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Summary Juniperus spp. are keystone shrubs in western North America and important climatic indicators in paleo‐records. However, a lack of taxonomic resolution among fossil species limits our ability to track past environmental changes. Plant macrofossils at Rancho La Brea (RLB) allow for reconstructions of juniper occurrence to species across 60 000 yr. We use microscopy, image analysis, species distribution modeling (SDM), and radiocarbon dating to identify an unknown Juniperus species at RLB and put it into chronological context with fossil Juniperus californica at the site to infer past environmental conditions. We identify the unknown taxon as Juniperus scopulorum Sargent, 1897. The Pleistocene occurrence of this species in California expands its known distribution and documents its extirpation. Temporal ranges of the two fossil junipers alternate, revealing a pattern of differential climatic sensitivity throughout the end of the Pleistocene. Occurrence patterns suggest sensitivity to temperature, moisture availability, and the presence of two mega‐droughts at c. 48–44.5 ka and c. 29.3–25.2 ka. Extirpation of both taxa by c. 13 ka is likely driven by climate, megafaunal extinction, and increasing fire. The extirpation of fossil junipers during these past climatic events demonstrates vulnerability of juniper species in the face of global change. [ABSTRACT FROM AUTHOR]

Tenebrionoids are not uncommon in late Mesozoic ambers, but their paleodiversity remains poorly explored, partly due to the insufficiently defined families of the extant fauna. Here, we describe and illustrate a new tenebrionoid beetle from mid‐Cretaceous Kachin amber, Glyphonotum hsiaoi gen. et sp. nov., which exhibits an unusual form of pronotal sulcus and tarsi. Based on morphological comparison and phylogenetic analyses, we tentatively place Glyphonotum in the extant family Pythidae, representing the first Mesozoic fossil of this family. [ABSTRACT FROM AUTHOR]

A new species of Ulnaria (Kützing) Compère is described from the Miocene Trout Creek fossil locality, situated in Harney County, Oregon, USA. The rock material from which the new species is described consisted of a subsample taken from the matrix harboring a plant fossil collected in 1932 or 1933 and archived at the Field Museum of Natural History, Chicago, Illinois, USA. The new species differs from all other taxa in the genus by possessing club-shaped valves that are asymmetric about the transverse axis, and frustules that are cuneate-shaped in girdle view. In addition to its unique shape, valves of the new species have a well-developed ocellulimbus (pore field) at each apex, a single rimoportula at one or both poles and closed girdle bands. The virgae are thick relative to the vimines, areolae possess exterior closing plates, and on larger specimens, the uniseriate striae can become biseriate near the valve margins. Material at the Trout Creek locality was deposited approximately 15.6–14 Ma during the Middle Miocene Climatic Optimum (MMCO). Given the diatom species co-occurring with the new Ulnaria species, it likely inhabited the plankton. [ABSTRACT FROM AUTHOR]

The Messel Pit is a Konservat-Lagerstätte in Germany, representing the deposits of a latest early to earliest middle Eocene maar lake, and one of the first palaeontological sites to be included on the list of UNESCO World Heritage Sites. One aspect of Messel that makes it so extraordinary is that its sediments are rich in different fossilised organisms – microfossils, plants, fungi, invertebrate animals and vertebrates – that are rarely preserved together. We present an updated list of all taxa, named or not, that have been documented at Messel, comprising 1409 taxa, which represent a smaller but inexactly known number of biological species. The taxonomic list of Labandeira and Dunne (2014) contains serious deficiencies and should not be used uncritically. Furthermore, we compiled specimen lists of all Messel amphibians, reptiles and mammals known to us. In all, our analyses incorporate data from 32 public collections and some 20 private collections. We apply modern biodiversity-theoretic techniques to ascertain how species richness tracks sampling, to estimate what is the minimum asymptotic species richness, and to project how long it will take to sample a given proportion of that minimum richness. Plant and insect diversity is currently less well investigated than vertebrate diversity. Completeness of sampling in aquatic and semiaquatic, followed by volant, vertebrates is higher than in terrestrial vertebrates. Current excavation rates are one-half to two-thirds lower than in the recent past, leading to much higher estimates of the future excavation effort required to sample species richness more completely, should these rates be maintained. Species richness at Messel, which represents a lake within a paratropical forest near the end of the Early Eocene Climate Optimum, was generally higher than in comparable parts of Central Europe today but lower than in present-day Neotropical biotopes. There is no evidence that the Eocene Messel ecosystem was a "tropical rainforest." [ABSTRACT FROM AUTHOR]

Two new extinct species of false skin beetles (Biphyllidae) are described and illustrated based on inclusions in Baltic amber: Biphyllus lar sp. nov. and Coelodiplus uuu-impressa gen. et sp. nov. The new Eocene fossils are compared with the related extant and extinct representatives of the family. A new monotypic genus is erected for one fossil; another Eocene species is classified within the extant genus Biphyllus Dejean, 1821. The holotype of Diplocoelus probiphyllus Vitali, 2010 (Baltic amber) is re-examined and its diagnosis is emended. The present paper brings the total number of described biphyllid species from European succinite to three. A key to all documented fossil species of the family is provided. Several aspects of insect use as paleoenvironment and climate proxies are briefly discussed. On the basis of the revealed biphyllid palaeofauna and analysis of the climatic extremes of the modern distribution of the family in the Northern Hemisphere, the lowest possible annual temperature in the area of amberiferous forests of Eocene Northern Europe is hypothesised. [ABSTRACT FROM AUTHOR]

The mid-Pliocene Upper Laetolil Beds at Laetoli in northern Tanzania have produced a rich diversity of fossil vertebrates, including the remains and traces of the early hominin, Australopithecus afarensis. The fauna from the older Lower Laetolil Beds, dating from 3.8 Ma to older than 4.3 Ma, has not previously been the subject of detailed analysis. This study, based on renewed palaeontological investigations, provides the first systematic account of the fauna, along with a brief synthesis of the geology and palaeoecology. Forty species of mammals are recorded from the Lower Laetolil Beds, dominated by bovids, equids, rhinocerotids, and proboscideans, in addition to reptiles, birds, and invertebrates. The mammal fauna is very similar to that from the Upper Laetolil Beds, and to a lesser extent to the Upper Ndolanya Beds, indicating a high degree of faunal continuity and provinciality in the Laetoli area during much of the Pliocene. Evidence indicates that the palaeoecology of the Lower Laetolil Beds was predominantly dry open woodlands-shrublands-grasslands, but dense woodlands and forests were present on the lower slopes of the nearby Satiman volcano. Water was more abundant than in the Upper Laetolil Beds, with seasonal rivers, marshes and shallow lakes bordered by riparian woodland. [ABSTRACT FROM AUTHOR]

