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The parasites of kob based on physical, faecal and blood examination has revealed that kob harbor ticks which include Rhipicephalus evertsi, Rhipicephalus nymph, Amblyomma variegatum; lice, Damalinia sp; and fed upon by by biting fly Culicoides. sp. Helminth ova found were Strongyl sp. and Fasciola. sp.; protozoan. Babesia sp. in blood and Eimeria sp. in faeces. The significance of tick as vector of parasites of domestic animals and the potential risk of kob as reservoir of the parasites these vectors transmit are discussed. The prevalence of Strongyl sp. and Fasciola sp. ova as reflection of worm burden and potential pasture contamination are highlighted

Assessing the population status of wildlife is useful for enhancing biodiversity management strategies in protected areas. This study was carried out within the framework of the Faro National Park (FNP) management plan. The objective was to provide baseline data on the status and population dynamics of the Buffon's kob (Kobus kob kob). The surveys were conducted in the month of June at the beginning of the rainy season. Data were collected along 145 line transects of 2.5 km apart and analysed using Distance 7.2 and Quantum GIS 3.4 software. The Buffon's kob population was estimated to comprise around 4,094 ± 1,303 individuals, with an overall density of 1.24 individuals/km². When compared to results from a study conducted in the year 2000, the concentration of Buffon's kob was higher in the northern part of the national park as opposed to the southern part, as was previously recorded. The population trend shows an extensive decline of approximately 80% compared to records of the year 2000. There was also a weak negative correlation between Buffon's kob density and human activities in the FNP.

The waterbuck (Kobus ellipsiprymnus), though widespread throughout Africa, is suspected to be declining overall. Data on population numbers and structure are lacking for many parts of its range, especially in West Africa, where the subspecies defassa is found. The aim of the present study was to evaluate the abundance, distribution and attributes of waterbuck populations from the Nazinga Forest Reserve, southern Burkina Faso. We investigated waterbuck population trends in the park using transect data collected in 1985–2019. For the more detailed analyses of population structure and distribution of the animals, we used census data gathered during 2019. Most animals were adults (46.6%), and the sex ratio was heavily skewed towards females (5:1). Most animals were concentrated along the larger rivers. There was no influence of poacher activity on waterbuck distribution. In the long term (1985–2019), the population dynamics of waterbuck can be roughly divided into two main periods: a phase of population increase from 1985 to 2005, and one of ongoing population collapse from 2007 to 2019. Although the declining population trend was obvious, coefficients of determination were low indicating that the years explained poorly the number of individuals and the number of sightings obtained. Waterbuck numbers in the Nazinga Forest Reserve are declining, but we found no single reason to explain this trend. It is likely that a combination of factors, including global warming (increased aridity) and illegal activities such as poaching, is responsible. Because there are probably multiple reasons for the observed waterbuck population decline in our study area, we suggest that a multifaceted approach should be adopted in order to enhance the conservation status of the local waterbuck populations.

White-eared Kob, Kobus kob leucotis, Lichtenstein & Peters, 1853, is known to occur in the Gambela-Boma landscape in western Ethiopia and South Sudan. They live in herds and are generally found near water, in such places as plains, woodlands, swamps, and flood plains. We deployed 36 camera traps in Omo National Park and one of them took two images of a White-eared Kob. This is the first documentation from Omo National Park and showing that its range extends further to Omo than previously known and, therefore, the entire area (Gambella to Omo) can be considered as a range extension.

Species delimitations by morphological and by genetic markers are not always congruent. Magnolia kobus consists of two morphologically different varieties, kobus and borealis. The latter variety is characterized by larger leaves than the former. For the conservation of M. kobus genetic resources in natural forests, the relationships between morphological and genetic variation should be clarified. We investigated variations in nuclear microsatellites, chloroplast DNA (cpDNA) sequences and leaf morphological traits in 23 populations of M. kobus over the range of species. Two genetically divergent lineages, northern and southern were detected and their geographical boundary was estimated to be at 39°N. The northern lineage consisted of two genetic clusters and a single cpDNA haplotype, while the southern one had multiple genetic clusters and cpDNA haplotypes. The northern lineage showed significantly lower genetic diversity than the southern. Approximate Bayesian computation indicated that the northern and southern lineages had experienced, respectively, population expansion and long-term stable population size. The divergence time between the two lineages was estimated to be 565,000 years ago and no signature of migration between the two lineages after divergence was detected. Ecological niche modeling showed that the potential distribution area in northern Japan at the last glacial maximum was very small. It is thus considered that the two lineages have experienced different population histories over several glacial-inter-glacial cycles. Individuals of populations in the central to northern part of Honshu on the Sea of Japan side and in Hokkaido had large leaf width and area. These leaf characteristics corresponded with those of variety borealis. However, the delimitation of the northern and southern lineages detected by genetic markers (39°N) was not congruent with that detected by leaf morphologies (36°N). It is therefore suggested that variety borealis is not supported genetically and the northern and southern lineages should be considered separately when identifying conservation units based not on morphology but on genetic markers., ファイル公開：2020/11/01

Assessing human-induced threatening factors to wildlife is the basis in determining and proposing appropriate conservation measures. The present study was conducted to assess impacts of human activities on wildlife in Gambella National Park, Southwest of Ethiopia, from October 2015 to March 2016, focusing on the case of Nile Lechwe (Kobus megaceros). The data in this study were gathered using questionnaires with structured interview and focus group discussion. Data on the threatening factors were compared among villages using chi-square test in SPSS version 20 software. Out of the 384 respondents, 139(36.2), 51(13.2) 49(12.8), 43(11.2) and 41(10.7%) of them informed that agricultural investment, illegal hunting, overgrazing, rice cultivation in the area and habitat loss, respectively, were the most predominant human factors affecting Nile Lechwe. Thus suggests that agricultural expansion and illegal hunting are those human activities with the highest impact on the Nile Lechwe. Before designing and implementing any development investment particularly large scale agricultural expansions, the government and other stakeholders should give consideration and attention to the rapidly declining natural resource beside to the development. Therefore, designing appropriate eco-friendly management with options must be adopted to mediate the effects and minimize future impacts. Key words: Agriculture, anthropogenic disturbance, habitats, Nile Lechwe, Gambella National park, wildlife.

