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African savannah elephants (Loxodonta africana) are key ecosystem engineers that migrate over large spatiotemporal scales foraging as they require copious amounts of food and water across habitable landscapes. Therefore a need to understand movement patterns arises in relation to vegetation type and landscape variability, moreso in forage depauparate arid areas such as Gonarezhou National Park (GNP) in Zimbabwe. The objectives of this study were to: (i) assess the performance of vegetation indices in modelling the distribution of African savannah elephants, and (ii) model future landscape variability in Gonarezhou National Park (GNP) in Zimbabwe. Maximum entropy (MaxEnt) algorithm was used to explore the relationship between vegetation indices and distribution of African savannah elephants in the GNP. The Soil Adjusted Vegetation Index (SAVI) performs better relative to other indices in modelling the distribution of African savannah elephants across all habitat types in the GNP. Cellular automata‐Artificial Neural Network (CA‐ANN) showed a significant future decrease (Kruskal Anova; p < 0.05) in landscape suitable to sustain large populations of African savannah elephants in the GNP by the year 2083. Future remote sensing reveals directional insights into the future consequences of current landscape management for African savannah elephant conservation which is a crucial in the sustainability of climate threatened arid protected areas such as the GNP. [ABSTRACT FROM AUTHOR]

A great challenge for African conservationists is human–wildlife conflict outside the boundaries of protected areas (PAs). This is particularly true when considering large mobile species, such as elephants, that often disperse from PAs into community land. In Zimbabwe's Hwange National Park, studies on elephant dispersal have focused on breeding herds. Here, we analyse the movements of five older, large males fitted with GPS collars. We found some dispersal outside Hwange, but large males were largely restricted to the Park, especially during the late dry season when water was scarce. Our findings may be useful to regional conservation decision makers. [ABSTRACT FROM AUTHOR]

Though there is consent that poaching is an important driver of wildlife declines in many African forests, we lack a deeper understanding of the factors behind population patterns in Afrotropical landscapes to inform conservation management. We surveyed line transects in four Cameroonian sites of different protection status in 2013/2014, namely Korup National Park, Banyang Mbo Wildlife Reserve, one timber concession and a section of an unprotected agroforestry matrix, and used data from 2007 for trend assessment. We used GLMs to identify predictors for mammal distribution based on a set of variables. While species compositions showed complex trend patterns, it is noteworthy that chimpanzee and elephant abundances declined significantly in the Banyang Mbo Wildlife Reserve. Although landscape management type affected the distribution of most mammal species, anthropogenic or environmental variables were strong predictors. Chimpanzee, elephant and drill abundances were higher at a larger distance to settlements. Vegetation density positively affected a number of species, including chimpanzee and red colobus. Several species were more abundant in rough terrain, probably because of reduced poaching. Our results suggest that protected areas in Southwest Cameroon had a mitigating effect on wildlife declines, mainly due to their remoteness and habitat quality, rather than management interventions. [ABSTRACT FROM AUTHOR]

Large herbivores have been described as agents of change in terrestrial habitats. Their effect on aquatic ecosystems are, however, underexplored. We raise the question of whether elephants and hippopotamus have the potential to significantly alter limnological properties and, indirectly, primary and secondary productivity within small and shallow freshwaters in arid and semi‐arid African landscapes. In this note we discuss hypothetical means by which elephants and hippopotamus alter shallow freshwater bodies. We further assimilate this with known ways by which these mega‐herbivores alter aquatic environments giving an overview of their potential functional role in structuring aquatic habitats. [ABSTRACT FROM AUTHOR]

Overpopulation in conservation areas with large populations of elephants reduces the diversity of the thicket vegetation of the Eastern Cape Province of South Africa. Monitoring species' specific impacts is a difficult task for reserves. This study investigates whether the Normalised Difference Vegetation Index (NDVI) can be used to evaluate the health of thicket in the Addo Elephant National Park successfully. To achieve this, different browsing intensities of elephants were used to establish a gradient of the full range of utilisation levels on thicket. These were obtained at different distances from the nearest available water sources for Mesic and Valley Thicket types. NDVI values were correlated to an elephant utilisation gradient, which was, in turn, directly related to distance from a water source. The lowest NDVI values (<0.2) were observed adjacent to watering holes where vegetation was completely degraded due to trampling by the daily congregation of animals. The highest NDVI values (>0.5 representing the densest stands of thicket vegetation) were obtained at distances exceeding 7 km from a water source where elephant densities were the lowest within the park. NDVI values could, therefore, be used to identify and monitor gradients of elephant impact on the integrity of thicket. [ABSTRACT FROM AUTHOR]

Equatorial Guinea in central Africa hosts rich biodiversity and a network of protected areas (PAs). However, infrastructure development has facilitated access to previously remote forests. This has likely increased poaching in PAs, thereby complicating efforts of agencies tasked with protecting threatened mammals. Reserva Natural de Río Campo (RNRC) in Equatorial Guinea was previously identified as a priority area for large mammals due to the presence of elephants and great apes and includes habitat for a diverse mammal community of commonly hunted species. To assess mammalian diversity in RNRC, we conducted a camera trap survey in 2017 and 2019. We used a two‐step modelling approach to quantify environmental and anthropogenic factors influencing mammal groups. We detected 32 terrestrial mammal species, including endangered forest elephant, western gorilla, chimpanzee, giant pangolin and white‐bellied pangolin. We found bushbuck and sitatunga closer to human‐dominated areas, while other common species were, in general, further from development. Monkey and pangolin abundance increased inward from the RNRC boundary. Endangered species appear restricted to northeast RNRC which connects to Campo Ma'an National Park in Cameroon. We recommend using our inventory and distributions of threatened mammals as starting points to determine effectiveness of future anti‐poaching and management strategies on mammal populations. [ABSTRACT FROM AUTHOR]