The present study describes two occurrences of clam shrimps ("Conchostraca"). Following a formbased classification of Schneider et al. (2005, 2022) and Schneider & Scholze (2018), the conchostracans have been classified as Pseudestheria form Oberbozen and Pseudestheria form Le Fraine. They come from the Guncina and Tregiovo formations, repectivey. These units are part of the northern Athesian Volcanic Group, North Italy. All of this material were obtained from grey coloured, fine-grained siliciclastic, lacustrine deposits of Kungurian (late Cisuralian, late Early Permian) age. Their carapace valves in lateral view are marked by concentric ribs. Often, growth lines can be observed to decrease in their preservation towards the umbonal area of the valve. An open nomenclature is applied to the taxonomy at the species level herein, because of either limited number of individuals or a plastic deformation due to tectonics. So far, the records reported herein are restricted geographically and stratigraphically to respective single occurrences. Further studies are recommended to establish their full biostratigraphic ranges. [ABSTRACT FROM AUTHOR]

Neuroptera, as a small relic group of Insecta undergoing a rapid species diversification during the Mesozoic Era, is known by diverse extinct endemic lineages preserved as impression fossils and in amber. The current understanding of Mesozoic neuropterans′ diversity has mainly focused on the adults, because the contemporaneous larvae have been fairly rare especially for the Jurassic lacewings. Herein, a new giant lacewing larva, Natator giganteus gen. et sp. nov., is described from the Middle Jurassic Daohugou Beds of China. The remarkable larva is characterized by its impressively large body size, distinctively elongated cervix, and presence of swimming hairs on legs, which provide direct evidence to reveal an aquatic habit for the Jurassic lacewing larva. The morphological analysis indicates this giant larva would have probably inhabited the benthic environments of Jurassic montane rivers and streams. In addition, its morphological specialization suggests that it might have adopted an ambush predation strategy to catch its prey. The finding enhances our knowledge of the species diversity and morphological plasticity for the Jurassic lacewing larvae, and reveals that the aquatic lineages of Neuroptera exhibited dramatically structural and ecological convergence across the evolutionary process. [ABSTRACT FROM AUTHOR]

Pterocarya (Juglandaceae) is disjunctly distributed in East Asia and the Caucasus region today, but its fossils are widely distributed in the Northern Hemisphere. We first inferred phylogeny with time estimation of Pterocarya under node‐dating (ND) based on plastomes of all eight extant species and tip‐dating (TD) based on plastomes and 69 morphological characters of 19 extant and extinct species, respectively. We compared the biogeographical reconstructions on the timetrees from ND and TD, respectively, and then compiled 83 fossil records and 599 current occurrences for predicting the potential distributions for the past and the future. The most recent comment ancestor of Pterocarya is inferred in East Asia at 40.46 Ma (95% highest posterior density [HPD]: 28.04–54.86) under TD and 26.81 Ma (95% HPD: 23.03–33.12) under ND. The current distribution was attributed to one dispersal and one vicariant event without fossils, but as many as six dispersal, six vicariant, and 11 local extinction events when considering fossils. Pterocarya migrated between East Asia and North America via the Bering Land Bridge during the early Oligocene and the early Miocene periods. With the closure of Turgai Strait, Pterocarya dispersed between East Asia and Europe through the Miocene. The potential distribution analyses indicated that Pterocarya preferred warm temperate regions across the Northern Hemisphere since the Oligocene, but the drastic temperature decline caused its extinction in high latitudes. Except for Pterocarya fraxinifolia and Pterocarya stenoptera, suitable habitats for this genus are predicted to contract by 2070 due to climate change. [ABSTRACT FROM AUTHOR]

Fruits of the caryophyllalean family Montiaceae are recognized from the latest Cretaceous Deccan Intertrappean beds of central India. Although initially interpreted as the sporocarp of an aquatic fern, Kuprianovites Nambudiri & Thomas was subsequently demonstrated to belong to the group Centrospermeae. The name Kuprianovites deccanensis Nambudiri & Thomas takes priority over subsequent synonyms, Centrospermocarpon chitaleyae Sheikh & Kubalkar, Deccanocarpon arnoldii Paradkar, and Portulacaceocarpon bhuterensis Borkar, Nagrale, Meshram, Korpenwar, & DD Ramteke. We studied anatomically preserved specimens using successive acetate peels and micro–computed tomography scanning. The fruits are thin-walled trivalved globose capsules bearing 30 or more seeds in free-central placentation. The seeds are ellipsoidal and bear strongly curved embryos. Although formerly assigned to the Portulacaceae as that family was traditionally circumscribed, the fossil is excluded from the Portulaca clade (i.e., Portulaceaceae s.s.) and conforms instead to the Montiaceae by its valvate, rather than circumscissile, capsules and valves with undifferentiated cell layers. [ABSTRACT FROM AUTHOR]

A remarkable new genus and species of cyclostome bryozoans is described from the Sea of Okhotsk off the Kamchatka Peninsula, Russia. Kamchatkapora ozhgibesovigen. et sp. n. has large, multilayered colonies, each layer constructed of numerous subcolonies joined at their outer edges. This colony-form was previously unknown in extant cyclostomes, although common in the geological past, especially during the Cretaceous period. The term 'stratocormidial' is here introduced for such colonies, and their taxonomy and geological history are reviewed. Six fossil stratocormidial cyclostome genera (Blumenbachium, Cellulipora, Centronea, Multifascigera, Reptomulticava and Semimulticavea) are illustrated and briefly characterised. [ABSTRACT FROM AUTHOR]

The first fossil rust fly (Diptera: Psilidae) from the Insect Bed of the Bembridge Marls of the Isle of Wight is also the first known fossil record of the subfamily Chylizinae. It is late Eocene: Priabonian in age, 34.2 Ma. Chyliza colenutti Ross, Zhou, Hoffeins & Crighton, sp. nov. is compared with 30 extant species of Chyliza and differs from them in having a relatively long 2nd basal cell compared to the discal cell. Some other fossil Acalyptratae records are reviewed and their probable ages updated. The phylogenetic position of the Psilidae is currently uncertain so the family may have appeared anytime from 70–40 million years ago. [ABSTRACT FROM AUTHOR]

