Tetanoptilon Bode 1953

Tetanoptilon Bode, 1953 Type species. Tetanoptilon brunsvicense Bode, 1953: 266 (by original designation) (Germany) (Early Jurassic). Comments. The genus is represented only by a single hind wing which was redescribed by Ponomarenko (1995). This specimen does not allow us to determine the systematic position of the genus since it is represented only by a hind wing with a set of unusual characteristics (e.g., 1r-m is crossvein-like; subcostal veinlets arising nearly perpendicular to Sc; the proximal branches of RP are widely spaced; intracubital area is broad basally). Based on the elongate and densely pectinate branching pattern of the hind wing CuP, we can preclude placement in Gumillinae, Spilosmylinae and Protosmylinae. The density of crossveins and pectinate branching of the posterior wing veins likely also reflected in the unknown forewing of this genus, which would also suggest that it cannot be placed in Mesosmylininae or Archeosmylidae. While we agree with its placement in Osmylidae, we cannot definitively place it in any of the remaining subfamilies (e.g., Stenosmylinae, Osmylinae, Porisminae, Eidoporisminae).
Published as part of Winterton, Shaun L., Martins, Caleb Califre, Makarkin, Vladimir, Ardila-Camacho, Adrian & Wang, Yongjie, 2019, Lance lacewings of the world (Neuroptera: Archeosmylidae, Osmylidae, Saucrosmylidae): review of living and fossil genera, pp. 1-99 in Zootaxa 4581 (1) on page 85, DOI: 10.11646/zootaxa.4581.1.1, http://zenodo.org/record/2633842
{"references":["Bode, A. (1953) Die Insektenfauna des Ostniedersachsischen Oberen Lias. Palaeontographica: Beitrage zur Naturgeschichte der Vorwelt (A), 103, 1 - 375.","Ponomarenko, A. G. (1995) [1996] Upper Liassic neuropterans (Insecta) from Lower Saxony, Germany. Russian Entomological Journal, 4, 73 - 89."]}