In Africa, most protected forests are in densely human‐dominated landscapes where human–wildlife conflict is intense. We documented farmer perceptions and responses to crop‐raiding wildlife from Kibale National Park, Uganda. Crop raiding was mostly (95%) by baboons (Papio anubis) and elephants (Loxodonta africana). While the financial loss caused by baboons and elephants did not differ, elephants were perceived as more damaging. Guarding and trenches were perceived as the most effective deterrent strategies for baboons and elephants, respectively. Distance from the park boundary and household income were significantly associated with a greater likelihood of crop raiding. Distance from the park, household head age and the species that raided crops, influenced whether a household applied one or more deterrent strategies. Households headed by women or older adults were most vulnerable, experiencing greater losses to raiding. Patterns of human–wildlife conflict around Kibale forest are complex, but the extent of crop damage was mostly determined by distance from the park and farm socio‐economic status and thus their ability to mitigate or deter raiding. Managing crop raiding requires collaboration between the park and affected farmers to ensure that mutually managed deterrent methods, such as trenching (elephants) and guarding (baboons), are effectively shared, applied and maintained. [ABSTRACT FROM AUTHOR]

Elephants face diverse threats from human activities and use temporal and social strategies to reduce human‐induced mortality risk. We used data from camera trap surveys in 2018–2019 (n = 1625 independent detection events from 11,751 sampling days) to investigate elephant responses to anthropogenic risk in the Ruaha‐Rungwa ecosystem, Tanzania. The study was conducted in one low‐risk and three high‐risk sites using 26–40 paired camera trap stations per site. Risk influenced the active periods, use of roads and water sources, social associations and behaviour of elephants. Elephants demonstrated significantly more night‐time and reduced daytime activity in the high‐risk sites relative to the low‐risk site. This higher night‐time activity in the high‐risk sites was observed for both males and females, though it was more pronounced for cow–calf groups than lone males. Foraging events and use of water sources were more frequent at night in the high‐risk sites. Elephants used roads as movement routes in the low‐risk site but avoided roads in the high‐risk sites. Males were significantly more likely to associate with other males and cow–calf groups in the high‐risk sites. Fewer occurrences of relaxed behaviours were observed in the high‐risk sites compared to the low‐risk site. We discuss the potential implications of our findings for elephant survival and reproduction. [ABSTRACT FROM AUTHOR]

African forest elephants (Loxodonta cyclotis) are a critically endangered and visually cryptic species that inhabits Central African rainforests. Using a 1250 sq‐km grid of 50 acoustic sensors in Republic of Congo, we investigated the landscape‐scale behavioural response of forest elephants to poaching events. We detected eight automatic weapon fire events over 1.5 years of recording (2017–2019). We examined detections of elephant vocalisations across grid sites within 10 km of the gunfire events in the 48 h surrounding the gunfire, and in paired 48‐h control periods free of gunfire. Fewer vocalisations were detected before gunfire events than during control periods, suggesting elephants show behavioural responses to poacher presence before any shots are fired. Immediately following the gunfire events, a significant increase in elephant calls was detected, which may indicate increased communication to facilitate group cohesion and movement. Elevated call levels dropped to lower‐than‐baseline rates after several hours, suggestive of a sustained response. These patterns indicate forest elephants respond to both poacher presence and to gunfire events themselves. As these patterns may reflect behaviours that mediate population processes, conservation strategies should account for the potential of poaching to impact populations via indirect effects on nontarget elephants in the area. [ABSTRACT FROM AUTHOR]

Translocation of elephants is used to mitigate human‐elephant conflict in Asia and Africa. However, few studies investigate how translocations affect the movements and social behaviour of individuals following their release, which may have important implications for whether translocated animals survive and succeed. Using GPS‐tracking data, we explored movements of five translocated bull elephants (Loxodonta africana) moved to Tsavo, Kenya, and compared them with five resident bull elephants. Position data was collected hourly for 1 year (March 2018–March 2019), and analysed to investigate home range, displacement rates, problematic behaviour and group size. Of the five translocated elephants, three were illegally killed and one continued to break fences and raid crops. Only one elephant stayed away from human settlement. We found group size and composition to be significantly different, with translocated elephants observed in smaller groups with no female elephant interactions. All elephants showed variation in home ranges and displacement rates, but differences were not significant between resident and translocated elephant groups. For future translocations, we recommend careful consideration of elephant social systems, elephant age, timing, release site and proximity to human settlements that might create human‐elephant conflict. This will improve chance for success of such high‐stake and expensive translocations. [ABSTRACT FROM AUTHOR]

In 2014, the elephant population in the Zambezi valley had reduced by at least 40% as compared to 2001 figures and the elephant density had reduced to 0.69 elephant/km SP 2 sp from 1.13 elephant/km SP 2 sp in 2001 (Dunham et al., 2015). Although anti-poaching efforts are in place (Lunstrum, 2014; Mukwazvure & Magadza, 2014), elephant poaching remains the major driver of elephant decline (e.g. in the Mid-Zambezi Valley, Zimbabwe). Following this decline, mainly related to poaching, anti-poaching strategies have increased within the Mid-Zambezi Valley to combat the illegal killing of elephants. Trends in elephant poaching in the Mid-Zambezi Valley, Zimbabwe: Lessons learnt and future outlook. [Extracted from the article]