A morphologically unusual fossil true bug genus, Palaeotanyrhina Poinar, Brown & Kóbor, 2022, placed into Cimicomorpha: Reduvioidea by its authors, is revisited based on an inclusion in mid-Cretaceous Burmese amber. The original description is supplemented with additional details and corrections. It is demonstrated that the placement of Palaeotanyrhina into Cimicomorpha was based on misinterpretations of the morphology of the holotype, and the genus is a member of the infraorder Leptopodomorpha. A phylogenetic analysis is provided for fossil and extant taxa of Leptopodomorpha. Parvilepta Yu, Zhuo & Chen, 2023, CretaleptusSun & Chen, 2019, Macrolepta Yu, Zhuo & Chen, 2023 (currently members of Leptopodidae: Leptosaldinae) and PalaeoleptusPoinar, 2009 (the type genus of Palaeoleptidae) are demonstrated to be phylogenetically closely related to Palaeotanyrhina. All the above mentioned genera are recognized to represent stem-group members of Leptopodomorpha: Leptopodidae: Leptopodinae: Leptopodini, and accordingly, they are transferred to the latter tribe. The following new subjective synonyms are proposed: Leptopodidae: Leptopodinae: Leptopodini Brullé, 1836 = Palaeoleptidae Poinar, 2009, syn. nov. = Palaeotanyrhinidae Poinar, Brown & Kóbor, 2022, syn. nov. Based on the analytic results, the subfamilies Leptosaldinae and Leptopodinae of Leptopodidae and the tribes Leotichiini and Leptopodini of Leptopodinae are considered valid, redefined, and a list of their included genera and species are provided. Arguments are provided in support of the placement of Leptosaldinea Popov & Heiss, 2016, formerly placed into Leptosaldinae, into the infraorder Dipsocoromorpha. [ABSTRACT FROM AUTHOR]

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Pseudoscorpions, one of the earliest terrestrial lineages dating back to the Middle Devonian period (c. 385 Ma), have a limited fossil record mainly as inclusions in amber. The available fossils are mostly from the Cenozoic era, making it challenging to comprehensively understand their evolutionary history and diversification over time. Here, we report the first fossils of the order from Spanish Lower Cretaceous (upper Albian) ambers. Two new genera and species in the family Garypinidae are described from Álava amber (Peñacerrada II outcrop, Álava Province): Cretogarypinus zaragozai gen. et sp. nov. and Ithioreolpium alavensis gen. et sp. nov. Remarkably, the holotype of Cretogarypinus zaragozai shows preservation of internal soft tissues. The variability observed in the arolium of both fossil garypinids serves as compelling evidence for the hypothesis of diphyletism within the clade, which was already well established before the Early Cretaceous. Additionally, an incomplete specimen consisting of an isolated palp ascribed to the family Pseudogarypidae (extant genus Pseudogarypus Ellingsen), is described from San Just amber (Teruel Province). These findings represent the earliest occurrences of the families Garypinidae (superfamily Garypinoidea) and Pseudogarypidae (superfamily Feaelloidea) in the fossil record. They not only contribute to our understanding of a diverse pseudoscorpion fauna in Spanish amber but also offer valuable insights into divergence time estimations for pseudoscorpion families and potentially for biogeographic studies. [ABSTRACT FROM AUTHOR]

Abstract The click beetles (Elateridae) represent the major and well-known group of the polyphagan superfamily Elateroidea. Despite a relatively rich fossil record of Mesozoic Elateridae, only a few species are described from the Upper Cretaceous Burmese amber. Although Elateridae spend most of their lives as larvae, our knowledge on immature stages of this family is limited, which is especially valid for the fossils. So far, only a single larval click beetle has been reported from Burmese amber. Here, we describe two larval specimens from the same deposit which based on their morphology unambiguously belong to the predominantly Southern Hemisphere subfamily Pityobiinae, being the most similar to the representatives of tribe Tibionemini. However, since the larvae of the closely related bioluminescent Campyloxenini have not yet been described, we place our specimens to Tibionemini only tentatively. One species of Pityobiinae was recently described from Burmese amber based on adults, and we discuss if it can be congeneric with the here-reported larvae. Recent representatives of the Tibionemini + Campyloxenini clade are known from South America and New Zealand, and this group is hypothesized to have a Gondwanan origin. Hence, the newly discovered Burmese amber larvae may further contribute to a recently highly debated hypothesis that biota of the resin-producing forest on the Burma Terrane, which was probably an island drifting northward at the time of amber deposition, had at least partly Gondwanan affinities. The discovery of enigmatic click beetle larvae in the Upper Cretaceous Burmese amber sheds further light on the palaeodiversity and distribution of the relatively species-poor Gondwanan clade of click beetles, which contain a recent bioluminescent lineage, as well as on the taxonomic composition of the extinct Mesozoic ecosystem.

Anatomically preserved material from Lower Devonian strata of the Battery Point Formation (Gaspé Peninsula, Quebec, Canada) offers a rare opportunity to reconstruct the sequence of events in the life of a Nebuloxyla mikmaqiana plant (early euphyllophyte) that led to wounding and the plant’s recovery after wounding. Using serial sections from cellulose acetate peels of the Nebuloxyla axis and volume renderings of the affected tissues, we show that the wound, centered around a branch and caused likely by an herbivore, resulted in removal of the extraxylary tissues of the branch starting from its very base, associated with bending of the xylem strand that supplied the branch. Downward bending of the branch xylem toward the base of the main axis led to the separation and displacement of a significant portion of it, leaving a short portion of the xylem strand at the base of the branch in its original position. The displaced strand of branch xylem, after stripping of the extraxylary tissues along with some of its own thickness, was left protruding outwards from the open wound surface. Subsequent development of wound response tissue filled the gap left by the wound around the protruding displaced xylem strand, surrounding it with wound periderm. This specimen provides evidence that plants as old as Nebuloxyla had the capacity to endure and survive traumatic events, and to deploy effective and sophisticated response mechanisms that had already evolved in the euphyllophyte clade by Emsian time. More broadly, this occurrence re-emphasizes the importance of permineralized fossils in documenting details of plant anatomy and development, even allowing glimpses into minutia of their daily life, such as interactions with their environment and immediate responses to these interactions.

A stakeholder engagement workshop was held in May 2024 as part of the "Community-driven enhancement of information ecosystems for the discovery and use of paleontological specimen data" project, which is funded under the United States National Science Foundation (NSF) Geosciences Open Science Ecosystem (GEO OSE) program. This report describes the activites and outcomes of the workshop.