Habitat degradation and fragmentation are eating deep into conservation areas and this is a serious threat to species diversity and abundance. Species like the antelopes have a sedentary and docile nature which makes them highly vulnerable to habitat degradation or human intrusion. The effect becomes complex as the remaining flora and fauna communities can be significantly impacted by changes in ecosystem structure and function. Population density, diversity and abundance of fauna species will either increase or decrease over time depending on the quality of the environment/habitat and the level of human interference or disturbance. Hence an updated checklist of species diversity and abundance is necessary to enable management and other stakeholders make pragmatic plans and policy towards sustainable species conservation. With the aid of a Global Positioning System (GPS), a 5 km transect was established per site and censured for Antelope species using the King Census method of enumeration. Descriptive statistics and ANOVA was used to analyze the data. Seven (7) species of Antelopes were recorded. Kobs (Kobus kob) were the most abundant (2019), while Reedbuck (Redunca redunca) was the least abundant with twenty-five (25) individuals. Kob is the most observed species in Oli Complex with 24.13%, ranking about 50% of kob in proportion. This was followed by roan antelope (Hippotragus equinus), and Red Flanked duiker, 4.02% and 3.63% respectively. Kobs had the highest density of 40.38 per square km followed by roan antelope (3.32) and RF duiker (2.36). Relative density followed a similar trend. The least encounter rate was observed in Sylvicapra grimmia (0.02) and increse further to Hippotragus equinus (0.4), Redunca redunca (0.06) and Alcelaphus buselaphus (0.09) respectively. It was low amongst Tragelaphus scriptus (0.2), and moderate, while it was very high amongst the kobs (5.0). The rate of encountering an antelope in the park is very high at a rate of 6.2 animals per kilometer. Species of antelopes are almost not found in other ranges due to anthropogenic activities around the park. These activities are fast entering into the core area of the park. Hence management should take effective measure to curb this fast-rising problem.

Background Brucellosis is a contagious zoonotic disease of great public health and economic significance especially in developing countries. The disease affects humans and several species of livestock and wildlife. Studies on Brucellosis in wildlife in Uganda have been limited to single populations particularly in Queen Elizabeth National Park. This study aimed at estimating the percentage of positive samples of Brucella spp. in wildlife in four major national parks of Uganda. This was a retrospective survey which utilized archived samples collected from wildlife during the annual disease surveillance activities between 2013 and 2017. Results A total of 241 samples from seven species namely African buffalo (Syncerus caffer, n = 109), African elephant (Loxodonta africana, n = 22), giraffe (Giraffa camelopardalis rothschildi, n = 41), Uganda kob (Kobus kob thomasi, n = 36), lion (Panthera leo, n = 6), plain zebra (Equus quagga, n = 25), and bushbuck (Tragelaphus scriptus, n = 2), were tested for antibodies using the Rose Bengal Plate Test. The overall percentage of positive samples in the four national parks was 31.1% (75/241; 95% CI: 25.6–37.2). Kidepo Valley National Park had a significantly higher percentage of positive samples of 55.9% (19/34; 95% CI: 39.5–71.1) compared to other sampled national parks (p p Brucella spp. detected in African elephant and bushbuck. Conclusion This study shows variations in percentage of positive samples with Brucella spp. between species and across national parks and notably a high percentage with Brucella spp. in wildlife in Uganda than that recorded elsewhere in sub-Saharan region of Africa. Potential for transmission to other wildlife and spill over to livestock is high especially in national parks with high livestock-wildlife interaction.

This study investigated feeding preference and habitat association of waterbucks in Nechisar National Park from November 2016 to August 2017 by total count and direct observation methods. From this survey, 20 plant species were observed to be the food items consumed by waterbucks. Waterbucks were mostly grazers than browser. The plant species consumed by waterbucks was highly significant between seasons (χ2 = 121.34, df = 1, p < 0.05 ). Out of 20 total plant species consumed, annual grass (11.5%), Leersia virginica (8.4%), and Cynodon dactylon (8.4%) were the most frequently consumed food items, but Tamarindus indica (2.9%), Balanites aegyptica (3.3%), and Acacia polycantha (2.36%) were the least consumed food items. There was significant difference between plant species consumed during both seasons (χ2 = 177.67, df = 19, p < 0.05 ). The Shannon–Wiener diversity index result revealed that there were different varieties of food items for waterbucks in dry season (H’ = 2.9) than in wet season (H’ = 2.6). Young leaves comprised the largest proportion of plant parts consumed by waterbucks compared to others. There was a highly significant difference in feeding plant parts in both seasons (χ2 = 88.5, df = 7, p < 0.05 ). Waterbucks spent more time on feeding in the morning and late afternoon during both seasons. The total number of waterbucks in each habitat varied due to food availability in different seasons. Different conservation measures should be taken against waterbuck reduction and create appropriate environment for waterbuck.