African savanna elephants (Loxodonta africana) have been recognised as ecosystem engineers, where their feeding habits have been shown to alter landscapes. Within small, fenced reserves, studies exploring elephant damage on trees and their recovery have overlooked secondary damages that could be contributing to tree mortality. The aim of this study is to assess the significance of both elephant damage and secondary damage, and the subsequent tree recovery. We identified secondary damage as insects and considered wood borers and termites in this study. This was conducted in in the small fenced Karongwe Private Game Reserve, South Africa. We analysed the level of damage, recovery and insect presence using vegetation transects, where all trees ≥2 m in height were surveyed (n = 1278 trees). Forty tree species were recorded, with 5 species accounting for 77% of the data set and used for further analysis. Termites were found to be more likely to colonise damaged trees without signs of recovery. However, wood borers were more likely to colonise damaged trees showing signs of recovery. Termites and wood borer presence on damaged trees was not dependent on tree height. We suggest carefully considering management approaches for elephant‐induced termite and wood borer damage on trees. [ABSTRACT FROM AUTHOR]

Our findings therefore highlight the need for further investigation of elephant population dynamics, distribution and space-use within the Malawi-Zambia Transfrontier Conservation Area to inform conservation management. Cross-border cooperation between Malawi-Zambia and Malawi-Mozambique and is therefore vital to the protection of these elephant populations. To examine potential human-elephant conflict in Malawi, we investigated the GPS locations that fell outside of protected areas and within Malawi's borders using QGIS v.3.12 (QGIS Development Team, 2020). Despite their ecological, cultural and economic importance, African savannah elephant (herein elephant; I Loxodonta africana i ) populations have experienced recent widespread declines (Chase et al., 2016; Wittemyer et al., 2014). [Extracted from the article]

In Togo, the illegal trade of African elephant (Loxodonta africana) ivory is widespread despite efforts made by the government to curtail these activities. By using data gathered from CITES institutions and natural resource management authorities in the country, we investigated the extent of ivory trafficking in Togo. We also conducted surveys in villages around protected areas, which still contain elephant populations to assess the species' current status in the country. Our surveys indirectly corroborated that there are probably no more than 150 elephants collectively within the country's protected areas, most of them in the Fazao‐Malfakassa National Park. We also estimated that a minimum of 41.65 tons of ivory (elephant tusks and carved objects) was intercepted in Togo between 2008 and 2018. Despite the fact that presently illegal elephant hunting is unlikely in Togo, we show that this country is the entry point for ivory from Central Africa (Cameroon and Gabon). We maintain that Togo's position as an intermediary country in the illegal ivory trade is a tangible threat to elephants throughout their range countries and is also likely to affect the dwindling population of these animals in the country if adequate enforcement is not implemented. [ABSTRACT FROM AUTHOR]

The southeastern hinge-back tortoise, I Kinixys zombensis i Hewitt, 1931 (Figure 1a,b), is a sub-Saharan tortoise that may be an important fruit consumer and seed disperser in the coastal forests of eastern Africa. Based on species comparisons, I K. zombensis i may also play a more important role as a seed disperser in its coastal forest ecosystem than savannah-dwelling hinge-back tortoises where vertebrate seed dispersal is dominated by small (Kuechly et al., 2010) and large (Bunney et al., 2017) herbivorous mammals. Seed dispersal by chelonians (i.e. tortoises, turtles and terrapins) is geographically and taxonomically widespread, more so than any other reptilian groups (Falcón et al., 2020). We propose that future studies assess the seed dispersal effectiveness of hinge-back tortoises, like I K. zombensis i , using current frameworks for quantifying the number of seeds dispersed and the quality of seed germination (e.g. Schupp et al., 2010). [Extracted from the article]

This study examined the African elephant's (Loxodonta africana) seasonal space use and movement using elephant telemetry data in Hwange National Park. The adaptive‐local convex hull (a‐LoCoH) was adopted to understand the elephant movement metrics and home range. Paired t test was used to compare the mean seasonal speed of each group (n = 10) of elephants, while the Wilcoxon signed‐rank test was used to determine differences in the 50% and 90% home range. Results show that mean elephant movement speed significantly differed (t = −3.07, df = , p < 0.01) for all elephant groups across seasons. Moreover, significant differences (W = 3, Z = 2.4973, p < 0.012) were observed between core a‐LoCoH home‐range distributions for the wet and dry seasons. However, no significant differences (W = 10, Z = 1.7838, p > 0.05) were detected between the two seasons for the total (90%) a‐LoCoH ranges. NDVI, Distance to water sources and Slope were positively related with elephant movement speed while Aspect and Distance to roads were negatively related with elephant speed. These findings underscore the importance of resource variability in driving elephant movement and foraging behaviour in a semi‐arid savannah ecosystem. [ABSTRACT FROM AUTHOR]

Expansion of human settlements and development have undeniable impacts on wildlife and habitats. Yet wildlife continues to persist in human‐modified environments. While shifting patterns of wildlife landscape use in response to development is well established, less is known about how risk avoidance by animals impacts their resource access. Our research addresses this question by assessing spatial and temporal patterns in access to water resources for savannah elephants (Loxodonta africana) relative to anthropogenic features in the Okavango Panhandle of Botswana. We combined GPS collar data from 39 elephants (2014–2018) with spatial data on permanent water sources and human development to quantify patterns of water access. We modelled a multi‐scale resource selection function that showed elephants selected water access points in areas with less human development (p < 0.0151). Elephants visited water more evenly across day and night in areas with no buildings nearby, but shifted to a significantly more nocturnal pattern in areas of higher building density (ANOVA, p < 0.001). Analysing spatial and temporal patterns of elephant resource access in the context of anthropogenic features provides compelling insight into the potential consequences of human‐modified landscapes on the behaviour, fitness, and persistence of threatened species and our coexistence with them. [ABSTRACT FROM AUTHOR]

