staphyliniden-Gattung Lesteva Latr. Verhandlungen der Kaiserlich- Koniglichen Zoologisch-Botanischen Gesellschaft in Wien, 53, 179 - 197.","Johansen, J. P. (1914) Danmarks rovbiller eller Billefam. Staphylinidae's danske Slaegter og Ar Territory. Bianco Lunos Bogtrykkeri, Kobenhavn, 660 pp.","Portevin, G. (1929) Histoire naturelle des Coleopteres de France. I. Adephaga. Polyphaga: Staphylinoidea. In: Encyclopedie Entomologique. Vol. 12. P. Lechevalier, Paris, pp. i - xii + 1 - 649.","Palm, T. (1948) Svensk Insekfauna. 9. Skalbaggar. Coleoptera. Kortvingar: Fam. Staphylinidae. Unterfam. Micropeplinae, Phloeocharinae, Olisthaerinae, Proteininae, Omaliinae. Almquist and Wiksells, Uppsala, 133 pp.","Lohse, G. A. (1956) Die mir aus Deutschland bekannt gewordenen Arten der Gattung Lesteva Latr. Entomologische Blatter fur Biologie und Systematik der Kafer, 51 (1 / 2), 51 - 60. [1955]","Lohse, G. A. (1964) Staphylinidae I. (Micropeplinae bis Tachyporinae). In: Freude, H., Harde, K. & Lohse, G. (Eds.), Die Kafer Mitteleuropas. Band 4. Goecke & Evers, Krefeld, pp. 1 - 264.","Toth, L. (1982) Magyarorszag Allatvilaga - Fauna Hungariae, Holyvak II. Staphylinidae II VII. Akademiai Kiado, Budapest, 110 pp.","Jeannel, R. & Jarrige, J. (1949) Biospeologica. LXVIII. Coleopteres staphylinides (Premiere Serie). Archives de Zoologie Experimentale et Generale, 86, 255 - 392.","Zanetti, A. (1987) Coleoptera. Staphylinidae. Omaliinae. In: Fauna d'Italia. Vol. 25. Calderini, Bologna, pp. i - xii + 1 - 472.","Zanetti, A. (2012) Unterfamilie Omaliinae. In: Assing, V. & Schulke, M. (Eds.), Freude-Harde-Lohse-Klausnitzer - Die Kafer Mitteleuropas. Band 4. Staphylinidae I. Zweite neubearbeitete Auflage. Spektrum Akademischer Verlag, Heidelberg, pp. 49 - 117.","Tronquet, M. (2006) Catalogue iconographique des Coleopteres des Pyrenees-orientales. Vol. 1. Staphylinidae. Revue de l`association roussillonnaise d`entomologie, 15, 1 - 79.","Mulsant, E. & Rey, C. (1880) Tribu des brevipennes. Onzieme famille: Omaliens. Douxieme famille: Pholidiens. Annales de la Societe Linneenne de Lyon, Series 2, 27, 1 - 430. https: // doi. org / 10.3406 / linly. 1880.3986","Jarrige, J. (1968) Sur quelques brachelytres des Alpes meridionales. L'Entomologiste, 24, 73 - 79.","Lohse, G. A. (1960) Die Arten der Gattung Lesteva Latr. (Coleoptera Staphylinidae) der iberischen Halbinsel. Bulletin Institut Royal des Sciences Naturelles de Belgique, 36 (26), 1 - 7.","Jarrige, J. (1972) Nouveaux coleopteres pyreneens. L'Entomologiste, 28, 155 - 159.","Tronquet, M. (2014) Staphylinidae (excl. Scydmaeninae, Pselaphinae, Scaphidiinae, Euaesthetinae, Leptotyphlinae). In: Tronquet, M. (Ed.), Catalogue des Coleopteres de France. Association Roussillonaise d'Entomologie, Perpignan, pp. 230 - 373","Schulke, M. & Smetana, A. (2015) Staphylinidae. In: Lobl, I. & Lobl, D. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 2. Hydrophiloidea - Staphylinoidea. Revised and Updated Edition. Brill, Leiden, pp. 304 - 1134."]}

Key to species of Lesteva of Portugal and Spain 1 Lateral sides of pronotum with deep impression. Forebody with very coarse punctation. Parameres wide (Fig. 88g in Zanetti (1987)). Body yellow-brown to brown. Body length: 3.20–4.00 mm.................................. L. sicula heeri - Lateral sides of pronotum without impression. Forebody with fine or moderately coarse punctation. Parameres narrow. Body usually darker....................................................................................... 2 2 Antennomeres 6–10 moderately short and wide, antennomere 10 slightly longer than broad.......................... 3 - Antennomeres 6–10 long and slender, antennomere 10 about or more than twice as long as broad..................... 6 3 Apical margins of elytra straight. Apical segment of metatarsus longer than basal tarsomere. Median lobe with longitudinal ridge (Fig. 88p in Zanetti (1987)). Body length: 3.30–4.70 mm......................... L. longoelytrata longoelytrata - Apical margins of elytra widely rounded.Apical segment of metatarsus about as long as or shorter than basal tarsomere. Median lobe without longitudinal keel........................................................................... 4 4 Median lobe narrow, parameres moderately short (Figs. 32, 34, 36). Body brown. Body length: 4.10–5.05 mm. Habitus as in Figs. 29–30.................................................................................. L. osellai - Median lobe different, parameres long. Body brown to reddish-brown........................................... 5 5 Pronotum wider, sharply narrowed posteriad. Apical portion of median lobe narrow, parameres very long, significantly exceeding apex of median lobe (Fig. 18). Body smaller: 3.20–4.00 mm. Habitus as in Fig. 11................ L. lusitana - Pronotum narrower, gradually narrowed posteriad.Apical portion of median lobe moderately wide, parameres slightly exceeding beyond apex of median lobe (Fig. 90a in Zanetti (1987)). Body larger: 4.00– 4.80 mm..................... L. monticola 6 Temples distinctly longer than eyes. Aedeagus as in Figs. 23, 25, 27. Body length: 3.00– 4.80 mm. Habitus as in Fig. 13................................................................................................ L. mateui - Temples about as long as or shorter than eyes............................................................... 7 7 Elytra significantly widened posteriad.................................................................... 8 - Elytra slightly widened posteriad........................................................................ 9 8 Elytra broader than long. Median lobe moderately wide; parameres narrow (Fig. 20, 22). Body length: 3.75–4.65 mm. Habitus as in Fig. 12.................................................................................. L. mariei - Elytra longer than broad. Median lobe very narrow; parameres wide (Fig. 3). Body length: 3.50–4.55 mm. Habitus as in Fig. 1......................................................................................... L. aterrima 9 Punctation of elytra fine and superficial. Aedeagus as in Fig. 16 or Fig. 92&scy; in Zanetti (1987). Body length: 3.60–4.60 mm. Habitus as in Fig. 10.......................................................................... L. hanseni - Punctation of elytra moderately coarse and deep........................................................... 10 10 Body yellow-brown to reddish-brown, usually with large yellowish spots on elytra. Median lobe, gradually widened toward apical third, with rounded apex (Fig. 6). Body length: 3.50–4.60 mm. Habitus as in Fig. 2................... L. balearica - Body darker, without spots on elytra. Median lobe different.................................................. 11 11 Median lobe pubescent; apical part of median lobe moderately narrow, strongly narrowed toward apex (Figs. 38, 41, 43, 45). Body length: 3.50–4.10 mm. Habitus as in Fig. 31................................................. L. pubescens - Median lobe without pubescence; apical part of median lobe wide, abruptly narrowing toward apex (Fig. 14). Body darker, without spots on elytra. Body length: 3.50–4.55 mm. Habitus as in Fig. 9............................... L. fontinalis, Published as part of Shavrin, Alexey V. & Zanetti, Adriano, 2021, Review of the genus Lesteva Latreille, 1797 (Coleoptera: Staphylinidae: Omaliinae Anthophagini) of the Iberian Peninsula, pp. 401-427 in Zootaxa 4966 (4) on page 424, DOI: 10.11646/zootaxa.4966.4.1, http://zenodo.org/record/4738667, {"references":["Zanetti, A. (1987) Coleoptera. Staphylinidae. Omaliinae. In: Fauna d'Italia. Vol. 25. Calderini, Bologna, pp. i - xii + 1 - 472."]}

Lesteva (s.str.) mateui Jarrige, 1954 (Figs. 8, 13, 23–28) Lesteva mateui Jarrige, 1954: 77 Lesteva cobosi Coiffait, 1970: 110 Lesteva cobosi: Outerelo 1980: 54, Lohse 1987: 135, Gamarra & Outerelo 2013: 355 Lesteva cazorlana Lohse, 1987: 135 syn.n. Type material examined. Paratypes of Lesteva mateui Jarrige, 1954, 2 &female;&female;: ‘ Corral del Veleta Sierra Nevada 3200 m J. Mateu’ , ‘PARATYPE’ (MHNH). Holotype of Lesteva cobosi Jarrige, 1954, &male; [The aedeagus is glued on the separate card]: ‘ &male; ’ , ‘F[uen]te. Bermejo. Jaén Iruela S [ierra]. Cazorla Mateau-Cobos coll.’ , ‘ Holotype’ , ‘ Lesteva (s.str.) cobosi Coiff. H. Coiffait det 1970’ (MHNH). Paratype, 1 &female;: same data as the holotype except for red printed label ‘Paratype’ (MHNH). Holotype of Lesteva cazorlana Lohse, 1987, &male; [without right antennomeres 6-11]: ‘ ESPAGNE JAEN | Sierra de Cazorla | Guadalquivir | (source) 25-VII-71 | A. SENGLET’ , ‘Holotypus’ , ‘ Lesteva | cazorlana | n.sp.’ , ‘Coll. | G.A. Lohse | MHNG-1994’ , ‘ HOLOTYPE | Lesteva | cazorlana sp. n. | Det. Lohse 1987 ’ , ‘MHNG | ENTO | 00087325’ , ‘ Lesteva (s.str.) | mateui Jarrige, 1954 | Shavrin A.V. det. 2021’ (MHNG) Material examined. SPAIN: ANDALUSÍA: 1 &male;: Albacete, Mundo River, Chorros del Rio Mundo. 01.06.2002. I. Ribeira & A. Cieslak leg. (cA); 1 &male;, 1 &female;: Jaén, Iznatoraf, Sima los Tejos. 30.10.2005. (cA); 3 &male;&male;: Cazorla Barranco de Teatinos. 1300 m a.s.l. 25.06.1975. (cZan); 5 &male;&male;, 2 &female;&female;: Sierra de Cazorla. 1400 m a.s.l. 06.10.1993. P. Wunderle leg. (cA, cW, cSh); 2 &male;, 5 &female;&female;: same data. 20.03.1974 (MHNG); 3 &male;&male;, 1 &female;: same Mts., Torre del Vinagre. 14.05.1975. H. Fülscher leg. (MHNG); 2 &male;&male;, 1 &female;: Guadalquivir (source), 25.07.1971. A. Senglet leg. (MHNG); 9 &male;&male;, 4 &female;&female;: Sierra Nevada, Capileira, Rio Nante, 36°99’N - 3°35’W. 1550 m a.s.l. 08- 09.05.2014. A.V. Shavrin leg. (cSh); 1 &male;, 4 &female;&female;: same locality. 1600 m a.s.l. 22.03.1994. V. Assing leg. (cA); 1 &male;: same data. 1900 m a.s.l. 23.03.1994. (cA); 6 &male;&male;, 1 &female;: same data. 1500 m a.s.l. 28.09.1993. P. Wunderle leg. (cA, cW, MHNG); 1 &female;: Sierra Nevada, Laguna de las Yeguas. 2900 m a.s.l. 18.05.2005. A. Anichtchenko leg. (cSh); 1 &female;, 6 exs.: same locality. 17.08.1966 (MHNH); 2 &male;&male;, 2 &female;&female;: Bano San Juan. 2900 m a.s.l. 18.08.1966. G. Ledoux leg. (MHNH); 1 &male;, 2 &female;&female;: same data. 2600 m a.s.l. 08.1967. G. Ledoux leg. (MHNH); 1 &male;: ‘ Sierra Nevada 17.7.[18]79. D.S. ’, ‘ Sharp Coll. 1905-303.’ (MHNG); 1 &male;, 1 &female;: ‘ Sa Nevada Hisp. VI.50 G.P. ’ (MHNH); 2 &male;&male;: Güejar-Sierra, Loma de la Cuna de los Cuartos. 1900-2100 m a.s.l. 14.06.1991 D. Wrase leg. (cZan); 1 &male;: Málaga, Las Navas de San Luis. M. Cobos leg. (MHNH); LOCALITY NOT SPECIFIED: 1 &male; [identified by G.A. Lohse]: ‘ Spain. ’ (BMNH). Redescription. Measurements (n=15): HW: 0.70–0.92; HL: 0.50–0.62; OL: 0.20–0.26; LT: 0.11–0.15; AL (mean): 2.30; PL: 0.63–0.85; PWMax: 0.84–0.96; PWMin: 0.59–0.70; EL: 1.38–1.65; ESL: 1.11–1.45; EW: 1.23– 1.53; MTbL (averaged): 1.00; MTrL (averaged): 0.60; AW: 1.14–1.44; AedL: 0.67–0.75; FL: 2.87–3.05; TL: 3.00– 4.80 (holotype of L. mateui: 4.20). Body yellow-brown to brown, with slightly paler elytra; antennae and legs yellow to yellow-brown, with darker apical portion of femora and sometimes tibia; mouthparts and tarsi yellow. Punctation of head dense and fine, slightly larger in middle and denser between longitudinal depressions and eyes, distance between punctures in middle about as broad as nearest puncture; neck with punctation similar to that of middle portion of head; punctation of pronotum denser and sometimes finer than that of head, slightly sparser in medioapical and mediobasal portions; scutellum with very fine and moderately dense, irregular punctation; punctation of elytra distinctly sparser and coarser than that of pronotum. Habitus as in Fig. 13. Head with markedly convex middle portion and very deep longitudinal depressions; eyes small, about twice as long as temples; distance between ocelli markedly shorter than distance between ocellus and posterior margin of eye. Antennae reaching anterior third or middle length of elytra when reclined, with long antennomeres 4–10; antennomeres 3–7 of approximately equal length, 8–10 slightly shorter and broader than 7. Pronotum 1.1–1.3 times as broad as long, markedly broader than head, usually very wide in anterior portion. Elytra distinctly broader than long, about twice as long as pronotum, significantly widened apicad. Basal metatarsomere slightly longer than combined length of tarsomeres 2–3 and distinctly longer than apical metatarsomere. Male. Apical margin of abdominal tergite VIII convex. Apical margin of abdominal tergite VIII concave. Aedeagus with moderately wide basal portion, gradually narrowing toward median lobe and slightly widened in apical third, with sharply narrowed apical portion and subacute apex; parameres nearly reaching apex of median lobe; internal sac long and moderately wide (Figs. 23, 25, 27). Lateral aspect of aedeagus as in Figs. 24, 26, 28. Female. Apical margins of abdominal tergite VIII and sternite VIII round. Comparative notes. Regarding body, size, coloration, the punctation of the forebody, the proportions of the antennomeres, and the shape of the elytra, L. mateui is similar to L. mariei (see above), from which it is distinguished by slightly longer elytra, a narrower median lobe with a differently shaped apical portion, the length of the parameres, and a different structure of the internal sac. Distribution. The species is distributed in South Spain (Fig. 8). Natural history. Specimens were collected at elevations from 1300 to 2900 m a.s.l. The specimens from Rio Nante (Sierra Nevada) were taken from wet mosses near the confluence of a stream and a river. Remarks. Lesteva mateui was originally described from several localities of Sierra Nevada. Lesteva cobosi was described from Sierra de Cazorla (Coiffait 1970) and later synonymized with L. mateui by Lohse (1987). In the same article, Lohse presented a new species without description: L. cazorlana Lohse, 1987, referring to specimens with somewhat larger eyes and a longer median lobe, which had been identified by Coiffait as L. mateui and illustrated by him in Figs. 3 g-i (Coiffait 1970). Lohse designated the specimen from his collection (see above) as the holotype of L. cazorlana. A study of material of this morphologically variable species revealed no significant differences between specimens from different localities. The proportions of the longitudinal length of the eyes in relation to the length of the temples, the width of the pronotum, and the apical portion of the median lobe may vary even in material from the same locality (Figs. 23, 25). Thus, L. cazorlana is synonymized with L. mateui., Published as part of Shavrin, Alexey V. & Zanetti, Adriano, 2021, Review of the genus Lesteva Latreille, 1797 (Coleoptera: Staphylinidae: Omaliinae Anthophagini) of the Iberian Peninsula, pp. 401-427 in Zootaxa 4966 (4) on pages 416-417, DOI: 10.11646/zootaxa.4966.4.1, http://zenodo.org/record/4738667, {"references":["Jarrige, J. (1954) Coleopteres de la Sierra Nevada. \" Staphylinidae \". Archivos del Instituto de Alimatacion, 2, 73 - 79.","Coiffait, H. (1970) Staphylinides nouveaux ou mal connus de region Palearctique occidentale. Bulletin de la Societe d'Histoire Naturelle de Toulouse, 106, 99 - 111.","Outerelo, R. (1980) Los Staphylinoidea de la Sierra de Cazorla. In: Viedma, M. (Ed.), Fauna de Cazorla. Invertebrados. Monografias. Instituto Nacional para la Conservacion de la Naturaleza, 23, 53 - 71.","Lohse, G. A. (1987) Staphyliniden-Studien. Entomologische Blatter fur Biologie und Systematik der Kafer, 83, 135 - 140.","Gamarra, P. & Outerelo, R. (2013) Los Estafilinidos (Coleoptera: Staphylinidae). In: Ruano, F., Tierno de Figueroa, M. & Tinaut, M. (Eds.), Los Insectos de Sierra Nevada. 200 anos de Historia. Vol. I. GRX servicios graficos, Granada, 543 pp."]}