A reliable character system is crucial to taxonomy and systematics, and it promises valid downstream inferences, e.g., estimates of diversity and disparity, reconstruction of evolutionary history, and even stratigraphic correlations. Modern taxonomy and systematics of extant cockroaches requires an integrative study involving multiple lines of evidence with emphasis on genital and reproductive characteristics and molecular data. In contrast, many fossil cockroach taxa published recently are based solely on forewings. Many studies have shown that forewing-based taxa are questionable. In order to find out how much of the phylogenetic signal we could ascertain from venational similarity, and how confident we could be, this study used forewing characters to reconstruct phylogenies of the genera of well-recognized family-group taxa. The intuitively reconstructed phylogeny of 75 extant genera failed to recover those taxa or their relationships. Parsimony analyses of various datasets all yielded strong polyphyly and chaotic relationships. In conclusion, the forewing of cockroaches is not a universally competent character system. The underlying causes are the complicated nature of veins and the limitations of current analytical techniques. The uncertainty in forewing-based taxonomy and systematics has been underestimated in the literature. Forewing-based fossil taxa warrant re-evaluation; some of them are herein deemed nomina dubia in their current state.

\nHighlightsA new and intriguing fossil Mallomonas species, M. enigmata, was uncovered from 10 strata in the Eocene Giraffe Pipe locality. The new species represents an ancient member of the section Punctiferae, and based on scale and bristle morphology is most closely related to the modern congener, M. nieringii. M. nieringii is a rare species known from a suite of acidic waterbodies on the Cape Cod peninsula, Massachusetts, USA. Only a few modifications of the body scales differ between M. enigmata and M. nieringii, and the new taxon was uncovered from a section of the Giraffe Pipe maar lake representing a shallow and acidic waterbody. Cysts of M. enigmata are oval, with a small pore surrounded by a narrow hyaline zone. The anterior ~ ¼ of the cyst surrounding the hyaline zone consists of a series of small, closely spaced, rimmed holes that penetrate the wall. Otherwise, the cyst wall is smooth and unornamented. Unlike any known modern species in the genus, a second type of body scale is produced, but found only attached to the cyst. It is hypothesized that these scales may represent unfinished body scales that became fused to the cyst during its development. A new fossil Eocene species of Mallomonas.A unique second scale morphotype attached to the cyst.Thrived in a shallow, humic and acidic freshwater pond.A new fossil Eocene species of Mallomonas.A unique second scale morphotype attached to the cyst.Thrived in a shallow, humic and acidic freshwater pond. [ABSTRACT FROM AUTHOR]

Plants and arthropods are primary drivers of terrestrial ecosystem function. Trace fossils of plant-arthropod interactions (PAIs) provide a unique window into assessing terrestrial ecosystem states through geological time and evaluating changes in herbivorous arthropod feeding guilds in the wake of global biotic crises. The end-Permian event (EPE; c. 252 Ma) resulted in the loss of keystone plant species from humid tropical and high-latitude ecosystems and the extinction of several major insect groups. The subsequent Early to Middle Triassic evinced diminished terrestrial productivity, punctuated by a series of second-order biotic crises that hindered recovery. Here, we survey records of Gondwanan Early to Middle Triassic floral assemblages for evidence of PAIs as an indication of ecosystem recovery following the EPE. We compiled a comprehensive dataset of fossil plant taxa and PAIs for lower Mesozoic strata of Gondwana, revealing an increase in specific and generic floral diversity from the Early to Middle Triassic. We noted a lack of PAIs reported from many localities with abundant fossil leaves, which might be interpreted to be a consequence of a post-EPE delay in the recovery of arthropod feeding guilds compared to the flora. However, by comparing floral assemblages between regions of Gondwana, our results also partly attribute the absence of PAIs to the relative paucity of palaeoichnological and palaeobotanical studies of this interval. To test for potential under-reporting of PAIs in the Triassic, we present a case study of the well-described Australian Middle Triassic Benolong Flora. In contrast to existing Australian Early to Middle Triassic PAI reports on only three plant specimens, this systematic investigation revealed 44 PAI traces comparable to published examples, hosted by 40 fossil plant fragments (7.77% of fragments assessed; N = 591). Margin-feeding traces constituted the dominant Functional Feeding Group (FFG) identified (23 examples: 3.72% of fragments assessed). Our review highlights several Early and Middle Triassic Gondwanan plant fossil-rich successions and existing collections that require further examination. We predict that investigations of these assemblages will greatly elucidate the relationships between rapidly changing environments during the Early and Middle Triassic and their effects on the plant and arthropod communities in the Southern Hemisphere. [ABSTRACT FROM AUTHOR]

Premise: Cladoxylopsids, one of the first lineages with complex organization to rise from the plexus of structurally simple plants that comprised the earliest euphyllophyte floras, are moniliformopsid euphyllophytes. They formed Earth's earliest forests by the Middle Devonian and are thought to have given rise to the equisetopsids and probably some fern lineages. The Lower Devonian (Emsian) Battery Point Formation (Quebec, Canada) contains previously unrecognized cladoxylopsids preserved anatomically. One of these provides new data on structural evolution among euphyllophytes and is described here. Methods: The anatomy and morphology of permineralized axes of the new plant were studied with light and electron microscopy on sections produced using the cellulose acetate peel technique. Morphological comparisons and phylogenetic analysis were used for taxonomic placement of the plant. Results: The plant represents a new species, Paracladoxylon kespekianum Chu et Tomescu, gen. et sp. nov., that has tracheids with modern‐looking bordered pits and the complex cauline vascular architecture characteristic of the genus Cladoxylon. Its dissected ultimate appendages have complex regular taxis and a pattern of vascularization that suggests bilateral symmetry. Conclusions: Paracladoxylon kespekianum is one of the largest Early Devonian euphyllophytes, among the oldest representatives of the cladoxylopsid group, and older than any species of the closely related Cladoxylon by at least 35 million years. It is also one of the oldest anatomically preserved representatives of the cladoxylopsid group. Its anatomical organization pushes the rise of complex vascular architecture among moniliformopsid euphyllophytes deeper in time than previously recognized. [ABSTRACT FROM AUTHOR]