Miombo woodland understory vegetation is scarce, it can be found along the few perennial rivers (3% riparian forests) and in post-cultivation regenerating woodland (0.3%) (Montfort et al., 2021). In Laikipia, Kenya, 70% of plants found in the elephant diet correspond to understory vegetation (Coverdale et al., 2016) and low-structure forests are important habitats for elephants in Borneo (Evans et al., 2018). Around the park, vegetation is a mosaic of small fields (<1 ha each), patches of regenerating woodland and open wooded savannahs that have lost their capacity to regenerate miombo woodlands (Figure 2). [Extracted from the article]

Transportation networks can be a major impediment to wildlife movements. We assessed the use of wildlife underpasses and culverts along a newly constructed railway in Kenya's Tsavo National Parks by African elephants (L. africana). We collared ten elephants with GPS satellite transmitters within 20 km of the railway in March 2016 and analysed their movement data to March 2019. Eight elephants used the underpasses although one did not cross the adjacent highway. The remaining two neither used the underpasses nor crossed the highway despite ranging in the vicinity. Their median speed significantly increased to 0.65 km/hr from 0.45 km/hr before crossing the railway, then slowed to 0.32 km/hr after crossing. Females in family groups moved faster than the lone bulls when using the underpasses. Seventy‐eight per cent of all crossings made were at night. The fast speeds and the nocturnal patterns are behavioural responses of elephants in risky landscapes or under stress. Disturbance from vehicles traffic on the adjacent highway and from newly developed human settlements may have limited use of underpasses. Wildlife crossing structures, signage and speed bumps along the highway; relocation of the illegal human settlements; and inter‐agency coordination are requisites for enhancing Tsavos' elephant habitat connectivity. [ABSTRACT FROM AUTHOR]

For the first time in Togo, we used camera trapping to investigate the mammal community in Fazao‐Malfakassa National Park (FMNP), a forest and savannah mosaic landscape subject to poaching and other detrimental anthropogenic activity. We compiled a species inventory, that is species occurrence, habitat use and activity patterns during dry and wet seasons. Based on images from >80 locations during 9,007 camera days, we identified 32 mammal species, which, when combined with other published studies, increases the total number of mammals (excluding bats) historically reported to 57 species. Our results confirm the presence of five mammal species evaluated as threatened according to the IUCN Red List, highlighting the significant conservation value of the FMNP. Specifically, it appears to be the only protected area in Togo where the African savannah elephant and the African forest elephant occur sympatrically, and only the second site currently known with direct observations of Walter's duiker. We also report the presence of poachers and other anthropogenic activity within the FMNP. We recommend that continued survey efforts should be combined with detailed ecological data collection, effective law enforcement, community outreach and eco‐tourism development, to safeguard the remaining mammal species communities necessary for a functioning ecosystem in the park. [ABSTRACT FROM AUTHOR]

Using local ecological knowledge to improve the effectiveness of detecting white-bellied pangolins (Phataginus tricuspis) using camera traps: A case study from Deng-Deng National Park, Cameroon Here, we evaluate the utility of local ecological knowledge (LEK) in tailoring camera-trap surveys, a detection method increasingly used for pangolins, to improve detection efficiency for occurrence and ecological studies for the white-bellied pangolin. For example, across 18 camera-trap surveys in the predicted range of white-bellied pangolins, 2,287 camera traps produced 275 five-day trap occasions with detections of this species (Khwaja et al., 2019). The results we present here are preliminary, and we recommend a more rigorous systematic study to compare the detection rates of white-bellied pangolins using different survey designs (e.g. random, standard-grid and arboreal placement) with a LEK-informed survey design at the same place, as well as investigating pangolin preference for particular tree species. [Extracted from the article]

African forest elephants (Loxodonta cyclotis Matschie, 1900) are ecological engineers that play a fundamental role in vegetation dynamics. The species is of immediate conservation concern, yet it is relatively understudied. To narrow this knowledge gap, we studied the drivers of daily movement patterns (linear displacements) of forest elephants—characterised by a set of geographical, meteorological and anthropogenic variables—in the Odzala‐Kokoua National Park, Republic of Congo. Explicitly, we used conditional random forest to model and disentangle the main environmental factors governing the displacements of six forest elephants, fitted with GPS collars and tracked over 16 months. Results indicated that females moved further distances than males, while the presence of roads or human settlements disrupted elephant behaviour resulting in faster displacements. Forest elephants moved faster along watercourses and through forest with understory dominated by Marantaceae forests and bais, but moved slower in savannahs. Finally, flood‐prone areas—described by elevation and accumulated precipitation—and higher temperatures prevented longer displacements. We expect these results to improve the knowledge on the species movements through different habitats, which would benefit its conservation management. [ABSTRACT FROM AUTHOR]

