Pseudoscalibregma Ashworth 1901

Genus Pseudoscalibregma Ashworth, 1901 Type-species: Scalibregma parvum Hansen, 1879. Diagnosis. Body elongate, arenicoliform. Prostomium T-shaped with lateral horns. Peristomium achaetous, surrounding prostomium dorsally and forming upper and lower lips of mouth ventrally. Parapodia of posterior segments with dorsal and ventral cirri; interramal papillae present; postsetal lamellae absent. Branchiae absent. Setae include capillaries, lyrate setae, and sometimes few inconspicuous spinous setae among capillaries of setigers 1���2, blunt, pointed, or bifurcated, representing homologues of lyrate setae; large conspicuous spines absent. Pygidium with long anal cirri. Remarks. Apart from a more variable morphology of dorsal and ventral cirri, the only character that separates Pseudoscalibregma from Scalibregma is the absence of branchiae. The presence of short spinous setae in the first 1���2 setigers as occurs in species of Scalibregma was confirmed by Imajima (2009) as part of his description of P. orientalis Imajima, 2009 from Japan and Bakken et al. (2014) as part of their redescription of the type species, P. parvum. I can also confirm after a re-examination of the holotype that P. usarpium Blake, 1981 from Antarctica also has these setae. Four species of Pseudoscalibregma have been reported from Antarctic seas: (1) P. bransfieldium (Hartman, 1967), (2) P. usarpium, (3) P. papilla Sch��ller, 2008, and (4) P. h ar t m a na e (Blake, 1981), new combination. A fifth species was found in the LIS-A collections and is described here. Additional descriptive notes are provided for P. bransfieldium. Character���N��. Setigers 10 14 �� 15 16 17 �� 18 20 22 �� 24 28 30 Length 630 ��m 850 �� 900 ��m 945 �� 990 ��m 960 �� 1,221 ��m 1300 �� 1875 ��m 2.6 �� 2.8 mm 3.5 mm 5.67 mm Width 70 ��m 220 �� 232 ��m 305 �� 325 ��m 353 �� 407 ��m 450 �� 580 ��m 0.62 mm 0.75 mm 0.95 mm Pr��st��mium Bulb��us,r��unded Bulb��us, R��unded, Inflated, lateral Bec��ming wide, Br��ad, thickened Lateral h��rns well Lateral h��rns fully extended apically p��inted apically bulges, tapering thickened acr��ss anteri��r margin, devel��ped devel��ped apically t�� r��unded subterminally, then lateral h��rns tip tapering t�� r��unded devel��ping anteri��r peak Perist��mium Singlering Single ring Single ring Single narr��w ring single ring d��rsally; Single ring d��rsally, Single ring d��rsally; Single ring d��rsally; d��rsally and d��rsally and d��rsally and d��rsally bec��ming ventrally enlarging ventral ring expanding ventral ring f��rming three rings ventrally ventrally ventrally ventrally expanded ventrally, anteri��r t�� m��uth t�� enc��mpass upper lip upper and l��wer lips ��f enc��mpassing the with pads devel��ping ��f m��uth m��uth m��uth Branchiae Absent Absent Absent Absent Absent 22 Setigers∶ ��ne Branchiae present Branchiae present branchial anlage, setigers 2 �� 5; with 1, 2, setigers 2 �� 5 with setigers 3 �� 4; 3, and 4 l��be(s) individual l��bes 24 Setigers∶ ��ne respectively branching branchial l��be ��n setigers 3 �� 5 D��rsal cirri Absent Absent Present, fr��m Present Present Present Present Present ab��ut setiger 8 �� 9 ventral cirri Absent Absent Absent Absent 1 �� 2 present, p��steri��r Present Present Present m��st setigers D��rsal pigmented Absent Absent Absent Absent Present Present Present Present glands setiger 4 Pygidium Bulb��us,r��unded, Bulb��us, divided Bulb��us, Bulb��us, divided int�� With additi��nal With additi��nal With additi��nal With 5 �� 6 additi��nal 2 weakly int�� 2 halves divided int�� 2 2 halves, with divisi��ns ��r l��bes divisi��ns ��r l��bes divisi��ns n��w distinct divisi��ns distinct devel��ped halves halves additi��nal