The genus Notiosorex is the only group of shrews in North America with adaptations to arid or semiarid zones. The genus was described with a single variable species, Notiosorex crawfordi , from which 5 new species have since been distinguished. To date, the phylogenetic relationships of Notiosorex species have only been partially analyzed and it is possible that there are still distinct species included within the catch-all of N. crawfordi. Here, we use geometric morphometrics on cranial and mandibular characters of the described extant species, 3 fossil species, and a distinctive population of N. crawfordi as a proxy for an integrated assessment of phylogenetic relationships of all Notiosorex species. Our results indicate that the population from the Altiplano Mexicano is more similar to, yet distinct from, N. villai than it is to N. crawfordi —we describe it as a new species. We also used the resulting tree to reconstruct phylogeographic history within the genus, which suggests that Notiosorex sp. nov. N. villai , N. evotis , and N. tataticuli all diverged allopatrically as populations from the ancestral area that is currently occupied by N. crawfordi (and in the deeper past by the fossil species N. harrisi), and pushed south into the Gulf Coast, the Altiplano, and Baja California within the last 5 million years following the end of the Miocene. [ABSTRACT FROM AUTHOR]

The remains of a unique fossil bony fish were discovered in late Eocene (39.1–36.7 Ma: NZ Kaiatan stage) mudstone at Burnside near Dunedin, New Zealand in the 1930s and subsequently named and described by Frederick Chapman. He interpreted the type specimen as being a large-scaled relative of the modern Thyrsites of the Gempylidae (Scombroidei: Trichiuroidea), known to be swift, large oceanic predators. However, Chapman is unlikely to have seen all of the fossil, and did not discuss the skull and caudal skeleton. Additional material now allows these to be included in the expanded description herein, including key morphologic features of the fish such as the presence of premaxillary fangs. This study describes the rather complex history of the specimen and re-examines this significant fossil fish in the University of Otago collections, giving a more complete understanding of Eothyrsites morphology, paleoecology and relationships. In summary, we suspect Eothyrsites represents an ancestral form of gempylid, closely related to the gemfish group, an important Southern Hemisphere macrofossil record from the Eocene seas around Zealandia. [ABSTRACT FROM AUTHOR]

Albatrosses are among the most intensely studied groups of living birds, yet their fossil record remains sparse. Despite modern albatrosses being more abundant and widespread in the Southern Hemisphere, the vast majority of fossil albatrosses identified to date come from Northern Hemisphere localities. Here, we describe Plotornis archaeonautes sp. nov., a new albatross species from the earliest Miocene that represents the earliest record of Procellariiformes in New Zealand and the earliest uncontroversial record of the clade Pan-Diomedeidae from the Southern Hemisphere. Phylogenetic analyses support the placement of Plotornis outside of the clade uniting all extant albatrosses. The new fossil reveals that stem lineage albatrosses were widespread by the onset of the Neogene. Although the humerus of Plotornis archaeonautes exhibits a short processus supracondylaris dorsalis, this early species may have possessed at least one of the unique ossifications associated with the patagial bracing system present in modern albatrosses. [ABSTRACT FROM AUTHOR]

Premise: The Cenozoic Macquarie Harbour Formation (MHF) hosts one of the oldest and southernmost post‐Cretaceous fossil plant assemblages in Australia. Coinciding with the Early Eocene Climatic Optimum (EECO) and predating the breakup of Australia from Antarctica, it offers critical data to study the diversity and extent of the Austral Polar Forest Biome, and the floristic divergence between Australasia and South America resulting from the Gondwana breakup. Methods: The micromorphology and macromorphology of new fossil plant compressions from the MHF were described and systematically analyzed. Previously published non‐flowering plant records were reviewed and revised. Macrofossil abundance data were provided. The flora was compared with other early Paleogene assemblages from across the Southern Hemisphere. Results: Twelve species of non‐flowering plants were identified from the macrofossil record. Conifers include Araucariaceae (Araucaria macrophylla, A. readiae, A. timkarikensis sp. nov., and Araucarioides linearis), Podocarpaceae (Acmopyle glabra, Dacrycarpus mucronatus, Podocarpus paralungatikensis sp. nov., and Retrophyllum sp.), and Cupressaceae (Libocedrus microformis). Dacrycarpus linifolius was designated a junior synonym of D. mucronatus. Further components include a cycad (Bowenia johnsonii, Zamiaceae), a pteridosperm (Komlopteris cenozoicus, Umkomasiaceae), and a fern (Lygodium dinmorphyllum, Schizaeaceae). Conclusions: The fossil assemblage represents a mixed near‐polar forest with a high diversity of conifers. The morphology and preservation of several species indicate adaptations to life at high latitudes. The coexistence of large‐ and small‐leaved conifers implies complex, possibly open forest structures. Comparisons with contemporaneous assemblages from Argentina support a circumpolar biome during the EECO, reaching from southern Australia across Antarctica to southern South America. [ABSTRACT FROM AUTHOR]

Australopiths are a group of early human ancestors that lived approximately 4 to 2 million years ago and are considered a key transitional form between apes and humans. Studying australopiths can help to understand the evolutionary processes that led to the emergence of humans and gain insights into the unique adaptations and characteristics that set humans apart from other primates. A bibliometric-based review of publications on australopiths contained in the Scopus database was conducted, analyzing approximately 2000 of them. The main authors, institutions, and countries researching this subject were identified, as well as their future development. The connections between authors, countries, and research topics were also analyzed through the detection of communities. The more frequent keywords in this subject are hominid, animal, human, South Africa, and Australopithecus afarensis. Four main research clusters were identified in the field of australopiths: palaeobiology, cranial evolution, locomotion, and mandible evolution and morphometry. The most important countries in terms of collaboration networks are South Africa, the UK, France, and Germany. Research on australopiths is ongoing, and new research clusters are expected to emerge, such as those focused on pre-australopiths and the molecular evolution and taxonomy of australopiths. Overall, this work provides a comprehensive overview of the state of research on australopiths and offers insights into the current direction of the field. [ABSTRACT FROM AUTHOR]

