Diverse downstream migration patterns of the anadromous Japanese grenadier anchovy Coilia nasus in the Chikugo River estuary and Ariake Sea, Japan

Variations in the downstream migration patterns of the anadromous Japanese grenadier anchovy Coilia nasus in the Chikugo River estuary and the inner part of the Ariake Sea, Japan were investigated using otolith Sr:Ca ratios over life history transects. Although most individuals migrated from freshwater or low salinity areas (i.e., upper estuary; salinities