Savannah tsetse flies avoid flying toward tsetse fly-refractory waterbuck (Kobus defassa) mediated by a repellent blend of volatile compounds in their body odor comprised of δ-octalactone, geranyl acetone, phenols (guaiacol and carvacrol), and homologues of carboxylic acids (C5-C10) and 2-alkanones (C8-C13). However, although the blends of carboxylic acids and that of 2-alkanones contributed incrementally to the repellency of the waterbuck odor to savannah tsetse flies, some waterbuck constituents (particularly, nonanoic acid and 2-nonanone) showed significant attractive properties. In another study, increasing the ring size of δ-octalactone from six to seven membered ring changed the activity of the resulting molecule (ε-nonalactone) on the savannah tsetse flies from repellency to attraction. In the present study, we first compared the effect of blending ε-nonalactone, nonanoic acid and 2-nonanone in 1:1 binary and 1:1:1 ternary combination on responses of Glossina pallidipes and Glossina morsitans morsitans tsetse flies in a two-choice wind tunnel. The compounds showed clear synergistic effects in the blends, with the ternary blend demonstrating higher attraction than the binary blends and individual compounds. Our follow up laboratory comparisons of tsetse fly responses to ternary combinations with different relative proportions of the three components showed that the blend in 1:3:2 proportion was most attractive relative to fermented cow urine (FCU) to both tsetse species. In our field experiments at Shimba Hills game reserve in Kenya, where G. pallidipes are dominant, the pattern of tsetse catches we obtained with different proportions of the three compounds were similar to those we observed in the laboratory. Interestingly, the three-component blend in 1:3:2 proportion when released at optimized rate of 13.71mg/h was 235% more attractive to G. pallidipes than a combination of POCA (3-n-Propylphenol, 1-Octen-3-ol, 4-Cresol, and Acetone) and fermented cattle urine (FCU). This constitutes a novel finding with potential for downstream deployment in bait technologies for more effective control of G. pallidipes, G. m. morsitans, and perhaps other savannah tsetse fly species, in ‘pull’ and ‘pull-push’ tactics., Author summary In our previous studies with tsetse fly-refractory waterbuck body odor, we found that certain subtle structural changes are associated with shifts in activities of some constituents from repellency to attraction. This led us to discovery of three potent tsetse attractants (ε-nonalactone, nonanoic acid and 2-nonanone). In the present study, we explored possible synergistic effects of blending of these compounds in different proportions to Glossina pallidipes and Glossina m. morsitans in the laboratory, followed by field studies with G. pallidipes. A three-component blend comprised of ε -nonalactone, nonanoic acid and 2-nonanone in 1:3:2 proportion gave 235% higher tsetse fly catches in the field compared with that of POCA and FCU. Thus, dispensing this odor blend in tsetse fly traps or insecticide treated targets is expected to suppress the tsetse flies more efficiently. It will also be interesting to see if the blend is similarly attractive to other savannah tsetse fly species.

Magnolia Flower is a crude drug used for the treatment of headaches, toothaches, and nasal congestion. Here, we focused on Magnolia kobus, one of the botanical origins of Magnolia Flower, and collected the flower parts at different growth stages to compare chemical compositions and investigate potential inhibitory activities against interleukin-2 (IL-2) production in murine splenic T cells. After determining the structures, we examined the inhibitory effects of the constituents of the bud, the medicinal part of the crude drug, against IL-2 production. We first extracted the flower parts of M. kobus from the bud to fallen bloom stages and analysed the chemical compositions to identify the constituents characteristic to the buds. We found that the inhibitory activity of the buds against IL-2 production was more potent than that of the blooms. We isolated two known compounds, tiliroside (1) and syringin (2), characteristic to the buds from the methanol (MeOH) extract of Magnolia Flower. Moreover, we examined the inhibitory activities of both compounds against IL-2 production and found that tiliroside (1) but not syringin (2), showed strong inhibitory activity against IL-2 production and inhibited its mRNA expression. Thus, our strategy to examine the relationship between chemical compositions and biological activities during plant maturation could not only contribute to the scientific evaluation of medicinal parts of crude drugs but also assist in identifying biologically active constituents that have not yet been reported.

Alkaloids are basic nitrogen-containing chemicals that have important physiological and pharmacological characteristics. Many vascular plant species contain alkaloids, and their roles in planta are of interest. However, the detailed distribution of alkaloids remains unclear because of their low water solubility and low concentrations in plants. In this study, we visualized the distribution of salicifoline, a water-soluble quaternary ammonium alkaloid, in the freeze-fixed stems of Magnolia kobus by cryo time-of-flight secondary ion mass spectrometry. Most of the salicifoline was distributed in living phloem tissues. In the xylem, salicifoline was detected in ray cells, lignifying wood fibres, and in vessels in the latest annual ring. The salicifoline distribution in the xylem varied with the cell wall formation stage. These results provide new insights into the storage, transportation, and role of the alkaloid salicifoline in M. kobus.

In West Africa, whether in forests or savannahs, most of the large mammal species have a scattered spatial distribution. Monitoring their population size represents a logistic, financial, theoretical challenge because counting of transects is not appropriate, unlike elsewhere in Africa. In this study, we (i) analyse the patterns of interannual (2006–2008) changes in population density of seven sympatric species of savannah herbivores in a protected area of Burkina Faso with two alternative methods: dung counts versus direct counts and (ii) quantify the economic costs for these two methods. The seven species of mammals we considered are the following: Hippotragus equinus, Alcelaphus busephalus, Sylvicapra grimmia, Tragelaphus scriptus, Ourebia ourebi, Kobus ellipsiprymnus and Phacochoerus africanus. A distance methodology was used to generate estimates with both methods. There were significant correlations between estimates of mammal density obtained using dung counts and direct counts, and an analysis of covariance revealed that the interannual trends in population densities were consistently detected with both methods. In addition, the dung count method costs 55% less. Thus, our study documents that dung counts can be used as a proxy of population size fluctuations for the seven studied species, offering a methodological alternative that is much less expensive, less sophisticated in terms of equipment compared to other methods and that can be performed by field staff with moderate professional qualification. We suggest that standardized campaigns of dung count surveys may be applied to all protected areas and savannah in Burkina Faso, West Africa, in order to improve evidence-based, large-scale conservation and management planning in the region.

