Revision of the Family Helotidae (Coleoptera: Cucujoidea) III: The Thibetana Group and a Checklist of Helota MacLeay Species

The thibetana species group of Helota MacLeay is revised. Five species are considered to be valid. The following synonyms are proposed: Helota sinensisOlliff, 1883 (=H. tonkinensisRitsema, 1905a); and H. thibetana Westwood, 1842 (=H. verrucosaRitsema, 1894; H. donckieriRitsema, 1907; H. westwoodiiRitsema, 1907). Lectotypes are designated for Helota sinensis Olliff, H. tonkinensis Ritsema, H. donckieri Ritsema, and H. westwoodii Ritsema. A key for the thibetana species group is provided. Figures of the protibia, penis, parameres, internal sac, and eighth abdominal tergite are given for each species. A checklist of the 23 valid species and subspecies of the genus Helota MacLeay is provided.