divisi��ns devel��ping devel��ping ��r l��bes devel��ping Pygidial cirri Absent 3 ∶ 1 mid-ventral, 3 cirri present 3 �� 5 cirri present 5 cirri present 5 cirri present 5 cirri present 5 cirri present 2 d��rs��lateral Spin��us setae, lyrate 1 �� 2 spines, 1 �� 2 spines 1 �� 2 spines 2 �� 3 spines setigers 3 �� 4 spines setigers 3 �� 4 spines setigers 1- 3 �� 4 spines setiger 1; 4 �� 5 spines setiger 1; , & capillaries setigers 1 �� 4; setigers 1-4; setigers 1 �� 4; 1 ��� 1 �� 4; 1 �� 2 lyrate setae 1 �� 4; 1 �� 2 lyrate setae 4; 2 �� 3 lyrate setae 2 �� 4 lyrate setae all 3 �� 4 lyrate setae all lyrate setae absent; lyrate setae 2 lyrate setae setigers 2-4; 3 �� 5 setigers 2-4; 5 �� 6 setigers 2-4, and setigers fr��m setiger 2; setigers fr��m setiger 2; 2 �� 3 capillaries all absent; 3 �� 4 setigers 3 ��� 4 capillaries all setigers capillaries all setigers present m��st middle 8 ��� 10 capillaries all 12 ��r m��re capillaries setigers capillaries all ��nly; 3 �� 4 and p��steri��r setigers; setigers all setigers setigers capillaries all 6 �� 8 capillaries all setigers setigers ventral gr����ve Absent Initial Medial paired Distinct, surr��unded L��ngitudinal r��w ��f L��ngitudinal r��w ��f L��ngitudinal r��w ��f Large pads in gr����ve devel��pment, pads present by paired pads paired pads paired pads paired pads fr��m setiger 2 t�� with paired pads p��steri��r end Nuchal ��rgans N��t apparent N��t apparent N��t apparent N��t apparent Large, lateral ciliated Large, lateral ciliated Lateral ciliated Lateral, ciliated gr����ves gr����ves anteri��r t�� gr����ves anteri��r t�� gr����ves anteri��r t�� anteri��r t�� d��rsal d��rsal perist��mial ring d��rsal perist��mial ring d��rsal perist��mial ring perist��mial ring
Published as part of Blake, James A., 2015, New species of Scalibregmatidae (Annelida, Polychaeta) from the East Antarctic Peninsula including a description of the ecology and post-larval development of species of Scalibregma and Oligobregma, pp. 57-93 in Zootaxa 4033 (1) on pages 74-75, DOI: 10.11646/zootaxa.4033.1.3, http://zenodo.org/record/289810
{"references":["Ashworth, J. H. (1901) The anatomy of Scalibregma inflatum Rathke. Quarterly Journal of Microscopical Science, 45, 237 - 309. [London]","Hansen, G. A. (1879) Annelider fra den norske Nordhavsexpedition i 1876. Nyt Magazin for Naturvidenskaern, 24, 1 - 17.","Imajima, M. (2009) Deep-sea polychaetes off Pacific coast of the northern Honshu, Japan. In: Fugita, T. (Ed.), Deep-sea Fauna and Pollutants off Pacific Coast of Northern Japan. National Museum of Nature and Science Monographs, 39, 39 - 192.","Bakken, T., Oug, E. & Kongsrud, J. A. (2014) Occurrence and distribution of Pseudoscalibregma and Scalibregma (Annelida, Scalibregmatidae) in the deep Nordic Seas, with the description of Scalibregma hanseni n. sp. Zootaxa, 3753 (2), 101 - 117. http: // dx. doi. org / 10.11646 / zootaxa. 3753.2.1","Blake, J. A. (1981) The Scalibregmatidae (Annelida: Polychaeta) from South American and Antarctic Seas, collected chiefly during the cruises of the R / V Anton Bruun, R / V Hero and USNS Eltanin. Proceedings of the Biological Society of Washington, 94, 1131 - 1162. Available from: http: // www. biodiversitylibrary. org / item / 107603 (Accessed 19 Oct. 2015)","Hartman, O. (1967) Polychaetous annelids collected by the USNS Eltanin and Staten Island cruises, chiefly from Antarctic seas. Allan Hancock Monographs in Marine Biology, 2, 1 - 387.","Schuller, M. (2008) New polychaete species collected during the expeditions ANDEEP I, II, and III to the deep Atlantic sector of the Southern Ocean in the austral summers 2002 and 2005 - Ampharetidae, Opheliidae, and Scalibregmatidae. Zootaxa, 1705, 51 - 68."]}