How do large-mammal communities reassemble after being pushed to the brink of extinction? Few data are available to answer this question, as it is rarely possible to document both the decline and recovery of wildlife populations. Here we present the first in-depth quantitative account of war-induced collapse and postwar recovery in a diverse assemblage of large herbivores. In Mozambique’s Gorongosa National Park, we assembled data from 15 aerial wildlife counts conducted before (1968–1972) and after (1994–2018) the Mozambican Civil War (1977–1992). Pre-war total biomass density exceeded 9,000 kg km-2, but populations declined by >90% during the war. Since 1994, total biomass has substantially recovered, but species composition has shifted dramatically. Formerly dominant large herbivores—including elephant (Loxodonta africana), hippo (Hippopotamus amphibius), buffalo (Syncerus caffer), zebra (Equus quagga), and wildebeest (Connochaetes taurinus)—are now outnumbered by waterbuck (Kobus ellipsiprymnus) and other small to mid-sized antelopes. Waterbuck abundance has increased by an order of magnitude, with >55,000 individuals accounting for >74% of large-herbivore biomass in 2018. By contrast, elephant, hippo, and buffalo, which totaled 89% of pre-war biomass, now comprise just 23%. These trends mostly reflect natural population growth following the resumption of protection under the Gorongosa Restoration Project; reintroductions (465 animals of 7 species) accounted for a comparatively small fraction of the total numerical increase. Waterbuck are growing logistically, apparently as-yet unchecked by interspecific competition or predation (apex-carnivore abundance has been low throughout the post-war interval), suggesting a community still in flux. Most other herbivore populations have increased post-war, albeit at differing rates. Armed conflict remains a poorly understood driver of ecological change; our results demonstrate the potential for rapid post-war recovery of large-herbivore biomass, given sound protected-area management, but also suggest that restoration of community structure takes longer and may require active intervention.

The article reflects the results of the study of history of introduction and acclimatization of species of Magnoliaceaе Juss. family in Volyn and Podillia. The research has identified the individual centres of introduction and sources of seed and planting material. The taxonomic composition of Magnoliaceae family in 18 dendrological facilities and green plantations in Volyn Podillia is described. These objects are as follows: Kamenetz-Podilsk and Kremenets botanical gardens, Lviv National University botanical gardens, Khorostkiv and Hermakivka, Bylokrinitsa dendrological park, Bilche-Zolote, Rayiv and Rivne parks, arboreta of Shatsky forestry college and Kozova school arboretum, rosary of Taras Shevchenko Kremenets Regional Humanitarian-Pedagogical Institute, Lutsk, Kamenetz-Podilsk, Ternopil, Rivne, and Berezhany. Collections of botanical gardens, arboreta, parks, seed banks are a vital guarantee of the preservation of species of magnolias. They are used to study the relicts, their distribution in ornamental horticulture and for repatriation in nature. In Volyn and Podillia the following species grow: Liriodendron tulipifera L., M. kobus DC., M. x loebneri Kache., M. acuminata L., M. officinalis Rehd.et Wils., M. tripetala L., M. salicifolia (Sieb.et Zucc.) Maxim., M. x soulangiana Soul.- Bod., 'Rosea', 'Lennei Alba', 'Alexandrina', M. sieboldii K. Koch., M. virginiana L., M. hypoleuca (Sieb.et Zucc.), M. stellata (Sieb.et Zucc.) Maxim., M. obovata Thumb., M. x kewensis Pearce, M. wilsonii (Finet et Gagnep.) Rehder, M. ashei Weatherby, M. liliflora Desr. The Red List of Plants (IUCN Red List Categories and Criteria) encouraged scientists and supporters of nature conservation to focus on gaining new knowledge on the strategy to protect these species. The research has determined that the greatest number of species is concentrated in the collections of the Botanical Garden of the Ivan Franko Lviv University, Kremenets Botanical Gardens and Khorostkovsky Dendropark. Our scientific research will be used to establish taxonomic composition and research representatives of Magnoliaceae family in all dendrological objects of Volyn Podillia.

A 19-year-old captive male waterbuck (Kobus ellipsiprymnus) exhibited traumatic pericarditis at necropsy. The animal weighed 182 kg at necropsy and revealed no remarkable findings in external observation. Severe pericardial adhesions with fibrosis, hepato-diaphragmatic adhesions, straw-colored ascites and hepatosplenomegaly were observed upon examining the internal organs. Perforations made by a 12-cm-long sharp-ended bamboo twig were detected in the reticulum, diaphragm, pericardium, lung and liver. Trueperella pyogenes was identified in pericardial fluid. To our knowledge, this is the first documented case of traumatic reticulopericarditis caused by a sharp-ended bamboo twig in a captive waterbuck.

The impact of abiotic and anthropogenic factors on the distributions of buffalo ( Syncerus caffer ), harte­beest ( Alcelaphus buselaphus ), roan antelope ( Hippotragus equinus ) and waterbuck ( Kobus defassa ) at Mole National Park was assessed by transect survey. Generalized linear mixed effects logistic re­gression was used to model mammal presence/absence as a function of ecological factors. Hartebeest inhabited highlands and avoided floodplains but buffalo and roan avoided floodplains by selecting both lowlands and highlands while waterbuck inhabited lowlands but not necessarily the floodplains. Fire, water availability and anthropogenic activities were limiting factors, which constrained habitat use to make some areas unexplored for foraging. Buffalo, roan and hartebeest did not inhabit areas close to the park’s boundaries. Herbivores need optimal environment almost free of constraints to construct their distribution patterns. Therefore, management should address the problems identified in this study to ensure the herbivores’ redistribution to maximise their use of resources for their effective conservation.