Lesteva (s.str.) hanseni Lohse, 1953 (Figs. 10, 16–17) Lesteva hanseni Lohse 1953: 588; Lohse 1955: 507, 1956: 55, 1960: 2, 1964: 59 Lesteva (s.str.) hanseni: Zanetti 1987: 363, 2012: 72, Tronquet 2006: 33 Material examined. PORTUGAL: CASTELO BRANCO: 1 &male;: Serra da Estrela, S Manteigas, 40°21‘20N 7°33‘ 39W. 1073 m a.s.l., bushes, under stones. 18.03.2002. A. Lompe leg. (cA); SPAIN: ARAGÓN: 1 &male;: Pirineos, Monte Perdido, Collata Añisolo. 2400 m a.s.l. 25.08.2005. A. Anichtchenko leg. (cSh); CATALUñA: 1 &male;, 2 &female;&female;: Seo de Urgel (Arfa), arr. de la Coma. 05-06.1962. G. Fagel leg. (MHNG); ANDALUSÍA: 1 &male;: ‘ Jaen (Espagne)’ (IRSN); 9 exs.: Jaen-Vaidepenas Rd., 18 km S Jaén. 1000 m a.s.l. 25.05.1967 (BMNH); 2 &female;&female;: ‘ Jaen (Sierra)’, ‘ Sammlung Cl- Müller’, ‘ hanseni sp.n. Lohse det.’ (ZSM); 1 &male;: Sierra de Cazorla, Torre del Vinagre. 14.05.1975. H. Fülscher leg. (MHNG); 1 &male;: Granada, Sierra de la Sagra, rio Bravate. 1100 m a.s.l. 30.04.1991. M. Tronquet leg. (cZan); 2 &female;&female;: Ronda, Puente de la Ventilla. 8- 20.05.1956. G. Fagel leg. (MHNG); 4 &male;&male;, 1 &female;: ‘ Ronda Dr. Cameron’ (BMNH); 1 &male;: Tarifa. 04.1992. P. Poot leg. (cW); 1 &female;: ‘ Andalusien’, ‘ Sammlung M. v. Pfaundler’ (ZSM); 1 &female;: ‘ Prov. Andalus. 20.03.1974 ’ (MHNG); LOCALITY NOT SPECIFIED: 1 &female;: ‘ Hispan. 6 Sz. ’, ‘ Samml. v.Seidlitz’, ‘ hanseni Lohse Lohse det.’ (ZSM); 1 &male;: ‘ Hispan. ’ (MHNG). Additional material. FRANCE: 1 &male;: Carnus, Eaux Bonnes. 14.06.1991. P. Wunderle leg. (cW); 1 &female;: ‘ Col de Aras Pyren. or.’ (MHNG); MOROCCO: 7 exs.: 11 km SE Asni, 50 km S Marrakesh 22.iv.1961. P.N. Lawrence (BMNH); 1 &male;: Tizi-n-Test Pass. 2100 m a.s.l. 10.04.1989. (ZMUC); 1 &male;: Haut Atlas, NE Tizi-n-Test, 30°54’12N 08°18’ 39W. 1710 m a.s.l., edge of stream. 26.12.2002. V. Assing leg. (cA); LOCALITY NOT SPECIFIED: 1 &male;: ‘ Atlas […illegible...]’ (BMNH). Remarks. Lesteva hanseni is widely distributed in Europe (Schülke & Smetana 2015). It was recorded from Morocco (Coiffait 1973) and southern Spain (Lohse 1960). Habitus as in Fig. 10. The shape of the apical portion of the median lobe is somewhat variable (Fig. 16 and Fig. 92c in Zanetti (1987)). Lateral aspect of aedeagus as in Fig. 17. The species is here recorded from Portugal for the first time., Published as part of Shavrin, Alexey V. & Zanetti, Adriano, 2021, Review of the genus Lesteva Latreille, 1797 (Coleoptera: Staphylinidae: Omaliinae Anthophagini) of the Iberian Peninsula, pp. 401-427 in Zootaxa 4966 (4) on pages 410-411, DOI: 10.11646/zootaxa.4966.4.1, http://zenodo.org/record/4738667, {"references":["Lohse, G. A. (1953) Eine neue Lesteva des nordlichen Mitteleuropa. Entomologiske Meddelelser, 26 (7), 587 - 590.","Zanetti, A. (1987) Coleoptera. Staphylinidae. Omaliinae. In: Fauna d'Italia. Vol. 25. Calderini, Bologna, pp. i - xii + 1 - 472.","Zanetti, A. (2012) Unterfamilie Omaliinae. In: Assing, V. & Schulke, M. (Eds.), Freude-Harde-Lohse-Klausnitzer - Die Kafer Mitteleuropas. Band 4. Staphylinidae I. Zweite neubearbeitete Auflage. Spektrum Akademischer Verlag, Heidelberg, pp. 49 - 117.","Tronquet, M. (2006) Catalogue iconographique des Coleopteres des Pyrenees-orientales. Vol. 1. Staphylinidae. Revue de l`association roussillonnaise d`entomologie, 15, 1 - 79.","Schulke, M. & Smetana, A. (2015) Staphylinidae. In: Lobl, I. & Lobl, D. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 2. Hydrophiloidea - Staphylinoidea. Revised and Updated Edition. Brill, Leiden, pp. 304 - 1134.","Coiffait, H. (1973) Staphylinides nouveaux ou mal connus du Maroc. Bulletin de la Societe des Sciences Naturelles et Physiques du Maroc, 53, 269 - 292.","Lohse, G. A. (1960) Die Arten der Gattung Lesteva Latr. (Coleoptera Staphylinidae) der iberischen Halbinsel. Bulletin Institut Royal des Sciences Naturelles de Belgique, 36 (26), 1 - 7."]}