African elephants have major impacts on vegetation, particularly at high densities. Knob‐thorns (Senegalia nigrescens) are typically ring‐barked by elephant, and high levels of mortality are common at high elephant densities. Our study aimed to test whether ivory palm clusters (Hyphaene petersiana) form a biotic refuge for knob‐thorn against elephant herbivory. We measured the density, damage and mortality of knob‐thorns in sites differing according to ivory palm presence and elephant density, and thus, the probability of knob‐thorn encounter by elephants. The site with palms and low elephant density, had a high density of knob‐thorns, but lower proportions of damaged and dead trees, than sites without palms but with similar or higher elephant density. In the former, knob‐thorns were associated with palm clusters, particularly saplings and young adults. In this site, low proportions of damaged and dead knob‐thorns were recorded in palm clusters, compared with outside clusters, and to those in the other sites. Our study also showed that juvenile palms which protected knob‐thorns, suffered low mortality in contrast to subadult palms. We have no evidence but implicate elephants and suggest that in palm clusters, subadult palms are more accessible to elephants than knob‐thorns because of the different methods of utilisation. [ABSTRACT FROM AUTHOR]

Elephant populations are in decline across the African continent, but recent aerial surveys show that populations in Uganda are increasing. However, threats such as poaching and habitat disturbance remain. Having a comprehensive knowledge of the ranging behaviour of Ugandan elephants is crucial to understanding where critical habitat for the species occurs. We investigated various aspects of ranging behaviour of 45 radio‐collared elephants (Loxodonta africana) in three areas—Queen Elizabeth Protected Area (QEPA), Murchison Falls (MFPA) Protected Area and Kidepo Valley (KVCA) Conservation Area. We also set Ugandan analyses in a continental context by comparison with home ranges reported in published literature. Elephants within KVCA had larger core ranges than elephants in QEPA or MFPA. Wet season ranges in KVCA were much larger than dry season ranges. The most important core areas in all three national parks were centred around water resources. Home range size was negatively correlated with net primary productivity (NPP) at Ugandan (N = 39 individuals) and continental (N = 17 sites) scales. This study indicates that, at a local scale, factors such as water source location are important in shaping elephant ranging behaviour. At larger scales, factors such as NPP are good predictors of elephant home range size. [ABSTRACT FROM AUTHOR]

Understanding the key drivers that influence the potential distribution of herbivore species in changing landscapes has been at the centre of enquiry in wildlife science for many decades. This knowledge is particularly important for keystone species like the African elephant (Loxodonta africana) whose population is declining even in conservation areas. The Sebungwe Region is part of the Kavango‐Zambezi Transfrontier Conservation Area and supports ~4,000 elephants. The Sebungwe Region has lost an estimated 76% of its elephant population over the years. This study aimed to determine how the elephant distribution in the Sebungwe Region was affected by human settlement and whether the patches for elephant distribution were large enough for elephant habitation. The prediction of the potential distribution of the elephant was based on presence‐only data modelled through an ensemble algorithm that combined several candidate models to enhance predictive ability. We observed that human settlement drives the potential distribution of elephants in the Sebungwe Region (test AUC = 0.95), and patches from the model were on average <1.5 km2. Our results provide initial insights into the key habitat factors that drive distribution of elephants in the Sebungwe landscape. Future conservation of the elephant could benefit from our study through systematic planning of settlements, which might help minimise human interaction with wildlife. [ABSTRACT FROM AUTHOR]

Specifically, ants forage for the honeydew that scale insects produce from tree sap, and, consequently, they may defend either the scale insect or the tree tissue from potential herbivores. We also tested how ants on the trees reacted to a perceived threat in order to learn whether or not the ants may defend the scale insects. Our survey of I S. nigrescens i trees across the herbivore exclosure treatment and exposed plots revealed the presence of ants, most of which were I Polyrhachis i sp. I , i in both unprotected and herbivore-protected trees (Figure b). Since the scale insect was only found in trees unprotected from herbivores, this suggests there may be a complex relationship between the plant, the ants, the scale insect, and herbivores. [Extracted from the article]

Dung counts were first used in the 1940s (Bennett, English, & McCain, [5]), and are mostly employed in woodland and forested habitats, where direct counts are difficult to apply (Acevedo et al., [1]). Mean dung persistence time of Cape buffalo dung was determined in JWPF in order to estimate dung density (Laing et al., [9]). The distribution of dung piles in different habitats includes the following: open forest = 14 dung piles (wet = 8; cry = 6), dense forest = 12 dung piles (6 dung piles during both seasons), plantation forest = 11 dung piles (wet = 7; dry = 4) and riparian forest = 13 dung piles (wet = 6; dry = 7). Hence, further follow-up of the remaining dung piles was stopped as reported by Barnes and Jensen ([3]) assuming that the dung decay time of the remaining dung piles has insignificant impact on the mean dung persistence time. [Extracted from the article]

The ability of animals to adapt to their changing environment will depend in part on shifts in their ranging patterns, but when and why individuals choose to move requires detailed understanding of their decision‐making processes. We develop a simple decision‐making model accounting for resource availability in habitually used ranges. We suggest that disparities between model predictions and animal tracking data indicate additional factors influencing movement decisions, which may be identified given detailed system‐specific knowledge. The model was evaluated using movement data from satellite‐tracked elephants (Loxodonta africana) inhabiting the Amboseli basin in Kenya, moving from savannah areas with low quality but constant resource availability, to areas with temporally constrained higher nutrient availability. Overall, the model fits the data well: There was a good correlation between predicted and observed locations for the combined data from all elephants, but variation between individuals in how well the model fits. For those elephants where model predictions were less successful, additional factors likely to affect movement decisions, including reproduction, anthropogenic threats, memory and perception are suggested. This protocol for building and testing decision‐making models should contribute to success in attempts to preserve sufficient space for large herbivores in their increasingly human‐dominated ecosystems. [ABSTRACT FROM AUTHOR]