Kobus Moolman prefaces A Book of Rooms with a quote by Georges Perec. The quote details the irrevocability of the past through memory. However, both Perec and Moolman not only recover memory, but are able to do so in great detail, specifically through thorough catalogic descriptions of spaces and objects that surround them in the domestic realm. Analysis of these catalogic descriptions forms the key component of this article. The structure of Moolman’s work, with different rooms used to demarcate different sections, and the significance of objects, further contribute towards his project of recovering the irrevocable. Comparison of Moolman’s project with that of Perec, with reference to Bachelard’s thoughts on the home, serves in the analysis of how the self is related to the concept of a house, with its many rooms in which are stored those things which contribute to an individual’s sense of identity. The generation of narrative via description and cataloguing of these various domestic objects and events is considered, with specific focus given to the ‘bed’ as it plays a significant role in the formation of the self and the recollection of memories.Keywords: Bachelard, catalogic description, domestic sphere, Kobus Moolman, Georges Perec

The well-being of a species fundamentally rests on understanding its biology, home range, and distribution. The highly seasonal distribution of kobs poses conservation and management difficulties, particularly because of the capricious nature of the ever-changing ecological and vegetation dynamics of the ecosystem. Assessing the distribution of kobs and their associated vegetation provides insight into the vulnerability and conservation status of the species. Species distribution and habitat suitability maps were developed and created respectively for the management of kobs in the Shai Hills Resource Reserve. Kob presence data collected was analyzed using the spatial analyst and Hawth's tool in the ArcGIS software where the gradients of kob distribution within the protected area landscape were plotted and mapped. Seven environmental variables including location, land cover/use, slope/elevation, nearness to dams and rivers, temperature, and rainfall were considered to have effect on kob distribution pattern and as such used in the development of species distribution and habitat suitability maps. The results indicated that kobs in the Shai Hills Resource Reserve (SHRR) assume a clumped or contagious distribution pattern where individual kobs are aggregated in patches. Rainfall, temperature, nearness to dams and rivers, slope/elevation, and land cover/use had influence in kob distribution. Of all the cataloged habitats, 86, 13, and 1% were moderately suitable, suitable, and unsuitable, respectively. Long-term survival of species depends on adequately large areas of suitable habitats and opportunities for home range activities between such areas. As such, it is recommended that suitable habitats for kobs be dedicated and designated as conservation areas, especially areas along the western boundary.

Setyawan AD, Sugiyarto. 2015. Diversity of Selaginella in the Bromo Tengger Semeru National Park, East Java. Pros Sem Nas Masy Biodiv Indon 1: 1312-1317. Eastern Java has a relatively drier climate than other parts of Java, but it still has quite a lot of cool mountain areas and provide enough water, for example, Bromo Tengger and Semeru Mountains complex. Selaginella is a herbaceous plant that need water for reproduction, thus it is important to know its existence in the dry climates region due to global climate change. This aims of this study was to determine the species diversity of Selaginella in the National Park of Bromo Tengger and Semeru (BTSNP), East Java. Field study was conducted in August 2007 and May 2015, and followed by observation of specimen collection of Herbarium Bogoriense, Research Center for Biology, Indonesian Institute of Sciences Cibinong, Bogor. From the field research, there were collected five herbarium specimens, while the herbaria collections observed were 49 sheets of specimens. The results showed that in this park and its border areas, there were found eight species of Selaginella, namely: S. ciliaris, S. intermedia, S. involvens, S. opaca, S. ornata, S. plana, S. remotifolia and S. singalanensis. However, the morphological characteristic of S. intermedia and S. singalanensis needs to be further confirmed since each of them was only observed from a single sheet herbarium, namely O Posthumus 1615 and Kobus Tosari 147, respectively. In Java, S. intermedia was mainly distributed in the western part of Java. Meanwhile, the presence of S. singalanensis in Java was relatively rare.

This article explores the possibility of an alignment of (a) theology (understood as study concentrating on one faith from the faith suppositions of that faith) and (b) a study of world religions, open to all regardless of faith in the context of an encompassing theory. This article argues that far from undermining the dignity and value of any religion, (b) could add value to the theological study of any one religion. This article proceeds to develop a model for (b). Utilising a methodology revolving around the strategies of careful historical criticism, faithful intentional understanding and imaginative yet trustworthy tendentional interpretation, this article explores a model harmonising the key concepts of metaphysical mysticism (understood as the peak of religious aspiration), conditionality (emphasising the unique singularity of individual religions), totality (emphasising the religious landscape as one coherent whole) and horizon (emphasising the non-absoluteness of all religions, petering out on, and arising from, silence).

The thesis proposes a study of late medieval plucked string musical instruments of the family of guitar and lute, based on a multidisciplinary approach that mixes historical, literary, linguistic-etymological, philological and iconographic aspects. The time span primarily considered ranges from the mid-1200s to the late 1400s, that is, from the first written evidence of these instruments in European languages to the invention of the tablature and the first treatises on organology. The sources used comprise Arab and Latin musical treatises, literary and historical documents in European languages, as well as iconographic evidence. The main purpose of the thesis is a comprehensive investigation of the history of medieval plucked string instruments, not limiting the scope of research to linguistic and organological problems, but also extending it to the performative aspects, the repertoire, the musicians and the literary and cultural context in which these instruments appeared. Another objective is to identify all the evidence connected to the instruments that form the object of this research, in order to perform an exhaustive analysis of the terminology and highlight a series of less known sources.

Dhaka Zoo with 2000 animal heads of 184 species from significant genetic diversity and five million visitors’ influx round the year is placing it a public health important spot. This study was conducted to investigate aspergillosis in animals at Dhaka Zoo to ascertain animal health, welfare and public health safety standard. One hundred and two necropsied tissue samples preserved in 10% neutral buffered formalin at necropsy from 36 animals of 25 different species were collected from Dhaka Zoo. Twenty five out of 36 study animals were suffering from granulomatous diseases. Among them 13 animals were suffering from Aspergillosis. Clinical history, nodular lesions from necropsy findings, granulomatous reactions along with fungal spores and characteristic radiating club on histopathology; dichotomously branching septate hyphae and mycelial conidiophore on special staining were revealed Aspergillosis in 13 animals of nine species that includes four rhesus macaques ( Macaca mulatta ), two samber deer ( Cervus unicolor ) and one of each species were nilgai ( Boselaphus tragocamelus ), horse ( Equus caballus ), stripped hyena ( Hyena hyena ), gayal ( Bos frontalis ), beisa oryx ( Oryx beisa beisa ), water buck ( Kobus L. leche ) and greater kudu ( Tragelaphus strepsiceros ). Present study provides evidence of existing Aspergillosis and similar long standing zoonotic diseases in majority of rest of the animals with health risk that shades health safety standard at Dhaka Zoo. DOI: http://dx.doi.org/10.3329/jbau.v11i2.19924 J. Bangladesh Agril. Univ. 11(2): 265-270, 2013

