Armascirus Den Heyer 1978

Armascirus Den Heyer, 1978 Historical review. The first Armascirus was described by Kramer (1881) as Scirus taurus. Berlese (1888) described S. taurus var. bison. Banks (1894) described S. quadripilus. Thor (1902) transferred S. taurus to Cunaxa. Banks (1914) described C. armata. Womersley (1933) reported C. taurus from Australia. Thor and Willmann (1941) transferred S. taurus var. bison to Cunaxa and raised it to full species status, viz. C. bison and transferred S. quadripilus to Cunaxa; they also redescribed and figured C. armata, C. bison, C. quadripilus, and C. taurus. Baker and Hoffmann (1948) synonymized S. quadripilus and C. armata with C. taurus; they followed Thor and Willmann (1941) in placing C. taurus var. bison in Cunaxa but declined to recognize it as a species and instead kept it as a variety or subspecies of C. taurus. Zaher et al. (1975) collected C. taurus in Egypt. Chaudhri (1977) described Dactyloscirus ebrius and D. fuscus from Pakistan. Den Heyer (1978) split Armascirus from Dactyloscirus and Cunaxa and raised the subfamily Cunaxinae to accommodate them, thus refining the definitions of all three genera; he transferred C. taurus and C. bison to the new; and described A. huyssteeni, A. lebowensis, A. limpopoensis, and A. albiziae. Kuznetzov and Livshitz (1979) redescribed and figured C. taurus and C. bison from Russia, either disagreeing with or being unaware of Den Heyer���s 1978 publication. Tseng (1980) reported A. taurus from Taiwan. Chaudhri (1980) described D. fixus from Pakistan. Den Heyer (1980) erected the tribe Armascirini and made Dactyloscirus and Armascirus the sole representatives. Gupta and Ghosh (1980) erected Indocunaxa, a monotypic genus with I. smileyi as the type species. Liang (1983) reported A. taurus from China. Michocka (1987) described D. rafalskii from Poland. A. mactator and A. pluri were described by Muhammad and Chaudhri (1991). Smiley (1992) described A. gimplei, A. anastosi, A. harrisoni, A. heryfordi, A. virginiensis, D. bakeri, and D. campbelli and transferred A. bison to Dactyloscirus. He also synonymized I. smileyi with A. taurus, though did not include Indocunaxa as a synonymy of Armascirus; this is confusing as Indocunaxa is monobasic and synonymizing the only species with A. taurus functionally synonymized the genus with Armascirus. Corpuz-Raros (1995) described A. garciai and A. makilingensis from the Philippines. Hu (1997) reported A. bison and A. taurus from China. Armascirus satianaensis and A. asghari were described by Bashir and Afzal (2005). Corpuz-Raros and Gru��zo described A. javanus. Corpuz-Raros (2008) described A. bifidus. Bashir, Afzal, and Khan described four species from Pakistan, A. akhtari, A. jasmina, A. sabrii, and A. gojraensis. Den Heyer and Castro (2008 b) reaffirmed the synonymization of Indocunaxa with Armascirus. Kal��z (2009) described A. cyaneus and A. cerris from Central Europe and transferred D. bison, D. campbelli, D. ebrius, D. fixus, D. fuscus, and D. rafalskii to Armascirus. Generic diagnosis. Den Heyer (1978) gives a detailed diagnosis of this genus. The following description highlights those features that are prominent or are regarded as being diagnostic. Palpi five segmented and end in strong claw. They extend beyond the subcapitulum by at least the last segment and are often adorned with an apophysis between the genua and tibiotarsi that tapers to a point; this apophysis shorter in males than in females. Basifemora complemented with a simple seta; telofemora complemented with a spine-like seta. These two segments fused though a dark line remains visible to differentiate them. Subcapitulum complemented with six pairs of setae (hg 1���4 and two pairs of adoral setae). It can be covered by integumental papillae which are either randomly distributed or form a polygonal, reticulated pattern. Female dorsal idiosoma with at least one sclerotized plate that bears two pairs of setose trichobothria (at and pt) and two pairs of simple setae (lps and mps). 0���4 other major plates and platelets may also be present. All plates, if present, covered by integumental papillae that form a reticulated pattern. Integument between plates striated. Seven pairs of setae, c 1���2, d 1, e 1, f 1 and h 1, present. Each seta, when not on a major plate or platelet, surrounded by a minute platelet only slightly larger than the setal socket. Cupule im present, usually laterad or in the proximity of e 1. Dorsal idiosoma of males similar except a single large plate complemented with c 1���2, d 1, e 1 and f 1 present. Female ventral idiosoma complemented by the coxal, genital, and anal plates. Coxal plates reticulated in the same manner as the dorsal plates. Coxae I and II often fused; coxae III and IV often fused. Coxae I���IV setal formula usually: males 3 - 1-3 - 3 or females 3 - 2-3 - 3. Genital plates each bear four setae; two pairs of genital papillae are visible underneath the plates. Anal plates bear one pair of setae (ps 1). Two pairs of setae (ps 2 and h 2) associated with but do not occur on the anal plates. Cupule ih present in close proximity to h 2. Integument between plates striated and bears 5���7 pairs of additional setae. Ventral idiosoma of males similar except the coxal plates are much more extensive. Sclerotized aedeagus is often visible in association with the genital plates. Legs comparatively long, at least �� the length and often longer than the body. Famulus on tarsi I normally shaped, not large and tri-pronged as in Dactyloscirus. Tarsi are constricted apically, resulting in large tarsal lobes.