As the human footprint expands worldwide, people and wildlife are coming into greater contact, and areas of human activity may be simultaneously associated with risk and reward for animals. To avoid human threats while exploiting opportunities, animals may adjust their spatiotemporal activity, using areas of anthropogenic disturbance at night when people are less active. We combined four camera trap datasets from Mozambique's Gorongosa National Park to evaluate the effects of roads and settlement on diel activity patterns of elephants (Loxodonta africana). We found high rates of elephant activity along the boundary of the park, where elephants can access cultivated crops, and along roads, which serve as movement corridors. However, elephants restricted their activity to the night and crepuscular periods in these areas of human disturbance, seeking refuge in the interior of the park away from roads and settlements during the day. Our findings suggest that a history of killing and antagonism has instilled a fear of humans in this elephant population, with implications for research, tourism and human–elephant coexistence. Our study highlights the utility of camera traps in monitoring human–wildlife conflict and habituation and demonstrates the value of integrating disparate camera trap datasets for comparative analyses on landscape or even continental scales. [ABSTRACT FROM AUTHOR]

Human encroachment on wildlife habitats is leading to increased fragmentation; hence, there is an increasing focus on improving connectivity between remaining habitat. Large, wide‐ranging species such as the African elephant, (Loxodonta africana), are particularly vulnerable due to their extensive habitat requirements. Wildlife corridors have been created to facilitate movement, with little knowledge to date on whether they serve their intended function as transit routes, or whether they simply extend the available habitat for occupancy. We collected data on elephant behaviour in the Mount Kenya Elephant Corridor, with the aim of quantifying the utility of the corridor. A grid of 25 camera traps was used to survey the 478 ha corridor over 11 weeks. Cameras recorded over 43,000 photos with 694 separate events triggered by elephants. Patterns of use varied spatially and temporally, indicating that certain areas were treated as habitat extension, while others were predominantly for transit. These differences were likely due to variation in vegetation cover and levels of human disturbance. Corridor use differed amongst individuals, suggesting that use may depend on both the characteristics of the corridor itself and the social or resource needs of individual elephants. [ABSTRACT FROM AUTHOR]

The utility of aerial survey data in modelling the distribution of wildlife has been questioned due to locational errors. Yet, little effort has been made to correct these errors. Here, a new method is proposed for correcting locational errors associated with aerial survey data. We further tested whether the correction improves model performance using elephant aerial survey data collected at three protected sites in Zimbabwe. Our correction method succeeded in shifting location by 143–177 m from the line of flight. Based on the true skill statistic, models built using corrected data in Hwange and Mana Pools national parks were significantly better than those from uncorrected data and only in Gonarezhou national park were no significant differences in model performance observed. These results underscore the benefit of applying the new correction method in modelling wildlife distribution. The other key result is that the correction seems to be more beneficial in heterogeneous landscapes than homogeneous ones for in the former, a small shift in location results in the sampling of different habitats than where sightings were made. The proposed method opens new opportunities for improving the accuracy of aerial survey data in species distribution modelling. [ABSTRACT FROM AUTHOR]

Hippos are gregarious animals, generally occurring as pods comprising females and their young, a dominant male and subordinate males (Klingel, [14]). There are photographs/videos showing hippos interacting with hippo carcasses (Breen, [4]; Harris, [10]; Hippo keeps crocs from dead juvenile, [11]; Hippopotamus (Hippopotamus amphibius), [12]; ViralHog, [20]) and defending carcasses from other animals (Csaba, [6]; Harris, [10]; Hippo keeps crocs from dead juvenile, [11]); providing anecdotal evidence that hippos may also display epimeletic behaviour towards dead conspecifics. The aggressive behaviour of the hippo towards the crocodiles (VideoS2) is also uncommon; hippos and crocodiles generally share this pool without conflict (personal observation). [Extracted from the article]

Abstract: Human–elephant conflict is a common conservation problem throughout Africa, but poorly studied where forest elephants (Loxodonta cyclotis) occur. Here, we investigated people's perceptions towards the impact of elephants around Nouabalé‐Ndoki National Park, northern Congo. We aimed to understand the perceptual differences amongst residents of four villages that varied substantially in the degree of conservation benefits received. We used a multivariate analysis to investigate how socio‐economic variables, such as employment, wealth, education and ethnicity, influenced perceptions. Using both quantitative and qualitative data, we found that the majority of respondents experienced elephant impacts, mainly through crop raiding. Residents of the village where the local conservation project is based had significantly more positive perceptions of elephants, whereas perceptions of farmers were mostly negative. We identified some misunderstandings regarding the responsibilities of elephant impact mitigation regarding benefit sharing, stakeholder involvement and lawbreaking, but also willingness to apply mitigation measures. To mitigate the prevalent impact of elephants, a comprehensive approach of measuring impact, exploring community‐based mitigating strategies and understanding of the root causes of the conflict is crucial. We propose a long‐term programme within logging concessions organizing groups of farmers to cooperate on mitigation techniques and explore alternative incomes. [ABSTRACT FROM AUTHOR]