Multiple land uses including tourism, hunting, and agriculture around protected areas can be a serious complication for wildlife management. We calculated habitat selection indices (Manly's alpha) for 10 bovid species in the Pendjari Biosphere Reserve in Benin, west Africa, to assess if habitat use differed in each bovid species between hunting and non-hunting zones. Presence/absence data was used in resource-selection functions based on a generalized linear mixed effect model to examine factors that explained bovid species distribution. We observed stronger avoidance of open habitat types in the hunting zone than in the non hunting zone for the hartebeest Alcelaphus buselaphus, oribi Ourebia ourebi, roan Hippotragus equines, kob Kobus kob, Waterbuck Kobus ellipsiprymnus defassa and reedbuck Redunca redunca. In contrast, in grey duiker Sylvicapra grimmia, red-flanked duiker Cephalophus rufilatus, bushbuck Tragelaphus scriptus and buffalo Syncerus caffer we found no differences in habitat use between hunte...

As an endangered species, Magnolia kobus is distributed in Jeju island in Korea with only about 500–1000 individuals. In this study, we presented a complete chloroplast genome of M. kobus which is 159,443 bp and has four sub-regions: 87,484 bp of large single copy and 18,783 bp of small single copy regions are separated by 26,588 bp of inverted repeat regions including 113 genes (79 unique genes, four rRNAs and 30 tRNAs). Phylogenetic analysis using chloroplast genomes showed that M. kobus is a sister of M. insignis and M. laevifolia clade.

This paper presents the phenology of seven taxa of the genus Magnolia L., pollen biology on germination and storage conditions, seed productivity, germinating ability of seeds and its dependence on stratification and germination conditions. It has been found that M. kobus DC, M. kobus var. boreales Sarg., M. obovata Thurb., M. officinalis Rehd. et Wils., M. salicifolia (Sieb. et Zucc.) Maxim., M. sieboldii K. Koch. and M. tripetala L. in cultivated condition produce pollen at a low viability rate (9.4 - 31.7%). Real seed productivity of the taxa being studied is less than their potential productivity. However, M. obovata , M. officinalis and M. tripetala are characterized by high seed germinating ability, up to 94%. Optimal germination conditions for seeds of M. tripetala , M. officinalis and M. obovata require protected ground (greenhouse) and stratification at 4oC during 30 days. The high germinating ability of M. sieboldii seeds in the open ground is explained by stratification duration and temperature required for this species. Sarcotesta effects on higher seed germinating ability have been observed in M. obovata only. DOI: http://dx.doi.org/10.3329/bjpt.v20i2.17390 Bangladesh J. Plant Taxon. 20(2): 163-170, 2013

A South African endemic antelope, the Grey Rhebok (Pelea capreolus), has long been an evolutionary enigma in bovid systematics—its phylogenetic intractability attributed to its curious combination of derived and primitive morphological attributes and the consequences of a rapid radiation. By using a combination of DNA sequences, chromosomal characteristics and quantitative and qualitative morphological features we show that the species is a sister taxon to a clade that comprises the waterbuck, reedbuck and allies. Our finding of few unambiguous synapomorphies reinforces suggestions of a rapid radiation and highlights the effects of incomplete lineage sorting, including the hemiplasic nature of several chromosomal rearrangements. We investigate these data to address the general question of what may have led to Pelea being both genetically and ecologically distinct from the Reduncini. We argue that its adaptation to exposed habitats, free of standing water, arose by vicariance prompted by increasing aridity of the extreme south/southwestern region of the African continent in the Miocene. Ancestral lineages leading to the extant Redunca and Kobus, on the other hand, retreated to water-abundant refugia in the north during these mostly globally cool phases. The mosaic of water-rich environments provided by the Okavango and the drainage systems in the southwestern extension of the East African Rift system are considered to have facilitated speciation and chromosomal evolution within these antelope.

The aim of this study was to investigate the potential use of leaf-level hyperspectral data to predict the density of sugarcane thrips Fulmekiola serrata (Kobus). A hand-held spectroradiometer was used to make the spectral measurements on spindle leaves of 4- to 5-month-old plants of sugarcane cv. N19 growing in commercial fields near Umfolozi, South Africa. A random forest algorithm followed by partial least squares regression was used for the analysis. Developed models were adequate to predict nymph numbers in December and adult numbers in March, but different models were needed for the thrips life stage assessed and the season when the estimation took place.

This article suggests a theoretical and methodological perspective primarily hinging on the categories of Horizon, Totality and conditionalism, with an outspoken mystical orientation, radically relativising yet simultaneously treasuring diverse religious expression. This model was developed with a view to interpreting the history of religions, in this case applied to the history of the Faculty of Theology at the University of Pretoria from 1917 to 2017.Utilising this perspective, the history of the faculty is analysed in terms of three qualitatively distinct yet continuous epochs, overlapping with the three epochs of South African history during the twentieth century: 1902–1948, 1948–1994 and 1994 to present. In particular the article focuses on two dimensions of theological existence at the University of Pretoria: firstly, its interaction with the state over this century, that is, its political existence during the decades prefiguring apartheid, during apartheid and during the aftermath of apartheid; secondly, its relationship with the wider world of religious pluralism over the past century, implying its notion of religious truth. Differences of emphasis and conflicts during the century, involving both sets of problems, are explained and understood conditionalistically and with reference to Totality and Horizon. Racial exclusion and religious exclusion are understood as mutually determining and are both informed by and dependent on a certain view of religious truth.In the context of its own ambit this article has a reconciliatory intention, not evaluating the mistakes of the past in terms of the categories of sin and guilt, but rather in terms of tragic misjudgements of situations: shortcomings in historical hindsight, sufficiently wide peripheral vision, realistic foresight and sufficient insight into the epochal conditions of the times and the essence of religion. Greed and hatred, seemingly ingrained in human nature, are taken to feed on such lack of insight.