Lesteva (s.str.) longoelytrata longoelytrata (Goeze, 1777) Staphylinus longo-elytratus Goeze, 1777: 729 Lesteva longelytrata: Luze 1903: 189, Johansen 1914: 589, Portevin 1929: 437, Palm 1948: 119, Jeannel & Jarrige 1949: 320, Lohse 1956: 56, 1960: 2, 1964: 59, Tóth 1982: 99 Lesteva (s.str.) longelytrata: Bordoni 1973: 234 Lesteva (s.str.) longoelytrata longoelytrata: Zanetti 1987: 353, 2012: 69, Tronquet 2006: 33 Lesteva (s.str.) longoelytrata: Shavrin 2014: 130 Staphylinus macroclytron Geoffroy, 1785: 164 Staphylinus bicolor Paykull, 1789: 21 Carabus dimidiatus Panzer, 1795: 367 Staphylinus multipunctatus Block, 1799: 117 Staphylinus obscurus Paykull, 1800: 388 Carabus staphylinoides Marsham, 1802: 464 Lesteva punctulata Latreille, 1804: 369 Anthophagus intermedius Gravenhorst, 1806: 221 Lesteva longula Mannerheim, 1830: 57 Lesteva impressa Stephens, 1834: 363 Lesteva nigripes Stephens, 1834: 363 Lesteva planipennis Stephens, 1834: 363 Lesteva rufitarsis Stephens, 1834: 362 Lesteva oblonga Motschulsky, 1858: 493 Lesteva major Mulsant & Rey, 1870: 117 [= 1871: 93] Lesteva longelytrata var. dorsalis Reitter, 1909: 184 Lesteva soror Smetana, 1967: 300 Material examined. SPAIN: PAIS VASCO: 1 &male;, 1 &female;: Torre. 2000 m a.s.l.. 15.11.1997. P. Poot leg. (cW); ARAGÓN: 1 &male;: Monte Perdido, Collata Añisolo. 2400 m a.s.l. 25.08.2005. A. Anichtchenko leg. (cSh); CASTILLA Y LÉON: 1 &male;: Astorga. Paganetti leg. (FMNH); 1 &male;: Palencia. Paganetti leg. (FMNH); MADRID: 1 &male;, 1 &female;: Cercedilla (MNCN); 50 &male;&male;, 42 &female;&female;: same loality. 05.1909. Bolivar leg. (MNCN); 1 &male;: Madrid. 06.1909 (MNCN); 1 &male;: Madrid. 1926. Lindberg leg. (MZHF); 1 &female;: Navacerrada. Martin leg. (IRSN); 1 &male;: Guadarrama, Eppelsheim leg. (SF); 1 &male;: ‘ Serra Guadarrama Escorial’ (NHMB); 2 &female;&female;: same locality. J. Ildefonso leg. (NMPC); EXTREMADURA: 1 &male;, 1 &female;: La Granja (IRSN, BMNH); ANDALUSÍA: 2 &male;&male;, 1 &female;, 13 exs.: Granada, Mt. Mulhacen. 2000 m a.s.l. 21.05.1967. M.E. Bacchus & B. Levey leg. (BMNH, MHNG); 1 &male;: Capileira. 1500 m a.s.l. 28.09.1993. P. Wunderle leg. (cW); 1 &female;: Sierra Nevada, Barranco de Cauchiles. 2000 m a.s.l. 23.07.1971. (cW); 1 ex.: ‘ Andalusia’ (ZIN); 2 &male;&male;, 1 &female;: ‘ Kraatz’, ‘ Andalusien’, ‘ Coll. K. Neumann’ (SF); 1 &male;: ‘ Sierra Nevada nivicola Kr. ’ (IRSN); LOCALITY NOT SPECIFIED: 1 &male;: ‘ Spanien’ (NHMB). Additional material. FRANCE: 1 &female;: Grottes de Bétharram (IRSN); 10 exs: Col de Peyresourde. 5,000 - 6,500 ft, in wet moss by stream. 04.09.1968 (BMNH); 1 &male;, 1 &female;: ‘ Lesteva pubescens Mnh. H. Pyrenees’ (MNCN); MOROCCO: 1 &male;: Sidi Ali Lake, 50 km S Ifrane. 6700 ft, island in lake, flood debris. 11.05.1961 (BMNH); ALGERIA: 1 &male;: ‘ mt. Babor Algérie’ (IRSN); LOCALITY NOT SPECIFIED: 3 exs.: ‘ Pyrenaei’ (ZIN). Remarks. Lesteva l. longoelytrata is a West Palaearctic subspecies, distributed in Orkney Islands, Faroe Islands, Iceland and Britain Isles through North Africa, Europe, Caucasus, Turkey, Lebanon, Cyprus, Turkmenistan and Iran (Shavrin 2014), replaced by L. l. maura Erichson, 1840 in Corsica and Sardinia, and by L. l. cretica Lohse & Steel, 1961 in Crete (Zanetti 1987). The status of these subspecies requires confirmation. In North Africa L. l. longoelytrata is known from Morocco (Koch 1941, Coiffait 1973) and Algeria (Fauvel 1878, 1886, 1902). This species is very common in the Iberian Peninsula (Gamarra & Outerelo 2008) and known from both Portugal and Spain (Kiesenwetter 1851, Heyden 1870, Quedenfeldt 1884, Champion & Chapman 1905, Lindberg & Bernhauer 1931, Koch 1940, Jeannel & Jarrige 1949, Jarrige 1954, Lohse 1960, Outerelo et al. 1998, etc.)., Published as part of Shavrin, Alexey V. & Zanetti, Adriano, 2021, Review of the genus Lesteva Latreille, 1797 (Coleoptera: Staphylinidae: Omaliinae Anthophagini) of the Iberian Peninsula, pp. 401-427 in Zootaxa 4966 (4) on page 411, DOI: 10.11646/zootaxa.4966.4.1, http://zenodo.org/record/4738667, {"references":["Goeze, J. A. E. (1777) Entomologische Beytrage zu des Ritter Linne zwolften Ausgabe des Natursystems. 1. Weidmanns Erben und Reich, Leipzig, xvi + 736 pp. https: // doi. org / 10.5962 / bhl. title. 45974","Luze G. (1903) Revision der palaarktischen Arten der Staphyliniden-Gattung Lesteva Latr. Verhandlungen der Kaiserlich- Koniglichen Zoologisch-Botanischen Gesellschaft in Wien, 53, 179 - 197.","Johansen, J. P. (1914) Danmarks rovbiller eller Billefam. Staphylinidae's danske Slaegter og Ar Territory. Bianco Lunos Bogtrykkeri, Kobenhavn, 660 pp.","Portevin, G. (1929) Histoire naturelle des Coleopteres de France. I. Adephaga. Polyphaga: Staphylinoidea. In: Encyclopedie Entomologique. Vol. 12. P. Lechevalier, Paris, pp. i - xii + 1 - 649.","Palm, T. (1948) Svensk Insekfauna. 9. Skalbaggar. Coleoptera. Kortvingar: Fam. Staphylinidae. Unterfam. Micropeplinae, Phloeocharinae, Olisthaerinae, Proteininae, Omaliinae. Almquist and Wiksells, Uppsala, 133 pp.","Jeannel, R. & Jarrige, J. (1949) Biospeologica. LXVIII. Coleopteres staphylinides (Premiere Serie). Archives de Zoologie Experimentale et Generale, 86, 255 - 392.","Lohse, G. A. (1956) Die mir aus Deutschland bekannt gewordenen Arten der Gattung Lesteva Latr. Entomologische Blatter fur Biologie und Systematik der Kafer, 51 (1 / 2), 51 - 60. [1955]","Lohse, G. A. (1960) Die Arten der Gattung Lesteva Latr. (Coleoptera Staphylinidae) der iberischen Halbinsel. Bulletin Institut Royal des Sciences Naturelles de Belgique, 36 (26), 1 - 7.","Lohse, G. A. (1964) Staphylinidae I. (Micropeplinae bis Tachyporinae). In: Freude, H., Harde, K. & Lohse, G. (Eds.), Die Kafer Mitteleuropas. Band 4. Goecke & Evers, Krefeld, pp. 1 - 264.","Toth, L. (1982) Magyarorszag Allatvilaga - Fauna Hungariae, Holyvak II. Staphylinidae II VII. Akademiai Kiado, Budapest, 110 pp.","Bordoni, A. (1973) Lesteva (Lestevina nov.) sbordonii n. sp. della Campania (Col. Staphylinidae). Redia, 54, 229 - 234.","Zanetti, A. (1987) Coleoptera. Staphylinidae. Omaliinae. In: Fauna d'Italia. Vol. 25. Calderini, Bologna, pp. i - xii + 1 - 472.","Zanetti, A. (2012) Unterfamilie Omaliinae. In: Assing, V. & Schulke, M. (Eds.), Freude-Harde-Lohse-Klausnitzer - Die Kafer Mitteleuropas. Band 4. Staphylinidae I. Zweite neubearbeitete Auflage. Spektrum Akademischer Verlag, Heidelberg, pp. 49 - 117.","Tronquet, M. (2006) Catalogue iconographique des Coleopteres des Pyrenees-orientales. Vol. 1. Staphylinidae. Revue de l`association roussillonnaise d`entomologie, 15, 1 - 79.","Shavrin, A. V. (2014) Review of the genus Lesteva Latreille, 1797 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini) of Iran and adjacent territories. Zootaxa, 3884 (2), 122 - 140. https: // doi. org / 10.11646 / zootaxa. 3884.2.2","Geoffroy, E. L. (1785) New species. In: Fourcroy, A. (Ed.), Entomologia Parisiensis; sive catalogus insectorum quae in agro Parisiensi reperiuntur; secundum methodum Geoffroeanum in sectiones, genera et species distributus; cui addita sunt nomina trivialia et fere trecentae novae species. Pars prima. Aedibus Serpentineis, Parisiis, pp. i - viii + 1 - 231.","Paykull, G. de (1789) Monographia Staphylinorum Sueciae. Johann, Edman, Upsaliae, 8 + 81 pp. https: // doi. org / 10.5962 / bhl. title. 146996","Panzer, G. W. F. (1795) Entomologia germanica exhibens insecta per germaniam indigena secundum classes, ordines, genera, species adiectis syonymis, locis, observationibus. I. Eleuterata. Felsecker, Normibergae, 8 (unnumbered) + 12 + 370 + 2 pp.","Block, F. L. H. von (1799) Verzeichniss der merkwurdigsten Insecten im Plauischen Grunde. In: Becker, W. G., Der Plauis-che Grund bei Dresden. Teil II. Frauenholz, Nurnberg, pp. 95 - 120.","Paykull, G. de (1800) Fauna Suecica. Insecta. Vol. 3. Joh. F. Edman, Upsaliae, 459 pp.","Marsham, T. (1802) Entomologia Britannica, sistens insecta Britanniae indigena, secundum methodum Linnaeanam disposita. Tomus I. Coleoptera. Londini: Wilks & Taylor, xxxi + 547 + [1] pp. https: // doi. org / 10.5962 / bhl. title. 65388","Latreille, P. A. (1804) Histoire naturelle, generale et particuliere, des crustaces et des insectes. 9. F. Dufart, Paris, 416 pp. https: // doi. org / 10.5962 / bhl. title. 15764","Gravenhorst, J. L. C. (1806) Monographia Coleopterorum Micropterorum. Henricus Dieterich, Gottingae, 248 pp. https: // doi. org / 10.5962 / bhl. title. 67769","Mannerheim, C. G. von (1830) Precis d'un nouvel arrangement de la famille des brachelytres de l'ordre des insectes coleopteres. s. n., St. Petersbourg, 87 pp","Stephens, J. F. (1834) Illustrations of British entomology; or, a synopsis of indigenous insects; containing their generic and specific dis-tinctions; with an account of their metamorphoses, times of appearance, localities, food, and economy, as far as practicable. Mandibulata, 5, 305 - 368. [Baldwin and Cradock, London]","Motschulsky, V. (1858) Enumeration des nouvelles especes de coleopteres rapportes de ses voyages. Bulletin de la Societe Imperiale des Naturalistes de Moscou, 30 (4), 490 - 517. [1857]","Mulsant, E. & Rey, C. (1870) Description de diverses especes nouvelles de coleopteres. Opuscules Entomologiques, 14, 105 - 122.","Mulsant, E. & Rey, C. (1871) Description de diverses especes nouvelles de coleopteres. Annales de la Societe Linneenne de Lyon, 18, 81 - 98. https: // doi. org / 10.3406 / linly. 1872.1881","Reitter, E. (1909) Fauna Germanica. Die Kafer des Deutschen Reiches. Nach der analytischen Methode bearbeitet. II Band. Schriften des Deutschen Lehrervereins fur Naturkunde 24. K. G. Lutz, Stuttgart, 392 pp., pls. 41 - 80. https: // doi. org / 10.5962 / bhl. title. 9570","Smetana, A. (1967) Wissenschaftliches Ergebnis der zoologischen Expedition des Nationalmuseum in Prag nach der Turkei. Coleoptera-Staphylinidae, Subfam. Oxytelinae. Acta Entomologica Musei Nationalis Pragae, 37, 297 - 324","Erichson, W. F. (1840) Genera et species Staphylinorum insectorum coleopterorum familiae. F. H. Morin, Berlin, 954 pp. https: // doi. org / 10.5962 / bhl. title. 59644","Koch, C. (1941) Recoltes de R. Paulian & A. Villiers dans le Haut-Atlas Marocain, 1938 (XIe note). Coleopteres Staphylinides. Annales de la Societe Entomologique de France, 110, 53 - 70.","Coiffait, H. (1973) Staphylinides nouveaux ou mal connus du Maroc. Bulletin de la Societe des Sciences Naturelles et Physiques du Maroc, 53, 269 - 292.","Fauvel, A. (1878) Les Staphylinides de l'Afrique boreale. Bulletin de la Societe Linneenne de Normandie, Series 3, 2, 83 - 162.","Fauvel, A. (1886) Les Staphylinides du Nord de l'Afrique. Revue d'Entomologie, 5, 9 - 100.","Fauvel, A. (1902) Catalogue des Staphylinides de la Barbarie de la Basse-Egypte et des Iles Acores, Maderes, Salvages et Canaries. Revue d'Entomologie, 21, 45 - 189.","Gamarra, P. & Outerelo, R. (2008) Catalogo iberobalear de los Omaliinae (Coleoptera: Staphylinidae). Boletin Sociedad Entomologica Aragonesa, 43, 211 - 231.","Kiesenwetter, E. A. H. von (1851) Enumeration des coleopteres trouves dans le midi de la France et en Catalogne. Annales de la Societe Entomologique de France, Series 2, 9, 385 - 440.","Heyden, L. (1870) Entomologische Reise nach dem Sudlichen Spanien, der Sierra Guadarrama und Sierra Morena, Portugal und den Cantabrichen gebirgen. Bei dr. G. Kraatz, Berlin. 217 pp.","Quedenfeldt, G. (1884) Beitrage zur Kenntniss der Staphylinen-Fauna von Sud-Spanien, Portugal und Marokko. Berliner Entomologische Zeitschrift, 28, 351 - 379. https: // doi. org / 10.1002 / mmnd. 18840280212","Champion, G. C. & Chapman, T. A. (1905) Another entomological excursion to Spain. Transactions of the Entomological Society of London, 1905 (1), 37 - 52. https: // doi. org / 10.1111 / j. 1365 - 2311.1905. tb 01154. x","Lindberg, H. & Bernhauer, M. (1931) Inventa entomologica itineris Hispanici et Maroccani, quod a. 1926 fecerunt Harald et Hakan Lindberg. Societas Scientarum Fennica, 3 (12), 1 - 29.","Koch, C. (1940) Resultados cientificos de los viajes entomologicos en Espana patrocinados por su Alteza el Principe Alessandro C. della Torre e Tasso. VI Congreso Internacional Entomologia, Madrid, 1935, pp. 369 - 390.","Jarrige, J. (1954) Coleopteres de la Sierra Nevada. \" Staphylinidae \". Archivos del Instituto de Alimatacion, 2, 73 - 79.","Outerelo, R., Gamarra, P. & Salgado, J. M. (1998) Los Staphylinidae cavernicolas del noreste de la Peninsula Iberica (I). Memoires de Biospeologie, 25, 111 - 137."]}