Wing venation pattern is particularly important in the taxonomy and classification of insects, especially for fossil material. There are recognized transformation series that apparently often represent a mainstream in the evolutionary trend of the wing venation of Hymenoptera. One notable instance is the gradual reduction of the subcosta veins from multiple branches to their total absence. Herein, we place four new genera and six new species in the family Blasticotomidae (=Xyelotomidae) of Hymenoptera. They are Xyelocerus abruptus sp. nov. Liberitoma tenella gen et sp. nov. Liberitoma compta sp. nov. Liberitoma incompleta sp. nov. Aduantoma insolita gen. et sp. nov. and Apertoma gen. nov. from the Middle Jurassic of China, and Enspeletoma oligocaenica gen. et sp. nov. from the Upper Oligocene of Germany. The newly discovered species of Liberitoma from China possessed a supernumerary hind branch of vein Sc in its forewing, which has apparently never been recorded before in other hymenopterans, and the unique venation pattern of Aduantoma suggests the possibility of two additional steps of vein Sc transformation before its complete loss. Based on the new findings, Blasticotomidae are proposed to be divided into four subfamilies: Blasticotominae Thomson 1871, Dahurotominae subfam. nov. Pseudoxyelocerinae subfam. nov. and Undatominae subfam. nov. [ABSTRACT FROM AUTHOR]

The Ediacaran of eastern Newfoundland preserves the world's oldest known eumetazoan body fossils, as well as the earliest known record of fossilized muscular tissue. Re-examination of the holotype of the eight-armed Haootia quadriformis in terms of its morphology, the arrangement of its muscle filament bundles, and hitherto undescribed aspects of its anatomy support its interpretation as a crown staurozoan. We also document several new fossils preserving muscle tissue with a different muscular architecture to Haootia, but with only four arms. This new material allows us to describe a new crown group staurozoan, Mamsetia manunis gen. et sp. nov. This work confirms the presence of crown group medusozoan cnidarians of the Staurozoa in the Ediacaran of Newfoundland circa 565 Ma. [ABSTRACT FROM AUTHOR]

Body size is of fundamental importance to our understanding of extinct organisms. Physiology, ecology and life history are all strongly influenced by body size and shape, which ultimately determine how a species interacts with its environment. Reconstruction of body size and form in extinct animals provides insight into the dynamics underlying community composition and faunal turnover in past ecosystems and broad macroevolutionary trends. Many extinct animals are known only from incomplete remains, necessitating the use of anatomical proxies to reconstruct body size and form. Numerous limitations affecting the appropriateness of these proxies are often overlooked, leading to controversy and downstream inaccuracies in studies for which reconstructions represent key input data. In this perspective, we discuss four prominent case studies (Dunkleosteus, Helicoprion, Megalodon and Perucetus) in which proxy taxa have been used to estimate body size and shape from fragmentary remains. We synthesise the results of these and other studies to discuss nuances affecting the validity of taxon selection when reconstructing extinct organisms, as well as mitigation measures that can ensure the selection of the most appropriate proxy. We argue that these precautionary measures are necessary to maximise the robustness of reconstructions in extinct taxa for better evolutionary and ecological inferences. [ABSTRACT FROM AUTHOR]

A reliable character system is crucial to taxonomy and systematics, and it promises valid downstream inferences, e.g., estimates of diversity and disparity, reconstruction of evolutionary history, and even stratigraphic correlations. Modern taxonomy and systematics of extant cockroaches requires an integrative study involving multiple lines of evidence with emphasis on genital and reproductive characteristics and molecular data. In contrast, many fossil cockroach taxa published recently are based solely on forewings. Many studies have shown that forewing-based taxa are questionable. In order to find out how much of the phylogenetic signal we could ascertain from venational similarity, and how confident we could be, this study used forewing characters to reconstruct phylogenies of the genera of well-recognized family-group taxa. The intuitively reconstructed phylogeny of 75 extant genera failed to recover those taxa or their relationships. Parsimony analyses of various datasets all yielded strong polyphyly and chaotic relationships. In conclusion, the forewing of cockroaches is not a universally competent character system. The underlying causes are the complicated nature of veins and the limitations of current analytical techniques. The uncertainty in forewing-based taxonomy and systematics has been underestimated in the literature. Forewing-based fossil taxa warrant re-evaluation; some of them are herein deemed nomina dubia in their current state. [ABSTRACT FROM AUTHOR]

Coccinelloid beetles have a sparse fossil record in the Mesozoic. Here, we describe and illustrate an unusual coccinelloid beetle, Yassibum yoshitomii gen. et sp. nov., from mid-Cretaceous Kachin amber. Yassibum stands out within the Coccinelloidea due to its notched profemora and the presence of antennal grooves on the elytral epipleura. Based on our phylogenetic analyses, we suggest that Yassibum is most likely related to the family Anamorphidae. The alternative placements are critically evaluated based on our comparison of the morphology. [ABSTRACT FROM AUTHOR]

Mass extinctions are major influences on both the phylogenetic structure of the modern biota and our ability to reconstruct broad-based patterns of evolutionary history. The most recent mass extinction is also the most famous—that which implicates a bolide impact in defining the Cretaceous/Palaeogene boundary (K/Pg). Although the biotic effects of this event receive intensive scrutiny, certain ecologically important and diverse groups remain woefully understudied. One such group is the freshwater ray-finned fishes (Actinopterygii). These fish represent 25% of modern vertebrate diversity, yet the isolated and fragmentary nature of their K/Pg fossil record limits our understanding of their diversity dynamics across this event. Here, we address this problem using diversification analysis of molecular-based phylogenies alongside a morphotype analysis of fossils recovered from a unique site in the Denver Basin of western North America that provides unprecedented K/Pg resolution. Our results reveal previously unrecognized signals of post-K/Pg diversification in freshwater clades and suggest that the change was driven by localized and sporadic patterns of extinction. Supported inferences regarding the effects of the K/Pg event on freshwater fish also inform our expectations of how freshwater faunas might recover from the current biodiversity crisis. [ABSTRACT FROM AUTHOR]

A stakeholder engagement workshop was held in May 2024 as part of the "Community-driven enhancement of information ecosystems for the discovery and use of paleontological specimen data" project, which is funded under the United States National Science Foundation (NSF) Geosciences Open Science Ecosystem (GEO OSE) program. This report describes the activites and outcomes of the workshop. [ABSTRACT FROM AUTHOR]

The research history of fossil Phasmatodea is reviewed in this paper, focusing on the current status of taxonomic and phylogenetic studies. The diversity, localities and ages of all reported fossil taxa of Phasmatodea all over the world are overviewed. The mimicry and defense behavior revealed by Phasmatodea fossils are summarized, involving their early evolution of wings and mimicry related body characters. The existing problems in the studies of fossil Phasmatodea are discussed, along with an outlook on future research of Phasmatodea. [ABSTRACT FROM AUTHOR]