The present study was conducted on six Magnolia species in the period 2006-2008 in the Botanical Garden of the Maria Curie-Skłodowska University in Lublin. The soil and leaves from the locations of the following species of Magnolia were subjected to evaluation: cucumber tree (Magnolia acuminata L.), Kobushi magnolia (Magnolia kobus DC), purple lily magnolia (Magnolia liliiflora L.), Japanese willow-leaf magnolia (Magnolia salicifolia Siebold et Zucc. Maxim.), saucer magnolia (Magnolia x soulangiana Soul.-Bod.), and umbrella magnolia (Magnolia tripetala L.). The studied plants were planted during the period 1968-1989 on grey-brown podzolic soil derived from loess. The significantly lowest content of phosphorus, potassium and magnesium as well as the lowest pH were recorded in the soil from the location occupied by the purple lily magnolia (M. liliiflora). The other sites were characterized by an optimal range of pH, high soil phosphorus and magnesium availability as well as medium potassium availability. An optimal content of nitrogen and calcium was found in the leaves of the cucumber tree (M. acuminata), of phosphorus and magnesium in the leaves of the Kobushi magnolia (M. kobus), and of potassium in the leaves of the Japanese willow-leaf magnolia (M. salicifolia). A low content of nitrogen and calcium was shown in the leaves of the purple lily magnolia (M. liliiflora), of phosphorus in the leaves of the saucer magnolia (M. x soulangiana), and of potassium in the indicator parts of the cucumber tree (M. acuminata). The study found that among the studied species the cucumber tree (M. acuminata) could be a good biostabilizer of nitrogen, while the Kobushi magnolia (M. kobus) a good biostabilizer of phosphorus and magnesium. In spite of significant differences in soil nutrient availability and clear variations in macronutrient contents in the indicator parts of the investigated plants, no visual symptoms of nutrient deficit were found in the Magnolia species under study.

Field immobilization of captive antelope may be required for medical examination, blood sample collection, and animal identification. The aim of this study was to evaluate the effects of a combination of butorphanol, detomidine, and midazolam (BDM) and its partial reversibility in Nile lechwe antelope (Kobus megaceros). Nine captive lechwes, weighing 28-64 kg, were immobilized, in February 2011, with butorphanol 0.20 ± 0.05 (mean ± SD) mg/kg, detomidine 0.20 ± 0.05 mg/kg, and midazolam 0.31 ± 0.08 mg/kg administered intramuscularly (IM) with a blowpipe. Physiologic parameters and depth of anesthesia were recorded when the animals became recumbent at 19.55 ± 8.36 min after darting (T0) and after 10 (T10), 20 (T20), and 30 (T30) min. An arterial blood sample was collected at T20. At the end of the procedures, immobilization was partially reversed with atipamezole 0.25 mg/kg IM. Quality of induction, immobilization, and recovery was scored. The BDM combination induced immobilization and lateral recumbency in 13.44 ± 5.61 min. Median induction score (scored 1 [excellent] to 4 [poor]) was 1 (range 1-2). Heart rate varied 40-104 beats/min, respiratory rate 16-108 breaths/min, and rectal temperature 36.5-40.3 C. Hyperthermia was observed and rapidly treated in three animals that demonstrated insufficient immobilization after darting. Arterial blood gas analyses revealed a mean pH of 7.43 ± 0.07, partial arterial pressure of CO(2) of 44.1 ± 6.0 mmHg, partial arterial pressure of O(2) of 74.0 ± 13.5 mmHg, and an arterial O(2) saturation of 94.77 ± 3.96%. Recovery was smooth and animals were walking in 13.44 ± 7.85 min. Median recovery score (1 = excellent to 4 = poor) was 1 (range 1-2). The BDM was effective in immobilizing captive healthy lechwes with minimal cardiorespiratory changes.

Always regarded as somewhat of an ‘outsider’ (the child of an Englishspeaking (Catholic) mother and an Afrikaans (Protestant) father in an exclusively Afrikaans milieu) and growing up extremely poor, seeing the hardship of others and realising how much talent was going to waste, Kobus Maree took a particular interest in gifted disadvantaged persons. A marginalised loner, he almost inevitably developed creative abilities and took a keen interest in creativity, giftedness and the education of gifted, disadvantaged learners. As an adult, his research showed that many teachers in South Africa have to contend with the generally poor socioeconomic background of learners. A dire need for appropriate teacher and learner support materials, and school environments that are not conducive to achievement (including inadequate facilities, overcrowded classrooms, lack of teacher and learner support materials). South Africa is at a critical stage in its education. It is therefore important for educators to teach emotional intelligence in their classrooms. Our biggest challenge will be to maintain and enhance vitality in gifted education in a dynamic, ever-evolving environment. A combination of scholarly leadership and strategic management to support gifted learners is important. We should do all we can to promote societal transformation and diversity, focussing anew on underrepresented groups (women and ethnic groups) who show promise and support them. The widest array of partners possible including the big institutional players, the entire teaching fraternity (including government departments), nongovernmental organisations and miscellaneous interest groups together should develop strategic, rolling five-year plans and make gifted education a priority.