3.5. Pseudodialycera Zanetti & Shavrin, gen. nov. Type species: Omalium striatipenne Aubé, 1850. Description. Body glossy, narrow, and parallel-sided, with flattened forebody, Dropephylla -shaped. Punctation of forebody moderately dense and coarse; punctation coarser and deeper on elytra, with punctures arranged in longitudinal striae. Pubescence very short, except for some long setae on pronotum and elytra. Surface of forebody without microsculpture. Length 2.30–2.60 mm. Head distinctly narrower than pronotum, distinctly broader than long; vertex slightly elevated between two short supraocular furrows, scarcely convergent posteriorly; portions between supraocular furrows and eyes with longitudinal wrinkles; eyes normally developed, moderately large, much longer than short and angular temples, with distinct postocular carina; posterior constriction marked with distinct narrow furrow. Ocelli moderately distinct, with short anteocellar pits. Anterior margin of labrum slightly emarginate. Mandibles abruptly curved and acute at apex. Maxillary palpi with apical segment long, clearly narrower and slightly shorter than the somewhat dilated preapical palpomere (Fig. 16). Basal segment of labial palpi short and wide, segment 2 narrower, as wide as long, segment 3 narrower than preceding segment, as long as wide, apical segment narrow and elongate. Gular sutures moderately narrowly separated from each other at level of anterior portions of eyes and significantly divergent porteriorly. Antenna slightly exceeding than combined length of head and pronotum when reclined; antennomere 6 not narrower than 5 and 7, apical five antennomeres forming a loose club, with moderately short setation, and antennomeres 7–10 distinctly transverse. Pronotum slightly transverse, with irregularly crenulate lateral margins. Prosternum with moderately long intercoxal process reaching middle of coxae, with distinct median longitudinal carina. Mesoventrite short, with partial median carina; intercoxal process acute. Scutellum large and triangular. Metaventrite moderately broad, convex, with short angular intercoxal process. Elytra short, extending to anterior margin of abdominal tergite III, with distinctly crenulate lateral margins and broad epipleura. Wings fully developed. Legs moderately short; procoxae transverse with prominent distal portion; mesocoxae oval; metacoxae large and subtriangular; tibiae distinctly widened apicad; tarsi short and narrow, tarsomere 5 distinctly longer than the combined length of the remaining four tarsomeres. Abdomen convex, long, slightly narrower than elytra, with a pair of small wing-folding patches in middle of tergite IV. Male. Medioapical surface of apical abdominal sternites without modifications.Aedeagus of the basic omaliinae type (Figs. 59–60), with moderately wide basal portion and median lobe, encased by a single sclerotized piece, with only ventral median line unsclerotized, with moderately thick and short cylindrical parameres bearing modified setiferous surface in distal parts, with apical setae 2+2. Female. Apical abdominal sternites without modifications. Genital segment (Fig. 68) with apical lobes of gonocoxites long and narrow, with thin and short styli; accessory abdominal sclerite moderately wide and long, forming median piece divided into two elongate branches, with paired sclerotized apical structures. Spermatheca not found. Immature stages. Unknown. Species included. Pseudodialycera striatipennis (Aubé, 1850). Differential diagnosis. The type species was originally included in Dialycera, but it lacks most of the main characters of this genus (maxillary palpi with apical segment conical, slightly longer than the penultimate segment (Fig. 16), antennomere 6 slightly narrower than 5 and 7, male abdominal sternite VII unmodified, median lobe with ventral surface without narrow appendage (Fig. 59), with wide apical portion and cylindrical parameres somewhat similar to those of Acrulia (Figs. 33, 35, 37), female genital segment with elongate gonocoxites and short styli (Fig. 68). It shares with Dialycera only the the crenulate lateral margins of the pronotum. Pseudodialycera, a new monotypical genus of the subfamily Omaliinae, belongs to the group of genera of Omaliini without elliptical plate on the ventral surface of the aedeagus (see above). Based on the general shape of the forebody, it resembles species of the genus Dropephylla Mulsant & Rey, 1880, but the maxillary palpi of Pseudodialycera (Fig. 16) have a long and narrow apical segment similar to that in Phloeonomus Heer, 1839 and allied genera (Phloeostiba Thomson, 1858, Paraphloeostiba Steel, 1960 and Xylostiba Ganglbauer, 1895). Besides, the aedeagus of all these genera has an elliptical plate on its ventral surface. In addition to the conspicuous differences of the aedeagus (see Zanetti 2012), Pseudodialycera can be distinguished from these genera by the following morphological features: from Phloeonomus, Paraphloeostiba and Phloeostiba primarily by the glossy surface of the pronotum and elytra, without miscrosculpture, and from Xylostiba by the shorter setation of the antenna. Pseudodialycera is best identified based on the small, parallel-sided and rather flattened body, crenulate lateral margins of the pronotum, and the long apical segment of the maxillary palpi being distinctly narrower than the penultimate segment. Natural history. All specimens were sifted from forest litter (see details below). Distribution. Caucasus: Georgia (Fig. 74). Etymology. The feminine generic name is a combinations of Greek adjective ψ&epsi;&upsi;δο (false) and the existing name Dialycera, the genus in which the type species had been originally included.

Key to species of the genus Dialycera 1 Pronotum and elytra with fine, but distinct setation. Aedeagus as in Fig. 55. Female accessory sclerite as in Figs. 65���66. Body length: 1.70���2.00 mm. Habitus as in Figs. 41���42. Aegean islands across Anatolia to Israel and Jordan........... D. aspera - Dorsal surface glabrous or with barely visible, very sparse and short setation...................................... 2 2 Body parallel-sided, head clearly darker than pronotum. Aedeagus as Fig. 57. Female accessory sclerite as in Fig. 67. Body smaller: 1.53���1.80 mm. Habitus as in Fig. 43. Anatolia to Lebanon, Israel and Jordan, and to Armenia........... D. minuta - Body with rounded lateral margins, head slightly darker than the rest of the body. Aedeagus as in Fig. 53. Female accessory sclerite as in Fig. 64. Body larger: 1.90���2.10 mm. Habitus as in Fig. 40. Iberian Peninsula, South France, Italy, Balkans (along the Adriatic Sea)......................................................................... D. distincticornis, Published as part of Shavrin, Alexey V. & Zanetti, Adriano, 2020, Revision of Acrulia Thomson, 1858 and Dialycera Ganglbauer, 1895, with taxonomic notes on related genera and description of a new genus from the Caucasus (Coleoptera: Staphylinidae: Omaliinae: Omaliini), pp. 1-63 in Zootaxa 4800 (1) on page 25, DOI: 10.11646/zootaxa.4800.1.1, http://zenodo.org/record/3896791

Two fossil species, Paraphloeostiba electrica sp.n. and Phyllodrepa antiqua sp.n. (Staphylinidae, Omaliinae), are described from Baltic amber. Their external and relevant internal structures are illustrated using propagation phase contrast synchrotron microtomography. The palaeobiogeogaphy of the two genera, the thermophilous Paraphloeostiba, the temperate Phyllodrepa, as well as palaeoenvironment of the amber forest are discussed in light of the new findings.

Eusphalerum swauki Hatch, 1957 Eusphalerum swauki Hatch, 1957: 83; Moore and Legner 1975: 191: Herman 2001: 461. Anthobium californicum sensu Hatch, 1944 (not Fauvel, 1878)., Published as part of Zanetti, Adriano, 2014, Taxonomic revision of North American Eusphalerum Kraatz, 1857 (Coleoptera, Staphylinidae, Omaliinae), pp. 1-80 in Insecta Mundi 2014 (379) on page 18, DOI: 10.5281/zenodo.5179446, {"references":["Hatch, M. H. 1957. The Beetles of Pacific North West. Part II: Staphyliniformia. University of Washington Press; Seattle. ix + 384 p.","Moore, I., and E. F. Legner. 1975. A catalogue of Staphylinidae of America North of Mexico (Coleoptera). Division of Agricultural Sciences, University of California Special Publication 3015: 1 - 514.","Herman, L. H. 2001. Catalog of the Staphylinidae (Insecta: Coleoptera). 1758 to the end of second millennium. I. Introduction, history, biographical sketches, and Omaliinae group. Bulletin of the American Museum of Natural History 265: 1 - 650.","Hatch, M. H. 1944. Studies on Northwestern Staphylinidae I. Anthobium Steph. The Pan-Pacific Entomologist 22 (3): 101 - 104.","Fauvel, A. 1878. Les Staphylinides de l'Amerique du Nord. Bullettin de la Societe Linneenne de Normandie (3) 2: 167 - 269."]}

Eusphalerum aurifluum (Fauvel, 1878) Anthobium aurifluum Fauvel, 1878: 201; Bernhauer and Schubert 1910: 39. Eusphalerum aurifluum (Fauvel); Moore and Legner 1975: 190; Herman 20: 418., Published as part of Zanetti, Adriano, 2014, Taxonomic revision of North American Eusphalerum Kraatz, 1857 (Coleoptera, Staphylinidae, Omaliinae), pp. 1-80 in Insecta Mundi 2014 (379) on page 23, DOI: 10.5281/zenodo.5179446, {"references":["Fauvel, A. 1878. Les Staphylinides de l'Amerique du Nord. Bullettin de la Societe Linneenne de Normandie (3) 2: 167 - 269.","Bernhauer, M., and K. Schubert. 1910. Staphylinidae I. In S. Schenkling, Coleopterorum Catalogus, 5 (19): 1 - 86. Junk; Berlin.","Moore, I., and E. F. Legner. 1975. A catalogue of Staphylinidae of America North of Mexico (Coleoptera). Division of Agricultural Sciences, University of California Special Publication 3015: 1 - 514."]}

Eusphalerum torquatum (Marsham, 1802) Silpha torquata Marsham, 1802. Eusphalerum torquatum; Moore and Legner 1975: 191; Zanetti 1987: 133; Herman 2001: 463. Material examined: (13 specimens). CANADA. Newfoundland 2 mm 2 ff Goulds 2.07.1985 leg. Morris (CNC); 2 mm 7 ff St. John’s 23.07.1949 leg. Brown (CNC). Published records. St. Johns and Topsail in 1965 (...); Nova Scotia in 1983 (CNC) (Brown 1967; Klimaszewski et al. 2013). Measurements. Head length: 0.24-0.29; head width: 0.51-0.68; pronotal length: 0.48-0.57; pronotal width: 0.68-0.83; elytral length: 0.94-1.09; elytral width: 0.88-1.05; length (clypeus to apex of elytra): 1.75-2.01; total length: 1.8-2.2. Description. Habitus as in Fig. 18. Head, pronotum, and elytra yellowish, elytra somewhat paler, neck often darkened; abdomen blackish or dark brown, prosternum yellowish, metasternum blackish; legs, antennae, and mouthparts yellowish, antennae darkened from antennomere 6-8. Head with prominent eyes, postocular carina present and well marked, temples short, strongly convergent caudad, medial margin of eyes without longitudinal wrinkles. Head flat, postantennal depressions very superficial and tentorial pits small. Between ocelli a small tubercle (vestigial ocellum?) is often present, as in some European specimens. Neck clearly separated from the head (Fig. 19). Punctation very superficial on strongly microsculptured, obsolete, ground. Antennae slightly elongate, antennomere 1 rather elongate, twice as long as wide, 2 ovoid elongate, 3 twice as long as wide, 4-7 longer than wide, 8-10 subquadrate, 11 twice as long as wide, rather ovoid, sharpened at apex. Pronotum transverse (ratio width/length = 1.4 on average), anterior margin wider than posterior in male, as wide as posterior in female, strongly convex in male, less convex in female, with a median superficial furrow in anterior two thirds, widest just in front of middle, lateral margins rounded at middle, convergent caudad in straight line, posterior angles marked and scarcely obtuse. Punctation dense but very superficial, ground with strong isodiametric microsculpture; pubescence long, yellowish, decumbent, directed caudally on disk, towards midline in front of posterior margin, depressions near posterior angles superficial, extending in front of middle of lateral margin. Elytra scarcely elongate (ratio length from scutellum to apex / combined width of elytra = 1.0), widened towards apex, apices truncate medially in male, prolonged in short lobes (Fig. 21) in female, punctation somewhat coarse, somewhat confluent on glossy ground, pubescence short but clearly visible, scarcely decumbent. Abdomen dull, microsculture well visible, pubescence long, decumbent. Tibiae straight in both sexes, tarsomere 5 of posterior tarsi slightly shorter than 1-4 together. Aedeagus as in Fig. 20. Accessory sclerites of female as in Fig 22, spermatheca as in Fig. 23. Comparative notes. Eusphalerum torquatum is easily separable from the other species living in Canada by its color pattern, which recalls light populations of E. californicum. The long pubescence, dense superficial punctation of pronotum, and impressed line that separates head from neck are distinctive. The form of the elytral apex of the female is also characteristic. The presence of a third vestigial ocellus is also recorded in the European species of the robustum group as well as occasionally in other omaliinegroup taxa. Distribution. North America: CANADA: Newfoundland and Labrador, Nova Scotia (Map 1). Palaearctic region: Iberian Peninsula, Great Britain, Central Europe, mostly in western regions, also reported from Japan (Zanetti 1987). Various forms and subspecies have been described from southern France and Spain, differentiated in color and form of elytra of the females, but their validity is doubtful (Zanetti 1987). The North American specimens belong to the typical form. Eusphalerum torquatum is very probably an adventive species in North America. Natural history. The species is regularly found on broom flowers, mostly Sarothamnus (Fabaceae), in Europe. This plant was introduced in North America and it is now widely distributed in the Pacific states, where it is considered a destructive invasive species, but it is not present in Newfoundland. The host plant(s) in this region is not known. Captures of E. torquatum in July., Published as part of Zanetti, Adriano, 2014, Taxonomic revision of North American Eusphalerum Kraatz, 1857 (Coleoptera, Staphylinidae, Omaliinae), pp. 1-80 in Insecta Mundi 2014 (379) on pages 11-12, DOI: 10.5281/zenodo.5179446, {"references":["Marsham, T. 1802. Entomologia Britannica, sistens Insecta Coleoptera Britanniae indigena, secundum methodum Linnaeanam disposita. Tomus I. Coleoptera. Wilks and Taylor; London, xxxi + 547 + [1] p.","Moore, I., and E. F. Legner. 1975. A catalogue of Staphylinidae of America North of Mexico (Coleoptera). Division of Agricultural Sciences, University of California Special Publication 3015: 1 - 514.","Zanetti, A. 1987. Fauna dItalia XXV. Coleoptera Staphylinidae Omaliinae. Calderini; Bologna. 472 p.","Herman, L. H. 2001. Catalog of the Staphylinidae (Insecta: Coleoptera). 1758 to the end of second millennium. I. Introduction, history, biographical sketches, and Omaliinae group. Bulletin of the American Museum of Natural History 265: 1 - 650.","Brown, W. J. 1967. Notes on the extralimited distribution of some species of Coleoptera. The Canadian Entomologist 99: 85 - 93.","Klimaszewski, J., A. V. Brunke, V. Assing, D. W. Langor, A. F. Newton, C. Bourdon, G. Pelletier, R. P. Webster, L. Herman, L. Perdereau, A. Davies, A. Smetana, D. S. Chandler, C. Majka, and G. G. E. Scudder. 2013. Synopsis of adventive species of Coleoptera (Insecta) recorded from Canada. Part 2: Staphylinidae. Pensoft; Sofia-Moscow. 360 p."]}