Abstract: The African elephant, Loxodonta africana, is under threat from habitat loss, poaching and human–elephant conflict. To mitigate for impact of habitat loss and reduce conflict, connectivity between elephant habitats can be improved through the protection of corridor areas. This study looks at elephant distribution and movement patterns within the Kasigau Wildlife Corridor (KWC) within the Tsavo Conservation Area in South‐east Kenya. Elephant presence data were obtained from observations by rangers during routine patrols across KWC, and were analysed in MaxEnt. The environmental factors predicting elephant distribution and density were tested, as well as the relationship between elephant maximum entropy and the presence and abundance of other wildlife. Seasonal variations in temperature and precipitation, plus presence of waterholes were found to play significant roles in elephant distribution across KWC. Higher elephant densities were not found to correlate with lower densities of other wildlife species; indeed, during the dry seasons, elephant presence was associated with greater wild herbivore densities. Besides illustrating the importance of the KWC for elephant conservation in the Tsavo ecosystem, both as a key corridor and habitat, this study also hopes to highlight the untapped utility of routine ranger patrol data, and encourage the use of such presence‐only data for deducing important knowledge for conservation of biodiversity. [ABSTRACT FROM AUTHOR]

This is the first study where elephant footprints as habitat for aquatic macroinvertebrate communities were assessed. Preliminary observations during the dry season in Kibale Forest, Uganda, indicated that water-filled footprints constituted the majority of stagnant ponds. Consequently, this study aimed at giving an overview of the diversity and ecology of those habitats and the capacity of elephants as ecosystem engineers. The fauna and abiotic factors (age, size, substrate, organic matter, pH, canopy cover, temperature, conductivity) of 30 water-filled natural elephant footprints were sampled, resulting in the record of 61 morphospecies among 27 families/orders. Species composition was dominated by Hydrophilidae and Dytiscidae and influenced by environmental variables, such as age and organic matter. To study the colonization process, 18 artificial footprints were created within different distances from the water source. After 5 days, 410 specimens were collected, with higher species richness in artificial footprints closer to a natural water source. We conclude that colonization of water-filled footprints is fast, they constitute important habitats with high diversity and variability, and they act as stepping stones for dispersal and add to the ability of elephants as ecosystem engineers. We emphasize the importance of elephants as a key species in ecosystem dynamics and conservation practice. [ABSTRACT FROM AUTHOR]

Knowledge of the distribution and nutrient values of key resources supporting the survival of wildlife species is integral for an effective conservation planning and management of the species. In the Miombo ecosystem of the Ugalla Game Reserve, African elephants ( Loxodonta africana Blumenbach 1797), eat soil, that is geophagy, from certain termite mounds. We mapped that all the geophagic termite mounds are exclusively situated in the flood plain. To understand why soils from some termite mounds are eaten, we collected and analysed soil samples from 10 geophagic termite mounds, seven nongeophagic termite mounds and 13 samples from the surrounding flood plain. Percentage of clay content did not differ significantly among the soil samples. Soils from geophagic termite mounds were richer in mineral elements compared with other soil samples. The results demonstrate that the driver for geophagic behaviour is related to rich mineral element contents found in geophagic termite mounds made of the mineral-enriching termites (Macrotermes). Thus, geophagic termite mounds play a role in elephant's dietary needs and possibly influence their movement patterns in Ugalla, as the elephants cannot obtain enough minerals from their feeds. Geophagic termite mounds should be protected from potential destructive land uses, such as airstrip construction. [ABSTRACT FROM AUTHOR]

The article presents a study which aims for the evaluation of the human-wildlife conflicts (HWCs) in Maramani Communal Area (MCA), Zimbabwe. The study uses the qualitative analysis for the identification of perceived wildlife species that cause damage. Results show that damage-causing animals in MCA include spotted hyena, elephant, and baboon.

In many elephant populations in Africa, adverse effects of poaching including altered age and sex structure and long calving intervals continue to negatively impact the rate of recovery. The study reported here characterized the demographic status of the elephant population in Meru National Park, Kenya, so as to determine the size, age and sex structure, and calving interval of the population and to compare its demographic parameters against those from the relatively stable elephant population at Amboseli. Additionally, a demographic study conducted 7 years earlier enabled us to determine the demographic performance of the population and to explicitly test the utility of the individual identification technique in monitoring intermittently studied populations. We found 392 elephants in the park. The proportion of older individuals was lower, and the calving interval was longer than estimates from Amboseli. We recognized 16% of elephants indexed in the previous study. Our results suggest that the individual identification technique may be useful in monitoring demographic changes including poaching in intermittently studied elephant populations. Overall, our results suggest that although the Meru elephant population showed signs of recovery, poaching continues to imperil its recovery. [ABSTRACT FROM AUTHOR]

We examined effectiveness of African savannah elephant dung as a protective barrier for seeds of three tree species, Acacia tortilis Hayne, Tamarindus indica L. and Ximenia aegyptiaca L. Seeds were collected from dung and underneath fruiting trees in Tarangire National Park, Tanzania. Experimental treatments were established to test: (i) the efficacy of dung in protecting seeds of A. tortilis from bruchid beetle infestation and the role of animals larger than insects in removing seeds; (ii) the same tenets as in Experiment 1, using seeds of T. indica; and (iii) the effect of distance on survival of seeds of X. aegyptiaca. Sites were established during two field seasons underneath conspeci-fic trees, where seed predation was likely highest. Repeated-measures two-way ANOVA indicated that there was no treatment effect for Experiment 1. For Experiments 2 and 3 in October 2013, seeds in dung experienced less beetle infestation than fresh seeds. Repeated-measures two-way ANOVA and Tukey's HSD indicated that treatment effect differed among all treatments. Passed seeds at distances ≥ 5 m experienced less beetle infestation than fresh seeds underneath conspecifics. African savannah elephants appear to be important seed dispersers of these three tree species. [ABSTRACT FROM AUTHOR]

