Discrimination and classification of rising and nonrising pitch patterns by European starling (Sturnus vulgaris)

In a series of four experiments, starlings learned to classify pitch patterns according to pattern structure. In Experiments 1 and 2, the starlings learned to discriminate a series of pairs of four-tone patterns in which one of the patterns rose in pitch and the other pattern did not. The starlings in these initial two experiments did not attend to pattern structure at all. Rather, they discriminated patterns according to the relative pitch of the initial pattern tones. In Experiment 3, the same starlings discriminated the same patterns from Experiments 1 and 2 in a new context that tempered the salience of individual pattern tones. The starlings initially failed to discriminate in the new context, but eventually learned to do so. In Experiment 4, the starlings classified novel pitch patterns according to their pattern structure. These experiments identify a hierarchy of relative pitch strategies, as well as a sophisticated conceptual capacity for the perception and classification of pitch patterns by starlings.