Eusphalerum horni (Fauvel, 1878) Anthobium horni Fauvel, 1878: 202; Bernhauer and Schubert 1910: 40; Blatchley 1910: 481. Eusphalerum horni (Fauvel); Moore and Legner 1975: 191; Downie and Arnett 1996: 439; Herman 2001: 425. Anthobium flavum Bernhauer, manuscript name., Published as part of Zanetti, Adriano, 2014, Taxonomic revision of North American Eusphalerum Kraatz, 1857 (Coleoptera, Staphylinidae, Omaliinae), pp. 1-80 in Insecta Mundi 2014 (379) on page 38, DOI: 10.5281/zenodo.5179446, {"references":["Fauvel, A. 1878. Les Staphylinides de l'Amerique du Nord. Bullettin de la Societe Linneenne de Normandie (3) 2: 167 - 269.","Bernhauer, M., and K. Schubert. 1910. Staphylinidae I. In S. Schenkling, Coleopterorum Catalogus, 5 (19): 1 - 86. Junk; Berlin.","Blatchley, W. S. 1910. An illustrated descriptive catalogue of the Coleoptera or beetles (exclusive of Rhynchophora) known to occur in Indiana [...]. W. S. Blatchley; Indianapolis. 1385 p.","Moore, I., and E. F. Legner. 1975. A catalogue of Staphylinidae of America North of Mexico (Coleoptera). Division of Agricultural Sciences, University of California Special Publication 3015: 1 - 514.","Downie, N. M., and R. H Arnett. 1996. The beetles of northeastern North America. Volume 1: Introduction: Suborders Archostemata, Adephaga and Polyphaga, thru Superfamily Cantharoidea. Sandhill Crane Press; Gainesville. 880 p.","Herman, L. H. 2001. Catalog of the Staphylinidae (Insecta: Coleoptera). 1758 to the end of second millennium. I. Introduction, history, biographical sketches, and Omaliinae group. Bulletin of the American Museum of Natural History 265: 1 - 650."]}

Eusphalerum margaretae Zanetti n. sp. Material examined (1 specimen) Holotype m Tennessee Sevier Co S. Gatlinburg 24.05.1977 leg. M. Thayer. Measurements. Head length: 0.22; head width: 0.44; pronotal length: 0.44; pronotal width: 0.57; elytral length: 0.88; elytral width: 0.83; length (clypeus to apex of elytra): 1.69; total length: 1.8. Etymology. The species, like E. thayeranum n. sp., is dedicated to its collector, Margaret K. Thayer, specialist of Staphylinidae Omaliinae. Description. Habitus as in Fig. 130. Head, pronotum and elytra yellowish; abdomen brown, somewhat paler at apex; prosternum yellowish, metasternum brownish, legs yellowish; antennae yellowish, slightly darkened from antennomere 7. Head with prominent eyes, postocular carina well marked; temples short, convergent caudad. Medial margin of eyes with microsculpture formed by meshes slightly lengthened, postantennal depression large not much deep, confluent with the rounded tentorial pits forming 2 superficial longitudinal depressions in front of ocelli, punctation very superficial and sparse, ground with isodiametric microsculpture, neck scarcely separated from head. Antennae rather short, antennomere 1 and 2 ovoid, 3 elongate, twice as long as wide, 4 and 5 longer than wide, 6 subquadrate, 7-10 transverse, 10 less than twice as wide as long, 11 twice as long as wide, cylindrical in the proximal part, conical at apex. Pronotum not much convex, without impressions, transverse (ratio width/length = 1.45), widest slightly in front of middle, anterior margin slightly narrower than posterior, lateral margins rounded, posterior angles scarcely obtuse and well marked, punctation very sparse and superficial, surface with isodiametric microsculpture, pubescence extremely short, scarcely visible, depressions near posterior angles distinct, extending anterad in front of middle. Elytra scarcely elongate (ratio length from scutellum to apex / combined width of elytra = 1.0), truncate at apex, clearly widened toward apex, punctation much stronger than on pronotum, irregular and scarcely confluent, ground glossy, pubescence extremely short and scarcely visible. Abdomen rather glossy, microsculpture superficial, with decumbent pubescence. Tibiae straight, not modified, tarsomere 5 of posterior tarsi as long as 1-4 together. Aedeagus as in Fig. 131, internal sac with 2 short spines. Female unknown. Comparative notes. Eusphalerum margaretae is close to E. thayeranum, which has different shaped spines in the internal sac of the aedeagus. It is also similar to E. fenyesi, but the latter lacks the spines of the internal sac. The unique known specimen has a brown metasternum (yellowish in E. thayeranum). Eusphalerum orientale is distinguished by the even more sparse and superficial punctation of the pronotum, and by the presence of 2 rows of fine spines in the internal sac of the aedeagus. Distribution. UNITED STATES: Tennessee (Map 3). Natural history. The specimen was collected in May., Published as part of Zanetti, Adriano, 2014, Taxonomic revision of North American Eusphalerum Kraatz, 1857 (Coleoptera, Staphylinidae, Omaliinae), pp. 1-80 in Insecta Mundi 2014 (379) on page 46, DOI: 10.5281/zenodo.5179446

Eusphalerum punctatum (Casey, 1894) Anthobium punctatum Casey, 1894: 429; Bernahauer and Schubert 1910: 42. Eusphalerum punctatum (Casey); Moore and Legner 1975: 191; Herman 2001: 450., Published as part of Zanetti, Adriano, 2014, Taxonomic revision of North American Eusphalerum Kraatz, 1857 (Coleoptera, Staphylinidae, Omaliinae), pp. 1-80 in Insecta Mundi 2014 (379) on page 47, DOI: 10.5281/zenodo.5179446, {"references":["Casey, T. L. 1894. Coleopterological notices. V. Annals of the New York Academy of Sciences 7: 281 - 606.","Moore, I., and E. F. Legner. 1975. A catalogue of Staphylinidae of America North of Mexico (Coleoptera). Division of Agricultural Sciences, University of California Special Publication 3015: 1 - 514.","Herman, L. H. 2001. Catalog of the Staphylinidae (Insecta: Coleoptera). 1758 to the end of second millennium. I. Introduction, history, biographical sketches, and Omaliinae group. Bulletin of the American Museum of Natural History 265: 1 - 650."]}

Eusphalerum rugulosum (Mäklin, 1853) Anthobium rugulosum Mäklin, 1853: 199; Fauvel, 1878: 199; Bernhauer and Schubert, 1910: 43. Eusphalerum rugulosum (Mäklin); Moore and Legner 1975: 191; Herman 2001: 453. Anthobium grayae Hatch, 1944: 102 n. syn. Eusphalerum grayae (Hatch); Hatch 1957: 83; Moore and Legner 1975: 191; Herman 2001: 425., Published as part of Zanetti, Adriano, 2014, Taxonomic revision of North American Eusphalerum Kraatz, 1857 (Coleoptera, Staphylinidae, Omaliinae), pp. 1-80 in Insecta Mundi 2014 (379) on page 34, DOI: 10.5281/zenodo.5179446, {"references":["Fauvel, A. 1878. Les Staphylinides de l'Amerique du Nord. Bullettin de la Societe Linneenne de Normandie (3) 2: 167 - 269.","Bernhauer, M., and K. Schubert. 1910. Staphylinidae I. In S. Schenkling, Coleopterorum Catalogus, 5 (19): 1 - 86. Junk; Berlin.","Moore, I., and E. F. Legner. 1975. A catalogue of Staphylinidae of America North of Mexico (Coleoptera). Division of Agricultural Sciences, University of California Special Publication 3015: 1 - 514.","Herman, L. H. 2001. Catalog of the Staphylinidae (Insecta: Coleoptera). 1758 to the end of second millennium. I. Introduction, history, biographical sketches, and Omaliinae group. Bulletin of the American Museum of Natural History 265: 1 - 650.","Hatch, M. H. 1944. Studies on Northwestern Staphylinidae I. Anthobium Steph. The Pan-Pacific Entomologist 22 (3): 101 - 104.","Hatch, M. H. 1957. The Beetles of Pacific North West. Part II: Staphyliniformia. University of Washington Press; Seattle. ix + 384 p."]}

Eusphalerum convexum (Fauvel, 1878) Anthobium convexum Fauvel, 1878: 201; Bernhauer and Schubert 1910: 39; Blatchley, 1910: 481. Eusphalerum convexum (Fauvel); Moore and Legner 1975: 190; Dearborn and Donahue 1994: 13., Published as part of Zanetti, Adriano, 2014, Taxonomic revision of North American Eusphalerum Kraatz, 1857 (Coleoptera, Staphylinidae, Omaliinae), pp. 1-80 in Insecta Mundi 2014 (379) on page 25, DOI: 10.5281/zenodo.5179446, {"references":["Fauvel, A. 1878. Les Staphylinides de l'Amerique du Nord. Bullettin de la Societe Linneenne de Normandie (3) 2: 167 - 269.","Bernhauer, M., and K. Schubert. 1910. Staphylinidae I. In S. Schenkling, Coleopterorum Catalogus, 5 (19): 1 - 86. Junk; Berlin.","Blatchley, W. S. 1910. An illustrated descriptive catalogue of the Coleoptera or beetles (exclusive of Rhynchophora) known to occur in Indiana [...]. W. S. Blatchley; Indianapolis. 1385 p.","Moore, I., and E. F. Legner. 1975. A catalogue of Staphylinidae of America North of Mexico (Coleoptera). Division of Agricultural Sciences, University of California Special Publication 3015: 1 - 514.","Dearborn, R. G., and C. P. Donahue. 1994. The forest insect survey of Maine. An annotated list of insects collected and recorded by the Maine Forest Service. Order Coleoptera. Beetles. Insect and Disease Division, Department of Conservation. Technical Report 32 (1993). ix + 102 p."]}

Eusphalerum subangulatum (Casey, 1894) Anthobium subangulatum Casey, 1894: 430; Bernhauer and Schubert 1910: 43. Eusphalerum subangulatum (Casey) Moore and Legner 1975: 191; Downie and Arnett 1996: 439; Herman 2001: 460., Published as part of Zanetti, Adriano, 2014, Taxonomic revision of North American Eusphalerum Kraatz, 1857 (Coleoptera, Staphylinidae, Omaliinae), pp. 1-80 in Insecta Mundi 2014 (379) on page 8, DOI: 10.5281/zenodo.5179446, {"references":["Casey, T. L. 1894. Coleopterological notices. V. Annals of the New York Academy of Sciences 7: 281 - 606.","Bernhauer, M., and K. Schubert. 1910. Staphylinidae I. In S. Schenkling, Coleopterorum Catalogus, 5 (19): 1 - 86. Junk; Berlin.","Moore, I., and E. F. Legner. 1975. A catalogue of Staphylinidae of America North of Mexico (Coleoptera). Division of Agricultural Sciences, University of California Special Publication 3015: 1 - 514.","Downie, N. M., and R. H Arnett. 1996. The beetles of northeastern North America. Volume 1: Introduction: Suborders Archostemata, Adephaga and Polyphaga, thru Superfamily Cantharoidea. Sandhill Crane Press; Gainesville. 880 p.","Herman, L. H. 2001. Catalog of the Staphylinidae (Insecta: Coleoptera). 1758 to the end of second millennium. I. Introduction, history, biographical sketches, and Omaliinae group. Bulletin of the American Museum of Natural History 265: 1 - 650."]}