Group dynamics related to distribution of African elephants in the semi-arid environment of Tsavo East National Park, Kenya was studied between 2007 and 2010. I examined the seasonal distribution of lone bulls, bull groups, family units and mixed groups and group size. Lone bulls were widely distributed in the dry season and localized in the wet season, whereas bull groups were localized in both seasons. On average bull group size did not differ with seasons or areas, but larger groups were observed in preferred areas in the dry season. Family units were widely distributed in the dry season and localized in the wet season, whereas mixed groups were localized in both seasons. Although family units and mixed groups tended to be large in all areas in the wet season, large groups were also observed in the dry season. Large family units were widely distributed, whereas large mixed groups were localized. The relationship between group dynamics and distribution was attributed to the combination of social, ecological and poaching factors. This study provided insight into distribution patterns that can be applied to the security of the Tsavo elephant population. [ABSTRACT FROM AUTHOR]

The article presents a study which examines the populations of Aloe (A.) ferox in the absence of black rhinos and elephants. The study uses the demographic patterns for the populations of A. ferox exposed to the different pressures of herbivore. Results show the absence of demographic bottleneck at the domestic herbivore-dominated sites.

Understanding factors influencing large herbivore densities and distribution in terrestrial ecosystems is a fundamental goal of ecology. This study examined environmental factors influencing the density and distribution of wild large herbivores in Gonarezhou National Park, Zimbabwe. Vegetation and surface water were predicted to have a stronger influence than anthropogenic-related disturbances (livestock grazing, fires, settlements and poaching) on the density and distribution of wild large herbivores. Aerial survey data for seven common wild large herbivores conducted in 2007 and 2009 and environmental data were collected. Only grass cover explained a significant proportion of the variation in large herbivore densities and distribution. Moreover, only two species densities significantly differed across the Gonarezhou, namely impala and zebra. In contrast, buffalo, elephant, giraffe, kudu and nyala densities did not differ significantly across the Gonarezhou. Overall, the findings only partly support the study prediction. The study results suggest the need to further investigate the roles of environmental factors at smaller scales in order to tease out their relative strengths in influencing density and distribution of large herbivores. [ABSTRACT FROM AUTHOR]

Long-term solutions to crop raiding by elephants ( Loxodonta africana) should be based on an understanding of their behaviour and ecology. The real and perceived risks from humans have been shown to affect elephant behaviour. This is evidenced by elephants predominantly raiding crops at night, avoiding the height of human activity. If such human avoidance behaviours are apparent, it might also be expected that elephants avoid risks associated with higher visibility and increased human activity as may occur during the full moon. However, elephant nocturnal crop-raiding behaviour in relation to lunar cycles has largely been a neglected factor in studies of human-elephant interactions. In this study around Mikumi National Park, Tanzania, we apply circular statistics in this context for the first time to show a significant decrease in crop raiding during the full moon and apply this method retrospectively to data from another site in West Africa with similar results. Additionally, a greater proportion of farms raided was guarded during the full moon than any other moon phase. Our results indicate that variations in crop raiding with lunar phase could be a general feature of elephant behaviour and thus could be used to design and time mitigation efforts. [ABSTRACT FROM AUTHOR]

Elephants have a major influence on vegetation structure, composition and ecosystem processes, and are primary agents of habitat change in Africa. At moderate-to-high population densities, elephants can damage vegetation, especially when enclosed in protected areas. This study examines the effects of elephant browsing on woody trees in Majete Wildlife Reserve ( WR), south-western Malawi. Regression analysis is used to assess the associations of six factors known to drive elephant browsing in other areas and determine which ones have the most influence on browsing at Majete WR. Twenty-four per cent of tagged trees had been subject to elephant browsing. The model with vegetation type, stem diameter and distance from permanent water correctly predicted browsing for 80% of the observations. Elephants mostly favoured riparian woodlands, followed by Acacia-dominated woodland and Brachystegia-dominated woodland. Browsing occurrence was negatively related to distance from permanent water and diameter at breast height( DBH). A larger number of trees, sampled at random and covering a larger portion of the reserve would provide more reliable estimates of browsing and related factors. Knowledge of time- and site-specific factors affecting elephant browsing can be used to forecast future habitat transformations and manipulate the range of the elephants within the reserve. [ABSTRACT FROM AUTHOR]

Since 1993, members of the national wildlife society have undertaken annual surveys of large mammals in the Zambezi alluvial woodlands of Mana Pools National Park, Zimbabwe. Data are collected along 36 systematically-arranged transects. We provide the first thorough assessment of the data from any survey within this long-term project. The transect data from 2011 were analysed with DISTANCE software to assess if the data were suitable for determining the densities of large mammals using distance sampling techniques. Successful application of distance sampling depended on observers using printed, large-scale, georeferenced satellite images onto which they mapped the location of animal groups detected. The assumptions of the distance sampling were well met and thus the 2011 survey provided reliable estimates of the densities of nine species of common large mammals on the Zambezi alluvium during the late dry season. Estimated density in this dry-season concentration area varied from 3.6 km−2 for kudu, to 204 km−2 for impala. The precision of the estimates ranged from a coefficient of variation of 7.9% for elephant, to 25.5% for buffalo. For elephant, warthog and baboon, the morning and afternoon densities differed significantly. [ABSTRACT FROM AUTHOR]