New notocupedin beetle in Cretaceous Burmese amber (Coleoptera: Archostemata: Ommatidae)

The archostematan family Ommatidae Sharp & Muir, 1912 represent one of the oldest beetle lineages, originating probably in the Permian or early Triassic (Tan et al., 2012; Cai & Huang, 2017; Zhang et al., 2018). Fossil ommatids have been known since the Triassic and, along with other members of the suborder Archostemata, are reminiscent of the earliest and most basal Permian beetles, with which they share a dorsoventrally compressed body, distinctly raised elytral veins, and characteristic rows of window punctures on the elytra (Crowson, 1962). As such, archaic ommatid beetles have sometimes been referred to as ‘living fossils’ (Jarzembowski et al., 2018). Only six extant ommatid species with a highly disjunct distribution are known (Hörnschemeyer & Beutel, 2016), but the family was much more diverse in the Mesozoic.