Eusphalerum fenyesi (Bernhauer, 1912) Anthobium fenyesi Bernhauer, 1912: 679; Scheerpeltz 1933: 1033. Eusphalerum fenyesi (Bernhauer); Hatch 1957: 82; Moore and Legner 1975: 190; Herman 2001: 423., Published as part of Zanetti, Adriano, 2014, Taxonomic revision of North American Eusphalerum Kraatz, 1857 (Coleoptera, Staphylinidae, Omaliinae), pp. 1-80 in Insecta Mundi 2014 (379) on page 40, DOI: 10.5281/zenodo.5179446, {"references":["Bernhauer, M. 1912. Zur Staphylinidenfauna von Nordamerica. V. Beitrag. Pomona College Journal of Entomology 4 (1): 678 - 683.","Scheerpeltz, O. 1933. Staphylinidae VII. Coleopterorum Catalogus 6 (129): 989 - 1500.","Hatch, M. H. 1957. The Beetles of Pacific North West. Part II: Staphyliniformia. University of Washington Press; Seattle. ix + 384 p.","Moore, I., and E. F. Legner. 1975. A catalogue of Staphylinidae of America North of Mexico (Coleoptera). Division of Agricultural Sciences, University of California Special Publication 3015: 1 - 514.","Herman, L. H. 2001. Catalog of the Staphylinidae (Insecta: Coleoptera). 1758 to the end of second millennium. I. Introduction, history, biographical sketches, and Omaliinae group. Bulletin of the American Museum of Natural History 265: 1 - 650."]}

Eusphalerum thayeranum Zanetti n. sp. Material examined (93 specimens) Holotype m 4 paratypes mm 2 paratypes ff British Columbia Garibaldi P.P., Cheakamus Lk. 3000 20.07.1973 leg. A. & Z. & D. Smetana (CNC) Other paratypes. CANADA. Alberta 1 m Ponaka (2 mi. S), Battle R. 6.08.1976 leg. J.M. Campbell (CNC); 1 m Waterton Lks. NP, Crypt Lake Tr. 4500' 22.06.1980 leg. JM Campbell (CNC); 1 m Waterton N.P., Lineham Creek Tr. 5900 6500 3.08.1976 leg. J.M. Campbell (CNC); 4 mm 2 ff Waterton N.P., Rowe Creek 5460 2.08.1976 leg. J.M. Campbell (CNC); 4 ff Waterton N.P., Rowe Creek 5400' 4.08.1976 leg. JM Campbell (CNC); British Columbia 1 m Courtney, Forbidden Plateau near Courtney Lookout 25.07.1979 leg. I.M. Smith (CNC); 3 mm 4 ff Klimpton Cr., Kootenay N.P. 4000 21.07.1971 leg. J.M. & B.A. Campbell (CNC); 3 mm 6 ff Kootenay N.P., Kindersley Pass 7000 23.07.1971 leg. J.M. & B.A. Campbell (CNC); 10 mm 11 ff Manning Pr. Pk, Rhododendron Flats Rhododendron flowers 21.06.1988 leg. J.M. Campbell (CNC); 1 m Manning Prov. Park, Sumallo Grove (Hwy5a) 650 m 10.07.1988 leg. Burckhardt & Loebl (MHNG); 1 m 3 f Manning Prov. Pk., Sumallo Grove Cornus flower 21.06.1988 leg. J.M. Campbell (CNC); 10 mm 10 ff Mount Cain 27.07/ 10.08.1996 leg. Winchester (cZan); 3 mm Osoyoos grass clippings 24.06.1988 leg. J.M. Campbell (CNC); 2 mm 2 ff Silver Star Prov. Pk. 6200 ground squirrel burrows (1 spec.) 8.08.1973 leg. R.H. Parry (CNC); 1 m 1 f Squamish, Diamond Head Trail 3500 10.08.1953 leg. G.J. Spencer (CNC); 1 m 2 ff Squaw Valley, nr. Mabel Lake 5.08.1982 leg. R. Baranowski (MZLU); 3 mm 6 f Wells Gray Prov. Pk., Falls Ck 30.06.1988 leg. J.M. Campbell (CNC); 10 mm 10 ff Mt. Cain 50 o 13’N 126 o 21’W 13.07/ 27.07.1997 leg. Corkum (cZan). USA. Idaho 1 m Bonner Co Sandpoint 20.06.1971 leg. N. M. Downie (FMNH); Indiana 1 m Tippecanoe Co 15.05.1976 leg. N.M. Downie (FMNH) [doubtful record]; Maine (?). 1 m Bascea (??) 2000 m 18.07.19?? (interrogation marks in the label) leg. N. M. Downie (FMNH) [doubtful record]; Oregon 1 m Clackamas Co jct US26 Ore 35 (1.3 m E) 3900 ft on flowers Achlys triphylla 12.07.1975 leg. A. Newton and M. Thayer (FMNH); 1 m 1 f Clackamas Co jct US26 Ore 35 (1.3 m E) 3900 on flowers Lupinus 12.07.1975 leg. A. Newton and M. Thayer (FMNH); 5 mm 16 ff Clackamas Co Mt. Hood N.F., Still Creek Cmpg. 4000' 31.07.1979 leg. JM & BA Campbell (CNC); 1 m Wallawa Co Eagles Cap Prim. Area, Moccasin Lk. 7500' 29.07.1981 leg. JM Campbell (CNC); 1 f Mt. Hood, Timberline Lodge Rd. 4800' 29.07.1979 leg. JM & BA Campbell (CNC); Washington 2 mm Clallam Co Olympic N.P., Hurricane Ridge Rd 625 m mixed coniferous forest, on flowers Rubus parviflora 9.07.1988 leg. A. Newton and M. Thayer (FMNH); 2 mm 2 ff Clallam Co Olympic N.P., Lake Creek at Hurricane Ridge Rd., 625 m mixed conifer forest, on flowers Clintonia uniflora 9.07.1988 leg. A. Newton and M. Thayer (FMNH); 9 mm 11 ff Whatcom Co Mount Baker 29.07.1931 leg. M. Sanderson (KSEM); 3 mm 1 f Whatcom Co Mt. Baker 1500 m 19.06.1987 leg. W.M. Downie (FMNH). Measurements. Head length: 0.20-0.25; head width: 0.42-0.55; pronotal length: 0.35-0.64; pronotal width: 0.42-0.70; elytral length: 0.85-1.12; elytral width: 0.79-1.03; length (clypeus to apex of elytra): 1.42-1.85; total length: 1.8-2.4. Etymology. The species is dedicated to one of its collectors, Margaret K. Thayer, specialist of Staphylinidae Omaliinae, who has helped me greatly in the study of North American Eusphalerum. Description. Habitus as in Fig. 126. Head, pronotum and elytra yellowish; abdomen dark brown, somewhat paler at apex (male) or entirely yellowish, very feebly darkened at apex (female); prosternum, metasternum, and legs yellowish; antennae yellowish, more or less darkened from antennomere 7, often entirely yellowish. Head with prominent eyes; postocular carina well marked; temples short, convergent caudad. Medial margin of eyes with microsculpture formed by slightly lengthened meshes; postantennal depression large, not very deep, confluent with the small, rounded tentorial pits forming 2 superficial longitudinal depressions in front of ocelli; punctation very superficial and sparse; ground with isodiametric microsculpture; neck scarcely separated from head. Antennae rather short, antennomere 1 and 2 ovoid, 3 elongate, twice as long as wide, 4 and 5 longer than wide, 6 subquadrate, 7-10 transverse, 10 less than twice as wide as long, 11 twice as long as wide, cylindrical in the proximal part, conical at apex. Pronotum not very convex, without impressions, transverse (ratio width/length = 1.4-1.5), widest slightly in front of middle, anterior margin slightly narrower than posterior, lateral margins rounded, posterior angles scarcely obtuse and well marked, punctation very sparse and superficial, ground with isodiametric microsculpture, pubescence extremely short, scarcely visible, depressions near posterior angles not much marked, extended anteriad in front of middle. Elytra scarcely elongate (ratio length from scutellum to apex / combined width of elytra = 1.0), truncate at apex in both sexes, clearely widened toward apex, punctation much stronger than on pronotum, irregular and scarcely confluent, ground glossy, pubescence extremely short and scarcely visible. Abdomen rather glossy, microsculpture superficial with decumbent pubescence. Tibiae straight in both sexes, not modified, tarsomere 5 of posterior tarsi as long as 1-4 together. Aedeagus as in Fig. 127, internal sac with 2 long spines. Accessory sclerites of female as in Fig. 128, spermatheca as in Fig. 129. Comparative notes. Eusphalerum thayeranum is very similar to E. fenyesi in external characters, but metasternum is usually yellowish instead of brownish. The presence of 2 long spines in the internal sac of the aedeagus is very characteristic; those of E. margaretae are much shorter. Distribution. CANADA: Alberta, British Columbia; UNITED STATES: Idaho, Indiana, Oregon, Washington (Map 3). The record from Indiana (Tippecanoe) is questionable, as it is outside the main geographical range of the species and the area does not possess the normal ecological conditions in which E. thayeranum lives. Its collector, N. M. Downie, lived and collected for decades in Tippecanoe Co., but also collected extensively in the western USA, so a label mixup may have occurred. The Maine (?) record is likewise doubtful. Natural history. The species was collected in a wide altitudinal range, between 600 and 2300 m. Reported host plants: Rhododendron sp. (Ericaceae), Cornus sp. (Cornaceae), Achlys triphylla (Sm.) DC. (Berberidaceae), Lupinus sp. (Fabaceae), Rubus parviflorus Nutt. (Rosaceae), Clintonia uniflora (Menzies ex Schult. & Schult.f.) Kunth (Liliaceae). The only macrohabitat reported on the labels is mixed conifer forest. Most captures between June and August., Published as part of Zanetti, Adriano, 2014, Taxonomic revision of North American Eusphalerum Kraatz, 1857 (Coleoptera, Staphylinidae, Omaliinae), pp. 1-80 in Insecta Mundi 2014 (379) on pages 44-45, DOI: 10.5281/zenodo.5179446

Xylodromus segmentarius (Mäklin, 1852) n. comb. Omalium segmentarium: Mäklin, 1852: 322. Anthobium segmentarium (Mäklin); Fauvel 1878: 200; Bernhauer and Schubert 1910: 43. Eusphalerum segmentarium (Mäklin); Moore and Legner 1975: 191; Herman 2001: 454. Examined type material. Holotype male Sitcha/Holmberg/166 [?]/ Omalium segmentarium Mäkl. /Mus. Zool. H:fors/Spec. typ. 2127/Mus. Hels. N:o 1766/ Xylodromus segmentarius (Mäklin 1852) det. A. Zanetti 2013/ Holotypus (red) (MZH). This species described from “insula Sitkha” (Alaska) was included in Eusphalerum (as Anthobium) by Fauvel on the basis of the form of the elytral apex (“... ad angulum interiorem apice producta; verisimile ad differentiam sexualem spectat;” (Mäklin, 1852). The examination of the type demonstrates that the species belongs to the genus Xylodromus Heer, 1839 (Fig. 153) and that it is not a junior synonym either of Xylodromus concinnus (Marsham, 1802) or of Xylodromus depressus (Gravenhorst, 1802), European species both also reported from North America. The validity of Xylodromus capito (Casey, 1894) (type from Wisconsin) requires examination of the type., Published as part of Zanetti, Adriano, 2014, Taxonomic revision of North American Eusphalerum Kraatz, 1857 (Coleoptera, Staphylinidae, Omaliinae), pp. 1-80 in Insecta Mundi 2014 (379) on page 59, DOI: 10.5281/zenodo.5179446, {"references":["Maklin, F. G. 1852. [New species and notes]. In: C. v. Mannerheim, Zweiter Nachtrag zur Kaefer-Fauna der Nord-Amerikanischen Laender des Russischen Reiches. Bulletin de la Societe Imperiale des Naturalistes de Moscou 25 (2): 283 - 387.","Fauvel, A. 1878. Les Staphylinides de l'Amerique du Nord. Bullettin de la Societe Linneenne de Normandie (3) 2: 167 - 269.","Bernhauer, M., and K. Schubert. 1910. Staphylinidae I. In S. Schenkling, Coleopterorum Catalogus, 5 (19): 1 - 86. Junk; Berlin.","Moore, I., and E. F. Legner. 1975. A catalogue of Staphylinidae of America North of Mexico (Coleoptera). Division of Agricultural Sciences, University of California Special Publication 3015: 1 - 514.","Herman, L. H. 2001. Catalog of the Staphylinidae (Insecta: Coleoptera). 1758 to the end of second millennium. I. Introduction, history, biographical sketches, and Omaliinae group. Bulletin of the American Museum of Natural History 265: 1 - 650.","Marsham, T. 1802. Entomologia Britannica, sistens Insecta Coleoptera Britanniae indigena, secundum methodum Linnaeanam disposita. Tomus I. Coleoptera. Wilks and Taylor; London, xxxi + 547 + [1] p.","Gravenhorst, J. - L. C. 1802. Coleoptera Microptera Brunsvicensia. Carolus Reichard; Brunswick. lxvi + 206 p.","Casey, T. L. 1894. Coleopterological notices. V. Annals of the New York Academy of Sciences 7: 281 - 606."]}