Published as part of Skvarla, Michael J. & Dowling, Ashley P. G., 2012, Some new armascirine cunaxids (Acari: Prostigmata: Cunaxidae) from the Eastern United States, pp. 1-34 in Zootaxa 3194 on pages 5-6, DOI: 10.11646/zootaxa.3194.1.1, http://zenodo.org/record/201630
{"references":["Den Heyer, J. (1978) Four new species of Armascirus gen. nov. (Prostigmata: Acari) from the Ethiopian region. Journal of the Entomological Society of South Africa, 41 (2), 217 - 239.","Kramer, P. (1881) Scirus. Arch. Natura. 81, 17, t. 3, fig. 9.","Berlese, A. (1888) Acari Austro-Americani quos collegit Aloysius Balzan. Bullettino della Societa Entomologica Italiana, 20, 171 - 222","Banks, N. (1894) New American Acarina. Transactions of the American Entomological Society, 21, 209 - 222.","Thor, S. (1902) Zur Systematik der Acarinenfamilien Bdellidae Koch, 1842, Grube 1859, Eupodidae Koch, 1842 und Cunaxidae Sig Thor, 1902. Verhandllungen der kaiserlich-kongiglichen zoologish-botanischen Gesellschaft in Wien, 159 - 165.","Banks, N. (1914) New Acarina. Journal of Entomology and Zoology, 6, 55 - 66","Womersley, H. (1933) On some Acarina from Australia and South Africa. Transactions of the Royal Society of South Australia, 57, 108 - 112.","Thor, S. & Willmann, C. (1941) Eupodidae, Penthalodidae, Penthaleidae, Rhagidiidae, Pachygnathidae, Cunaxidae. In: Thor, S. & Willmann, C. (Eds). Das Tierreich, Eine Zusammenstellung und Kennzeichung der rezenten Tierformen, Lieferung 71 a. Walter De Gruyter & Co, Leipzig, pp. 164 - 175.","Baker, E. W. & Hoffmann, A. (1948) Acaros de la familia Cunaxidae. Anales de la Escuela Nacional de Ciencias Biologicas Mexico, 5 (3 - 4), 229 - 273.","Zaher, M. A., Soliman, Z. R. & El-Bishlawy, S. M. (1975) Studies on population dynamics of soil predaceous prostigmatid mites in Giza, Egypt. Zeitschrift fur Angewandte Entomologie, 79, 440 - 443.","Chaudhri, W. M. (1977) Descriptions of the mites of the Family Cunaxidae (Acarina) from Pakistan. Pakistan Journal of Agricultural Science, 14 (2 - 3), 41 - 52.","Kuznetzov, N. N. & Livshitz, I. Z. (1979) Predatory mites of the Nikita Botanical Gardens (Acariformes: Bdellidae, Cunaxidae Camarobiidae). Trudy Gosudarstvennogo Nikitskogo Botanicheskogo Sada, 79, 51 - 105.","Tseng, Y. H. (1980) Taxonomical study of the mite family Cunaxidae from Taiwan (Acarina: Tromoidiformes). Quarterly Journal of the Taiwan Museum, 33 (3 - 4), 253 - 277.","Chaudhri, W. M. (1980) Studies on the biosystematics and control of mites of field crops, vegetables and fruit plants in Pakistan, second annual report. University of Agriculture Faisalabad, Pakistan, 80 pp.","Den Heyer, J. (1980) A new classification system for the family Cunaxidae (Actinedida: Acarida). Publications of the University of the North, Series A 23, 1 - 12.","Gupta, S. K. & Ghosh, S. K. (1980) Some prostigmatid mites (Acarina) from Andaman and Nicobar Islands. Records of the Zoological Survey of India, 77, 189 - 213.","Liang, G. W. (1983) Notes on four species of mite (Acarina: Cunaxidae) in China. Natural Enemies of Insects, 5 (3), 104 - 107.","Michocka, S. (1987) Mites (Acari) of the Bdellidae and Cunaxidae families in Poland. Monografie Fauny Polski, 14, 1 - 127.","Muhammad, T. & Chaudhri, W. M (1991) Two new cunaxid mites of the genus Armascirus (Acarina: Cunaxidae) from Pakistan. Pakistan Entomologist, 13 (1 - 2), 50 - 55.","Smiley, R. L. (1992) The predatory mite family Cunaxidae (Acari) of the world with a new classification. Indira Publishing House, West Bloomington, Michigan, 356 pp.","Corpuz-Raros, L. A. (1995) Philippine predatory mites of the family Cunaxidae (Acari). 2. Genera Armascirus Den Heyer and Dactyloscirus Berlese. The Philippine Agriculturist, 78 (2), 159 - 173.","Hu, S. J. (1997) Cunaxid mites recorded in China. Journal of Ninbo Teachers College, 15 (1), 56 - 59.","Bashir, M. H. & Afzal, M. (2005) New cunaxid mites of the genus Armascirus from Punjab-Pakistan. Pakistan Journal of Agricultural Sciences, 42 (3 - 4), 117 - 121.","Corpuz-Raros, L. A. (2008) Additional species of Cunaxidae and first report of the subfamily Neobonzinae (Cunaxidae, Acari) from the Philippines. Asia Life Sciences, 17 (1), 71 - 89.","Den Heyer, J. & Castro, T. M. M. G. (2008 b) A new Neotropical genus of the family Cunaxidae (Acari: Prostigmata: Bdelloidea). Zootaxa, 1843, 35 - 46.","Kaluz, S. (2009) Two new Palearctic mite species of the family Cunaxidae (Acari: Prostigmata). Zootaxa. 2198, 27 - 40."]}