Two new genera and species of Ithonidae, i.e. Stictopolystoechotes sparsulus Gao, Xu et Wang gen. et sp. nov. and Pycnopolystoechotes striatus Gao, Xu et Wang gen. et sp. nov., are described from the Middle Jurassic Jiulongshan Formation, Inner Mongolia, China. The two new genera can be assigned to the Polystoechotes genus-group of Ithonidae by the combination of following characters: elongated forewing, Sc and RA fused distally, crossveins in radical sector sparse except for the gradate series, and simple CuP branches. The known wing markings among the Jurassic Polystoechotes genus-group were outlined, which suggested the diversification of wing markings was possibly positively associated with the species radiation of this lineage during this period. Moreover, a key to fossil genera of the Polystoechotes genus-group was provided. LSID: urn:lsid:zoobank.org:pub:CAEEFF67-876D-4709-BE4A-18F5AB04B01D Stictopolystoechotes: urn:lsid:zoobank.org:act:666ED6B2-B132-4CF4-BFB0-ED6B86EB30F4 Stictopolystoechotes sparsulus: urn:lsid:zoobank.org:act:6C96432B-205F-4F39-A647-3E0048169A92 Pycnopolystoechotes: urn:lsid:zoobank.org:act:1D1343DF-844E-4243-BAB9-242F6232EEBF Pycnopolystoechotes striatus: urn:lsid:zoobank.org:act:B4EFBE6B-5011-4103-A65F-77D65BEC0B85 [ABSTRACT FROM AUTHOR]

We describe the first fossil species of the extant genus Eulomalus (Coleoptera: Histeridae: Dendrophilinae: Paromalini) from the Eocene Baltic amber. Eulomalus balticussp. nov. has a flattened body shape typical for the Histeridae living under the bark of decaying trees. The newly described species is the only European representative of the genus, which is presently distributed chiefly in the Indomalayan realm. [ABSTRACT FROM AUTHOR]

Periderm is a well‐known structural feature with vital roles in protection of inner plant tissues and wound healing. Despite its importance to plant survival, knowledge of periderm occurrences outside the seed plants is limited and the evolutionary origins of periderm remain poorly explored. Here, we review the current knowledge of the taxonomic distribution of periderm in its two main forms – canonical periderm (periderm formed as a typical ontogenetic stage) and wound periderm (periderm produced as a self‐repair mechanism) – with a focus on major plant lineages, living and extinct. We supplement the published occurrences with data based on our own observations and experiments. This updated body of data reveals that the distribution of wound periderm is more widespread taxonomically than previously recognized and some living and extinct groups are capable of producing wound periderm, despite canonical periderm being absent from their normal developmental program. A critical review of canonical and wound periderms in extant and fossil lineages indicates that not all periderms are created equal. Their organisation is widely variable and the differences can be characterised in terms of variations in three structural features: (i) the consistency in orientation of periclinal walls within individual files of periderm cells; (ii) the lateral coordination of periclinal walls between adjacent cell files; and (iii) whether a cambial layer and conspicuous layering of inward and outward derivatives can be distinguished. Using a new system of scoring periderm structure based on these criteria, we characterise the level of organisation of canonical and wound periderms in different lineages. Looking at periderms through the lens provided by their level of organisation reveals that the traditional image of periderm as a single generalised feature, is best viewed as a continuum of structural configurations that are all predicated by the same basic process (periclinal divisions), but can fall anywhere between very loosely organized (diffuse periclinal growth) to very tightly coordinated (organized periclinal growth). Overall, wound periderms in both seed plants and seed‐free plants have lower degrees of organisation than canonical periderms, which may be due to their initiation in response to inherently disruptive traumatic events. Wound and canonical periderms of seed plants have higher degrees of organisation than those of seed‐free plants, possibly due to co‐option of the programs responsible for organizing their vascular cambial growth. Given the importance of wound periderm to plant survival, its widespread taxonomic distribution, and its early occurrence in the fossil record, we hypothesise that wound periderm may have had a single origin in euphyllophytes and canonical periderm may have originated separately in different lineages by co‐option of the basic regulatory toolkit of wound periderm formation. In one evolutionary scenario, wound periderm regulators activated initially by tissue tearing due to tensional stresses elicited by woody growth underwent heterochronic change that switched their activation trigger from tissue tearing to the tensional stresses that precede it, with corresponding changes in the signalling that triggered the regulatory cascade of periderm development from tearing‐induced signals to signalling induced by tension in cells. [ABSTRACT FROM AUTHOR]

The evolution of the extinct megatooth shark, Otodus megalodon, and its close phylogenetic relatives remains enigmatic. A central question persists regarding the thermophysiological origins of these large predatory sharks through geologic time, including whether O. megalodon was ectothermic or endothermic (including regional endothermy), and whether its thermophysiology could help to explain the iconic sharks gigantism and eventual demise during the Pliocene. To address these uncertainties, we present unique geochemical evidence for thermoregulation in O. megalodon from both clumped isotope paleothermometry and phosphate oxygen isotopes. Our results show that O. megalodon had an overall warmer body temperature compared with its ambient environment and other coexisting shark species, providing quantitative and experimental support for recent biophysical modeling studies that suggest endothermy was one of the key drivers for gigantism in O. megalodon and other lamniform sharks. The gigantic body size with high metabolic costs of having high body temperatures may have contributed to the vulnerability of Otodus species to extinction when compared to other sympatric sharks that survived the Pliocene epoch.

Many modern extinction drivers are shared with past mass extinction events, such as rapid climate warming, habitat loss, pollution and invasive species. This commonality presents a key question: can the extinction risk of species during past mass extinction events inform our predictions for a modern biodiversity crisis? To investigate if it is possible to establish which species were more likely to go extinct during mass extinctions, we applied a functional trait-based model of extinction risk using a machine learning algorithm to datasets of marine fossils for the end-Permian, end-Triassic and end-Cretaceous mass extinctions. Extinction selectivity was inferred across each individual mass extinction event, before testing whether the selectivity patterns obtained could be used to 'predict' the extinction selectivity exhibited during the other mass extinctions. Our analyses show that, despite some similarities in extinction selectivity patterns between ancient crises, the selectivity of mass extinction events is inconsistent, which leads to a poor predictive performance. This lack of predictability is attributed to evolution in marine ecosystems, particularly during the Mesozoic Marine Revolution, associated with shifts in community structure alongside coincident Earth system changes. Our results suggest that past extinctions are unlikely to be informative for predicting extinction risk during a projected mass extinction.