Eusphalerum newtoni Zanetti n. sp. Material examined (77 specimens) Holotype m 3 paratypes mm 1 paratype f California Marin Co Inverness, (3.1 mi. NW of) 200 ft Alnus forest, on flowers Heracleum lanatum 22.05.1976 leg. A. Newton and M. Thayer (FMNH) Other paratypes. CANADA. British Columbia 4 mm 2 ff Bowser 5/ 19.06.1955 leg. W.J. Brown & R. Coyles (CNC); 1 m Mission City 3.06.1953 leg. W. Mason (CNC); 1 m Prince Rupert, Mt. Hays 1 2000 29.06.1968 leg. Campbell & Smetana (CNC); 2 mm 1 f Vancouver Is., Port Renfrew 12.07.1984 leg. L. Huggert (MZLU). USA. Oregon 2 mm 1 f Douglas Co Reedsport (5 km S) 7.06.1984 leg. R. Danielsson (MZLU); Washington 1 f Clallam Co Fairholm (6 mi. ESE), Olympic Nat. Forest sweeping on forest roads and meadows 11.08.1988 leg. R. Baranowski (MZLU); 4 mm 5 ff Clallam Co Olympic NP, Lake Creek at Hurricane Ridge Rd. 625 m mixed conifer forest on flowers Tiarella trifoliata 09.07.1988 leg. A. Newton M. Thayer (FMNH); 11 mm 9 ff Clallam Co Olympic NP, Lake Creek at Hurricane Ridge Rd. 625 m mixed conifer forest on flowers Clintonia uniflora 09.07.1988 leg. A. Newton M. Thayer (FMNH); 2 ff Clallam Co Olympic NP, Lake Creek at Hurricane Ridge Rd. 625 m mixed conifer forest on flowers Cornus canadensis 09.07.1988 leg. A. Newton M. Thayer (FMNH); 1 m 3 f Clallam Co Olympic NP, Lake Creek at Hurricane Ridge Rd. 625 m mixed conifer forest on flowers Asteraceae 09.07.1988 leg. A. Newton M. Thayer (FMNH); 1 m Swob ?? 24.05.19?? leg. L. Russell (FMNH); California 3 mm 1 f Colusa Co Goat Mt. (?) 30.05.1959 (CNC); 1 m Marin Co Taylor State Pk. flowers Rubus parviflorus 8.05.1949 leg. H.P. Leech (CNC); 6 mm 9 ff Marin Co Alpine Lk 11.06.1964 leg. Doyen (FMNH); 1 m Mendocino 26.05.1955 leg. Helfer (CNC); 1 f Napa Co Calistoga (11.4 mi. N) 1600 21.05.1976 leg. A. Newton and M. Thayer (FMNH). Etymology. The species is dedicated to one of its collectors, the student of evolution of Staphylinidae Alfred F. Newton. Measurements. Head length: 0.27-0.33; head width: 0.53-0.59; pronotal length: 0.40-0.53; pronotal width: 0.64-0.75; elytral length: 0.99-1.35; elytral width: 0.88-1.11; length (clypeus to apex of elytra): 1.59-2.03; total length: 1.7-2.9. Description. Habitus as in Fig. 141. Head from yellowish slightly darkened on vertex to brown with anterior part medially yellowish; pronotum yellowish, sometimes darkened on discal part; elytra yellowish; abdomen brown with paler apex in male, entirely yellowish in female; prosternum yellowish; metasternum brownish yellow (male) or yellowish (female); legs yellowish; antennae yellowish at base, more or less darkened from antennomere 6. Head with prominent eyes, postocular carina well marked, temples short and convergent caudad, medial margin of eyes with microsculpture forming longitudinal wrinkles, postantennal depressions confluent caudally with the deep tentorial pits forming 2 longitudinal depressions in front of ocelli. Neck wide, not separated from the head. Punctation rather sparse and superficial on microsculptured ground. Antennae moderately elongate, antennomere 1 about twice as long as wide, 2 ovoid, 3 thin, elongate, twice as long as wide, 4-5 longer than wide, 6 subquadrate, 7-10 slightly transverse, 11 twice as long as wide, cylindrical at base and conical at apex. Pronotum transverse (ratio width/length = 1.4 on average), convex, with a median longitudinal impression in large males, a narrower furrow in the small males and in females. Pronotum widest in anterior half, anterior margin slightly narrower than posterior, lateral margins rounded in anterior third, usually convergent caudad in straight line in posterior two thirds, posterior angles marked and scarcely obtuse, punctation rather sparse and superficial, ground with isodiametric microsculpture, pubescence extremely short, scarcely visible, depressions near posterior angles wide, extending in front of middle of lateral margin. Elytra scarcely elongate (ratio length from scutellum to apex/combined width of elytra = 1.1), truncate at apex in both sexes, scarcely widened towards apex, punctation stronger than on pronotum, dense, rather confluent, ground glossy, pubescence extremely short, scarcely visible. Abdomen rather glossy, microsculpture superficial, with decumbent pubescence. Tibiae straight in both sexes, middle tibiae of male slightly curved internally at apex, tarsomere 5 of posterior tarsi slightly shorter than 1-4 together. Aedeagus of the type (California) as in Fig. 142, with a large copulatory sclerite, divided in 2 branches in the basal portion and curved in lateral view. In the Northern populations, from British Columbia to Oregon, the copulatory sclerite is somewhat longer. Accessory sclerites of female as in Fig. 143, spermatheca as in Fig. 144. Comparative notes. Eusphalerum newtoni is similar to E. pothos, from which the less defined median groove on the pronotum, which is more convex in the males, is somewhat distinctive. Eusphalerum parvispiculum is even more similar, and only the shape of the aedeagus and especially the presence of a characteristic large copulatory sclerite allows sure identification. Distribution. CANADA: British Columbia; UNITED STATES: Oregon, California (Map 7). Natural history. The species was documented at low altitude, up to 625 m a.s.l.. Some records are from mixed coniferous forest, one from Alnus forest. Known host plants: Tiarella trifoliata L. (Saxifragaceae), Clintonia uniflora (Menz. ex Sch. & Sch.) Kunth (Liliaceae), Cornus canadensis L. (Cornaceae), Heracleum maximum Bartram (= H. lanatum) (Apiaceae), Rubus parviflorus Nutt. (Rosaceae), and unidentified Asteraceae. Captures from May (California) to August (Washington)., Published as part of Zanetti, Adriano, 2014, Taxonomic revision of North American Eusphalerum Kraatz, 1857 (Coleoptera, Staphylinidae, Omaliinae), pp. 1-80 in Insecta Mundi 2014 (379) on pages 55-56, DOI: 10.5281/zenodo.5179446

Eusphalerum klimaszewskii Zanetti, n. sp. Material examined. (21 specimens) Holotype m 1 paratype m 1 f 1 spec. incomplete, British Columbia Mt. Cain 50 o 13’N 126 o 22’W 23.06/ 7.07.1997 leg. N. Winchester (MCSNV). Other paratypes: CANADA. British Columbia 2 mm 1 f Mt. Cain 50 o 13’N 126 o 21’W 23.05/ 9.06.1997 leg. N. Winchester (cZan); 3 ff Mt. Cain 50 o 13’N 126 o 22’W 23.05/ 9.06.1997 leg. N. Winchester (cZan); 3 spec. incomplete Mt. Cain 50 o 13’N 126 o 22’W 9.06/ 23.06.1997 leg. N. Winchester (cZan); 1 spec. incomplete Mt. Cain 50 o 13’N 126 o 22’W 23.05/ 9.06.1997 leg. N. Winchester (cZan); 1 m 1 f Mt. Cain 50 o 15’N 126 o 25’W 23.05/ 9.06.1997 leg. N. Winchester (cZan); 1 f Mt. Cain 50 o 15’N 126 o 25’W 9.06/ 23.06.1997 leg. N. Winchester (cZan); 1 f Mt. Cain 50 o 15’N 126 o 25’W 23.06/ 7.07.1997 leg. N. Winchester (cZan); 1 f Aberni (38 mi. W) 24/ 25.05.1968 leg. Campbell & Smetana (CNC); 2 ff Carmanah Valley (upper) UTM 10UCJ 802998 21.06/ 3.07.1991 leg. N. Winchester (FMNH). Measurements. head length: 0.33-0.35; head width: 0.64-0.66; pronotal length: 0.64-0.65; pronotal width: 0.75-0.81; elytral length: 1.44-1.48; elytral width: 1.29-1.33; length (clypeus to apex of elytra): 2.36-2.70; total length: 2.7-2.9. Etymology. The species is dedicated to Jan Klimaszewski, specialist of Staphylinidae, who provided me most specimens of the type series. Description. Habitus as in Fig. 14. Head, pronotum, abdomen, and ventral surface blackish, sometimes brown; elytra yellowish to brown, always lighter than rest of body; legs, antennae, and mouthparts yellowish, antennae sometimes very feebly darkened at apex. Head with strongly prominent eyes, postocular carina extremely reduced, almost absent, temples very short, strongly convergent caudad, medial margin of eyes without longitudinal wrinkles, microsculpture formed by elongate meshes. Clypeus prolonged anterad. Postantennal depressions not much impressed, slightly confluent with the small tentorial pits. Neck separeatd from head. Punctation very dense and rather strong, somewhat confluent, ground with superficial microsculpture. Antennae elongate, all antennomeres longer than wide, 4 clearly narrower than 5, 11 almost ovoid, more than twice as long as wide. Pronotum scarcely transverse (ratio width/length = 1.2 on average), impressed medially in front of posterior margin, convex, widest in front of middle, somewhat narrowed anterad, anterior margin clearly shorter than posterior, lateral margins rounded in anterior two thirds, convergent caudad in almost straight line, posterior angles marked and slightly obtuse. Punctation very dense and rather coarse, ground glossy with very superficial microsculpture, pubescence very long, whitish, depressions near posterior angles narrow, extended in front of middle of lateral margins. Elytra scarcely elongate (ratio length from scutellum to apex / combined width of elytra = 1.1), scarcely widened towards apex, truncate in middle in both sexes, punctation coarser and somewhat less dense than on pronotum, confluent on glossy ground, pubescence long, whitish and decumbent. Abdomen dull, microsculpture clearly visible, formed by almost isodiametric meshes, pubescence long, decumbent. Tibiae straight in both sexes, not modified, tarsomere 5 of posterior tarsi somewhat shorter than 1- 4 together. Aedeagus as in Fig. 15. Accessory sclerites of female as in Fig 16, spermatheca as in Fig. 17. Comparative notes. Dark color, long white pubescence, rather narrow pronotum, and long yellowish antennae make this species easily recognizable among North American Eusphalerum. The related species of the subangulatum group are light and have a wider pronotum. Distribution. CANADA: British Columbia (Map 4), known only from Vancouver Island. Natural history. All specimens were collected in forest sites. No information about host plants is available, as specimens were collected mostly by traps. Captures between May and beginning of July., Published as part of Zanetti, Adriano, 2014, Taxonomic revision of North American Eusphalerum Kraatz, 1857 (Coleoptera, Staphylinidae, Omaliinae), pp. 1-80 in Insecta Mundi 2014 (379) on pages 10-11, DOI: 10.5281/zenodo.5179446

Eusphalerum orientale (Bernhauer, 1912) Anthobium orientale Bernhauer, 1912: 678; Scheerpeltz 1933: 1036; Hatch 1944: 101. Eusphalerum orientale (Bernhauer); Moore and Legner 1975: 191; Herman 2001: 444. Anthobium frosti Bernhauer, 1928: 40 n. syn.; Scheerpeltz, 1933: 1033. Eusphalerum frosti (Bernhauer); Moore and Legner 1975: 190., Published as part of Zanetti, Adriano, 2014, Taxonomic revision of North American Eusphalerum Kraatz, 1857 (Coleoptera, Staphylinidae, Omaliinae), pp. 1-80 in Insecta Mundi 2014 (379) on page 41, DOI: 10.5281/zenodo.5179446, {"references":["Bernhauer, M. 1912. Zur Staphylinidenfauna von Nordamerica. V. Beitrag. Pomona College Journal of Entomology 4 (1): 678 - 683.","Scheerpeltz, O. 1933. Staphylinidae VII. Coleopterorum Catalogus 6 (129): 989 - 1500.","Hatch, M. H. 1944. Studies on Northwestern Staphylinidae I. Anthobium Steph. The Pan-Pacific Entomologist 22 (3): 101 - 104.","Moore, I., and E. F. Legner. 1975. A catalogue of Staphylinidae of America North of Mexico (Coleoptera). Division of Agricultural Sciences, University of California Special Publication 3015: 1 - 514.","Herman, L. H. 2001. Catalog of the Staphylinidae (Insecta: Coleoptera). 1758 to the end of second millennium. I. Introduction, history, biographical sketches, and Omaliinae group. Bulletin of the American Museum of Natural History 265: 1 - 650.","Bernhauer, M. 1928. Ein neues Anthobium aus Nordamerica (Col. Staph.) Deutsche entomologische Zeitschrift 1928: 40."]}

Eusphalerum luteipes Zanetti n. sp. Material examined (6 specimens) Holotype m 2 paratypes mm 2 paratypes ff California Ventura Co Murietta Trail LPNF 34.5009 oN 119.3899 oW 26.03.2006 leg. Caterino (SBMNH, 1 paratype cZan). Other paratypes. USA. California 1 m 1 f (FMNH) [identified as A. lutipes Casey (recent label), manuscript name]. Doubtful identification: 2 ff G.P. Mackenzie, Mt. Wilson 2.05.1942 (FMNH). Measurements. Head length: 0.20-0.24; head width: 0.46-0.49; pronotal length: 0.38-0.44; pronotal width: 0.59-0.64; elytral length: 0.79-0.88; elytral width: 0.75-0.81; length (clypeus to apex of elytra): 1.38-1.51; total length: 1.7-2.1. Etymology. The name means with yellow legs (luteus = golden yellow, pes = foot in Latin). It is reported as “ lutipes ” (manuscript name of Casey) on a label of an ancient specimen (FMNH), and it has been maintained, though with a spelling correction. Description. Habitus as in Fig. 109. Head, pronotum and elytra brown; abdomen dark brown; ventral thoracic surface dark brown; legs yellowish; antennae yellowish, darkened from antennomere 7, or entirely yellowish. Head with prominent eyes, postocular carina evident, temples strongly convergent caudad in straight line, medial margin of eyes with microsculpture tending to form longitudinal wrinkles. Surface of head scarcely impressed, postantennal depressions and tentorial pits superficial, confluent in 2 longitudinal depressed areas. Neck not separated from the head. Punctation rather dense and strong, ground glossy almost without superficial microsculpture on the disc. Antennae rather long, antennomere 1 and 2 ovoid elongate, 3 twice as long as wide, 4-6 longer than wide, 7 subquadrate, 8-10 transverse, 11 twice as long as wide, conical at apex. Pronotum transverse (ratio width/length = 1.5 on average), convex, widest in the middle, anterior margin slightly shorter than posterior, lateral margins strongly rounded, posterior angles obtuse and scarcely marked. Punctation dense and incised and superficial on glossy ground with very superficial microsculpture, pubescence very short scarcely visible, depressions near posterior angles superficial, extending in front of middle of lateral margins. Elytra scarcely elongate (ratio length from scutellum to apex / combined width of elytra = 1.0), scarcely widened towards apex, punctation coarse and dense, somewhat confluent on glossy ground, pubescence very scarcely visible. Abdomen rather glossy, microsculpture superficial, pubescence decumbent. Male with widened mesofemora and mesotibiae curved and sunken in apical half, widened in an acute internal tip (Fig. 113), tarsomere 5 of posterior tarsi longer than 1-4 together. Aedeagus as in Fig. 110. Accessory sclerites of female reduced (Fig. 111), scarcely visible, spermatheca as in Fig. 112. Comparative notes. The small size, entirely dark color, presence of postocular carina (though not marked), absence of pronotal microsculpture, and especially shape of male middle tibiae are distinctive. Eusphalerum luteipes is similar to E. caterinoi, but in the latter the male middle tibiae are unmodified, the pronotum is distinctly microsculptured, and the aedeagus is different (Fig. 103 vs. 110) in shorter parameres with less numerous setae and in internal structures. Distribution. UNITED STATES: California (southern) (Map 8). Natural history. The types were collected at low altitude, in a chaparral habitat, early in spring (March)., Published as part of Zanetti, Adriano, 2014, Taxonomic revision of North American Eusphalerum Kraatz, 1857 (Coleoptera, Staphylinidae, Omaliinae), pp. 1-80 in Insecta Mundi 2014 (379) on pages 37-38, DOI: 10.5281/zenodo.5179446

Eusphalerum diversicolle (Casey, 1894) Anthobium diversicolle Casey, 1894: 428; Bernhauer and Schubert 1910: 39. Eusphalerum diversicolle (Casey); Moore and Legner 1975: 428; Herman 2001: 423., Published as part of Zanetti, Adriano, 2014, Taxonomic revision of North American Eusphalerum Kraatz, 1857 (Coleoptera, Staphylinidae, Omaliinae), pp. 1-80 in Insecta Mundi 2014 (379) on page 20, DOI: 10.5281/zenodo.5179446, {"references":["Casey, T. L. 1894. Coleopterological notices. V. Annals of the New York Academy of Sciences 7: 281 - 606.","Bernhauer, M., and K. Schubert. 1910. Staphylinidae I. In S. Schenkling, Coleopterorum Catalogus, 5 (19): 1 - 86. Junk; Berlin.","Moore, I., and E. F. Legner. 1975. A catalogue of Staphylinidae of America North of Mexico (Coleoptera). Division of Agricultural Sciences, University of California Special Publication 3015: 1 - 514.","Herman, L. H. 2001. Catalog of the Staphylinidae (Insecta: Coleoptera). 1758 to the end of second millennium. I. Introduction, history, biographical sketches, and Omaliinae group. Bulletin of the American Museum of Natural History 265: 1 - 650."]}

Eusphalerum uncinatum Zanetti n. sp. Material examined (135 specimens) Holotype m 5 paratypes mm 7 ff Oregon Clackamas Co Camp Crk. (3.5 mi. SE) 2300-2400' Rhododendron 27.06.1974 leg. A. & D. Smetana (CNC) Other paratypes. CANADA. British Columbia 18 mm 18 ff Manning Pr. Pk Rhododendron Flats Rhododendron flowers 21.06.1988 leg. J.M. Campbell (CNC); 6 mm 5 ff Manning Prov. Pk., Rhododendron Trail sweep Rhododendron flowers 20.06.1988 leg. L. LeSage (CNC); 2 mm Mt. Revelstoke 6000 25.07.1972 leg. G.J. Spencer (CNC); 1 m Silver Star Prov. Pk. 6200 8.08.1973 leg. R.H. Parry (CNC). USA. California 10 m 33 ff El Dorado Co Fallen Leaf 6500 1/ 29.07.1935 leg. F.E. Blaisdell (CNC); 1 m El. Dorado Co Fallen Leaf 21.06.1915 leg. R.Hopping (CNC); 3 mm 2 ff Fresno Co Huntington Lake swept 3.08.1962 leg. C. D. Johnson (FMNH); 6 mm 3 ff Fresno Co Sierra NF John Muir Tr 37.2583 oN 118.8654 oW 20.08.2006 leg. Caterino (SBMNH); 1 m 1 f Placer Co Lake Tahoe, Tahoe Pines 6200 10.08.1969 leg. A. Smetana (CNC); 2 mm Tehama Co Wilson Lake 5300 27.06.1966 leg. C.W. OBrien (CNC); 2 f 1 m Lake Tahoe 23.06.1925 leg. E.H.Nast (CNC); 2 mm Fresno Co Huntington Lake 3.08.1962 leg. C.D. Johnson (FMNH); Oregon 1 m Clackamas Co jct US26 Ore 35 (1.3 m E) 3900 on flowers Lupinus 12.07.1975 leg. A. Newton M. Thayer (FMNH); 1 m 1 f Douglas Co Lamola Lk. (3.5 mi. S), Umpqua Nat. For. 4400' currant, Pinus contorta, rotted wood 22.06.1972 leg. E.H. Benedict (CNC); Washington 2 mm 2 ff Chelan Co Leavenworth (10 km NW) 30.06.1984 leg. R. Baranowski (LMZ). Measurements. head length: 0.31-035; head width: 0.53-0.62; pronotal length: 0.46-0.55; pronotal width: 0.72-0.83; elytral length: 0.98-1.14; elytral width: 0.99-1.18; length (clypeus to apex of elytra): 1.66-2.23; total length: 2.1-3.1. Etymology. The adjective uncinatum means hooked in Latin. It refers to the form of median lobe of the aedeagus in lateral view. Description. Habitus as in Fig. 149. Head and pronotum reddish yellow; neck sometimes somewhat darkened in male; elytra yellowish;, abdomen brown, somewhat paler at apex (male) or entirely yellowish (female); prosternum yellowish; metasternum brown, sometimes yellowish in female; legs yellowish; antennae yellowish, more or less darkened from antennomere 6, often entirely yellowish. Head with prominent eyes; postocular carina well marked; temples short, convergent caudad. Medial margin of eyes with microsculpture forming longitudinal wrinkles; postantennal depressions deep, confluent with tentorial pits, forming 2 longitudinal depressions in front of ocelli; punctation superficial and sparse; ground with microsculpture isodiametric in middle and on neck, longitudinal in depressions in front of ocelli; neck wide, not separated from head. Antennae robust, antennomere 1 and 2 ovoid, 3 elongate, twice as long as wide, 4, 5, and 6 subquadrate, 9-10 transverse, 10 almost twice as wide as long, 11 twice as long as wide, cylindrical in the proximal part, conical at apex. Pronotum convex, often with a small impression medially near anterior margin, transverse (ratio width/length = 1.4-1.6), widest in middle, anterior margin slightly narrower than posterior, lateral margins usually rounded, sometimes almost rectilinear, posterior angles scarcely obtuse and well marked, punctation sparse and rather superficial, ground with isodiametric microsculpture, pubescence extremely short, scarcely visible, depressions near posterior angles rather wide. Elytra scarcely elongate (ratio length from scutellum to apex / combined width of elytra = 1.0), truncate at apex in both sexes, moderately widened toward apex, punctation stronger than on pronotum, irregular and rather confluent, ground glossy, pubescence extremely short and scarcely visible. Abdomen rather glossy, microsculpture superficial with decumbent pubescence. Tibiae straight in both sexes, not modified, tarsomere 5 of posterior tarsi slightly shorter than 1-4 together. Aedeagus as in Fig. 150, hooked near apex in lateral view, without large copulatory sclerites. Accessory sclerites of female as in Fig. 151, spermatheca as in Fig. 152. Comparative notes. Eusphalerum uncinatum is similar to E. pothos, from which the absence of a defined median groove on pronotum is somewhat distinctive. Eusphalerum newtoni and E. parvispiculum are similar, too, and only the shape of the aedeagus, especially the presence of a hook near the apex, visible in lateral view, is distinctive for E. uncinatum. Distribution. CANADA: British Columbia; UNITED STATES: California, Oregon, Washington (Map 9)., Published as part of Zanetti, Adriano, 2014, Taxonomic revision of North American Eusphalerum Kraatz, 1857 (Coleoptera, Staphylinidae, Omaliinae), pp. 1-80 in Insecta Mundi 2014 (379) on pages 57-58, DOI: 10.5281/zenodo.5179446

Eusphalerum chatzimanolisi Zanetti n. sp. Material examined (2 specimens) Holotype m 1 paratype m California Fresno Co Sequoia NF FS 13 S21 36.8105 oN 118.9670 oW 8.06.2006 leg. Caterino & Chatzimanolis (SBMNH) Measurements. head length: 0.25-0.27; head width: 0.48-0.51; pronotal length: 0.44-0.46; pronotal width: 0.64-0.70; elytral length: 1.20-1.33; elytral width: 0.96-10.3; length (clypeus to apex of elytra): 1.85-1.97; total length: 2. Etymology. The species is dedicated to one of its collectors, Stylianos Chatzimanolis, student of systematics of Staphylinidae. Description. Habitus as in Fig. 93. Head, pronotum, and elytra yellowish, elytra somewhat paler, neck somewhat darkened; abdomen brown (male); prosternum yellowish; metasternum pale brown; legs, antennae, and mouthparts yellowish, antennae strongly darkened from antennomere 6. Head with prominent eyes, postocular carina absent, temples short, strongly convergent caudad, medial margin of eyes microsculptured without longitudinal wrinkles. Head rather flat, postantennal depressions very superficial, tentorial pits small but well impressed. Neck not separated from head medially. Punctation rather sparse, superficial, ground with isodiametric microsculpture. Antennae elongate, antennomere 1 and 2 segments twice as long as wide, 3 long and thin, three times as long as wide, 4 -5 twice as long as wide, 6 and 7 longer than wide, 8 -10 wider than long, 11 twice as long as wide, ovoid at base and conical at apex. Pronotum transverse (ratio width/length = 1.5), convex, widest just in front of middle, anterior margin much narrower than posterior, lateral margins strongly convergent anteriad, rounded in middle, and scarcely convergent caudad in almost straight line, posterior angles marked and scarcely obtuse. Punctation very sparse, superficial, and irregular, ground with isodiametric microsculpture, pubescence very short, scarcely visible, depressions near posterior angles scarcely impressed. Elytra strongly elongate (ratio length from scutellum to apex / combined width of elytra = 1.2), scarcely widened towards apex, rounded at apex, punctation much denser and stronger than on pronotum, confluent on glossy ground, pubescence very short, almost invisible. Tibiae straight, not modified, tarsomere 5 of posterior tarsi shorter than 1-4 together. Sternite VII emarginate medially as in Fig. 95. Aedeagus as in Fig. 94 (internal sac partly extruded). Female unknown. Comparative notes. Eusphalerum chatzimanolisi is very similar to E. convexum and E. fraternum in external characters: only the very sparse pronotal punctation is distinctive. The shape of the posterior margin of abdominal sternite VIII of the male is also different, recalling E. carolinensis, which differs in having the posterior half of the lateral margins of the pronotum convergent. The shape of the internal sac of the aedeagus is also characteristic. Distribution. UNITED STATES: California (Map 1). Natural history. The type specimens were collected in June in forest at about 1800 m a.s.l. (altitude inferred from coordinates), host plants not known., Published as part of Zanetti, Adriano, 2014, Taxonomic revision of North American Eusphalerum Kraatz, 1857 (Coleoptera, Staphylinidae, Omaliinae), pp. 1-80 in Insecta Mundi 2014 (379) on pages 31-32, DOI: 10.5281/zenodo.5179446