Recent paleontological and geological research in Pakistan has found numerous fossils belonging to different groups of prehistoric organisms, including reptiles. Pakistan is rich in paleontological evidence of vertebrates, especially tracks/footprints of Mesozoic reptiles. These reptilian fossils are significant for paleo biogeographical studies because the region is connected to Asia in the north and east, and Eurasia and Africa in the north and west. It was attached to Gondwana in the past (Jurassic and Pre-Jurassic), so Mesozoic vertebrates show distinct affinities with Gondwana, Cenozoic vertebrates show affinities with Eurasia and migrated from the Indo-Pak subcontinent via the ancient Indus system of the West Indus River (and vice versa). Neogene (Siwalik Age) deposits in India and Pakistan have produced prehistoric reptiles containing numerous fossils, many of which were named in the 19th century. Recent geological and paleontological research conducted in Pakistan at the beginning of the new millennium/third millennium (2000-2019) found more than 3,000 fossils divided into 45 biological groups (taxa). [ABSTRACT FROM AUTHOR]

Shaligrams are the sacred fossil ammonites of the Himalayas. Viewed primarily as manifestations of Hindu gods, Shaligrams are obtained by pilgrimage to Himalayan Nepal and are then brought home to families and communities all over South Asia and the Diaspora as household deities. But Shaligrams also contain a variety of natural characteristics that are read and interpreted through long-standing oral traditions that use these features to both determine which specific deity is manifest within the stone and to link each Shaligram with a body of religious stories and local folklore. Therefore, the semiotic interpretation of Shaligrams instantiates ritual practices by which each stone becomes both an object and a text; able to be read by those fluent in its symbolic language. This practice then blurs the line between categories of object and archive; where fossils become literal texts and stones become storytellers. [ABSTRACT FROM AUTHOR]

Beetles have a remote evolutionary history dating back to the Carboniferous, with Mesozoic fossils playing a pivotal role in elucidating the early evolution of extant families. Despite their exceptional preservation in amber, deciphering the systematic positions of Mesozoic trogossitid‐like beetles remains challenging. Here, we describe and illustrate a new trogossitid‐like lineage from mid‐Cretaceous Kachin amber, Foveapeltis rutai Li, Kolibáč, Liu & Cai, gen. et sp. nov. Foveapeltis stands out within the Cleroidea due to the presence of a significant large cavity on each hypomeron. While the exact phylogenetic placement of Foveapeltis remains uncertain, we offer a discussion on its potential affinity based on our constrained phylogenetic analyses. [ABSTRACT FROM AUTHOR]

Between 1862 and 1867, Albert Gaudry (1824–1908) published Animaux fossiles et géologie de l'Attique, a seminal work describing and depicting the rich collection of mammalian fossils that he assembled during his scientific expeditions in Attica, Greece. This paper will focus on how Gaudry's collection and reconstruction of the extinct fauna of Attica articulated itself with geopolitical, cultural, and economic concerns regarding the newly independent Greek state and its place within Europe. Since the Greek War of independence against the Ottomans between the 1820s and 1830s, European nations sought to secure a series of strategic advantages in the region. France for example invested a lot of capital, military, and cultural resources in the newly independent Greek state in hope of cementing France's influence in the Mediterranean. Between 1853 and 1854, Gaudry was sent on a mission by the French government to evaluate the agricultural production of countries of the 'Orient', including Greece. Gaudry was tasked with gathering information intended to facilitate the advancement of agriculture in France's colonies as well as the planning of future investments in the Eastern Mediterranean. In his report published in 1855, Gaudry explicitly defined the restoration of Greece's agricultural industry as a sine qua non condition for Greece's establishment as a European state. It was during this official mission that Gaudry learned about the existence of exceptionally rich fossil sites in Attica. He returned on multiple occasions to collect thousands of fossil specimens from these sites. In Animaux fossiles et géologie de l'Attique and in his subsequent publications on the same fossils, Gaudry regularly articulated the reconstruction of extinct mammals and their environment with inspired descriptions of the ruins of Athens. In his writings, the reconstruction of a Greek geological past inhabited by a grandiose and harmonious megafauna echoed the celebration of Ancient Greece as the cradle of Western civilization. This paper will comment on the complex interplay between three temporalities which informed Gaudry's work on the mammalian fossils of Attica: (1) the reconstructed paleontological past, (2) the idealized Ancient Greece, and (3) the present geopolitical context. [ABSTRACT FROM AUTHOR]

Among the most widely promoted examples of fossil folklore is a supposed link between the Central Asian horned dinosaur Protoceratops and the griffin, a gold-guarding mythical creature combining features of lions and birds. First proposed in the 1990s, this geomyth postulates that tales of Protoceratops fossils were transmitted westward along trade routes from Asian gold mines to inform griffin lore among the ancient Greeks. An evaluation of the Protoceratops–griffin link, however, finds it uncompelling. Not only does it ignore established histories of griffin art and myth, but no convincing connections occur between Protoceratops and central aspects of griffin lore, such as gold guarding. In fact, Protoceratops fossils occur hundreds of kilometres from the nearest gold deposits, subverting suggestions that they inspired the griffin's association with gold. Interpretations of ancient literary references to griffins as pertaining to Protoceratops are unconvincing, and suggested anatomical similarities between griffins and Protoceratops are selectively identified. We regard the Protoceratops–griffin link as an 'ex post facto geomyth': an effort to find significance in superficial, inconsequential readings of geological phenomena and mythology. We posit that the allure of ancient cultures sharing our modern fascination with dinosaurs has denied this idea due scepticism. [ABSTRACT FROM AUTHOR]

We describe a diverse aquatic insect assemblage from McGraths Flat, a Miocene Lagerstätte in central New South Wales, Australia that includes representatives of Sialidae, Limoniidae, Chironomidae and Chaoboridae. The aquatic insect fossils from this deposit consist predominantly of larvae. These include a new species of phantom midge (Chaoborus, Chaoboridae), three morphotypes of non‐biting midges (Chironomidae), one morphotype of cranefly (Limoniidae) and one morphotype of alderfly (Sialidae). The large number of fossil specimens enabled us to study the ontogeny of the new midge species. We discerned growth rates in fossil larvae, using morphometry of all four instars of Chaoborus. The simultaneous presence of taxa associated with still water and taxa associated with flowing water supports the hypothesis that McGraths Flat was deposited in an isolated water body (oxbow lake/billabong) with influence from a river during high water events. [ABSTRACT FROM AUTHOR]