Six species of Magnolia L., growing in the Botanical Garden of the Maria Curie-Skłodowska University in Lublin which is located on the north-western outskirts of the city of Lublin, were studied during the period 2006-2008. The aim of this study was to determine the dynamics of leaf and flower development in these Magnolia species under the climatic and soil conditions of Lublin as well as to evaluate the size of their leaves and long shoots. The present study included the following species: M. acuminata L., M. kobus DC, M. liliiflora Desr., M. salicifolia Siebold et Zuch, M. x soulangiana Soul.- -Bod., and M. tripetala L. The magnolia trees were planted in 1968-1989 on grey-brown podzolic soil derived from loess. The study area is affected by the continental climate which is characterized by large annual amplitude of temperatures, long summers as well as long and cold winters. The long-term average annual air temperature for this region is 8.1oC, while the long-term average annual rainfall reaches 541.6 mm. The obtained results showed that, among the Magnolia species studied, M. salicifolia flowered earliest under the climatic and soil conditions of Lublin, while M. tripetala was the last to bloom. Full leaf development, on average for the study period, was recorded earliest in M. salicifolia and M. liliiflora, whereas it occurred latest in M. tripetala. The leaves of M. tripetala were characterized by the greatest length and width, while the smallest leaves were recorded in M. kobus. M. liliiflora was found to show the smallest annual shoot increments and the lowest number of leaves.

Thirty-eight taxa of Zygomycetes distributed in 15 genera were recorded from tapir (Tapirus terrestris), camel (Camelus bactrianus), horse (Equus caballus), deer (Cervus elaphus), agouti (Dasyprocta aguti), donkey (Equus asinus), llama (Llama glama) and waterbuck (Kobus ellipsiprymnus) dung collected at the Reserva Ecológica de Dois Irmãos located in Recife, State of Pernambuco, Northeast Brazil. The samples were collected on a monthly basis from June 2005 to May 2006, taken to the laboratory and incubated in moist chambers. Higher number of taxa was observed in the excrements of tapir, followed by deer and donkey. The highest number of species was detected for Mucor, followed by Pilobolus. Statistical analyses showed significant differences in richness of Zygomycetes taxa between the herbivore dung types. Differences of species composition, however, were weak. Seasonality influenced the Zygomycetes species composition but not its richness. Variations in taxa composition between ruminants and non-ruminants dung were non significant.

International audience; In group-forming prey species, theory assumes that individuals within groups should scan independently of one another, with vigilance sequences being relatively unpredictable, making interscan durations highly variable. We attempted to detect any divergence from randomness in the scanning process in three mammalian prey species phylogenetically and geographically separated and exposed to different levels of predation: waterbuck, Kobus ellipsiprymnus defassa, under a high observed predation risk, eastern grey kangaroo, Macropus giganteus, still experiencing occasional predation and European roe deer, Capreolus capreolus, under a very low natural predation risk. Our results revealed that the focal interscan duration increased when the duration of the preceding interscan increased, whatever the studied species and the predation risk that its individuals experienced, and decreased with the preceding scan duration in two species under, respectively, occasional and low predation risks. The exponential distribution was the tested model that fitted the observed distributions of interscan durations least well. We discuss what can trigger non-randomness in scanning, through a non-homogenous Poisson process, at both intra-individual and inter-individual levels, particularly with regard to previous studies that have demonstrated synchronisation of vigilance in such mammals. Our results suggest the need to reconsider any assumption of randomness in scanning in the basic model predicting form and frequency of scanning behaviour by prey species.

Reproductive effort should negatively correlate with reproductive value, yielding a pattern of increased effort with age. According to the terminal investment hypothesis, females near the end of their reproductive life span should devote more resources to reproduction than those near the start of their reproductive careers. We tested predictions of the terminal investment hypothesis by evaluating 38 years of reproductive life-history data collected from Nile lechwe (Kobus megaceros), an ungulate species living at San Diego Zoo's Wild Animal Park. The maximum reproductive success of Nile lechwe matched predictions of models of lifetime reproductive effort, with the relative mass of newborn calves providing an accurate indicator of the costs of reproduction. Newborn mass was significantly correlated with maternal age, and neonatal males tended to be heavier than neonatal females. Older dams were more likely to produce sons than daughters, dams that produced sons were more likely to die than were da...

This study investigated composition and selectivity in diet for waterbuck in the Pendjari National Park in north-western Benin, through the use of micrographic analysis of faecal samples. Three plant species (Panicum anabaptistum, Echinochloa stagnina and Andropogon gayanus) were regularly consumed all year round. Meanwhile, three other species (i.e., Hyparrhenia involucrata, Acroceras amplectens and Oryza barthii) are mostly found in its diet during the beginning of the rainy season. During the dry season, long life grasses (>40%) and tree forage (about 35%) were the most dominant life form in the diet. On the contrary at the beginning of the rainy season, annual species (> 50%) were dominant. In conclusion, the waterbuck has a grazer regime when plant species are abundant and a mixed diet during the dry season. Waterbuck’s food niche breath, defined by Hespenheide [Ecology and Evolution of communities. Harvard Univ. Press, 1975], was lower than 1, implying this antelope does not eat all food categories in a proportional way. Shannon diversity index showed that the diet was more diversified during the rainy season and less diversified at the end of the dry season. Based on [Ecology, 64 (1983), 1297] diet selectivity index, waterbuck exerted a positive selection on the major graminaceous species. Resume Une etude a ete realisee sur le regime et la selectivite alimentaire du waterbuck dans le Parc National de la Pendjari au Nord-Ouest Benin a partir d’une analyse micrographique des echantillons de crottes. Trois especes vegetales (Panicum anabaptistum, Echinochloa stagnina et Andropogon gayanus) sont particulierement consommees en toutes saisons. La consommation d’especes annuelles comme Hyparrhenia involucrata, Acroceras amplectens et Oryza barthii, est observee en saison humide. Durant la saison seche, l’alimentation du waterbuck est un regime mixte de graminees vivaces (> 40%) et de fourrage ligneux (environ 35%), alors que pendant la saison des pluies ce sont les herbacees annuelle (> 50%) qui dominent. Le waterbuck est donc un animal paisseur en periode d’abondance alimentaire. La largeur de la niche alimentaire du waterbuck est inferieure a l’unite. Donc, le waterbuck ne consomme pas toutes ces categories alimentaires dans les memes proportions. L’indice de diversite de Shannon indique que le regime alimentaire du waterbuck est plus diversifie en saison humide qu’en fin de saison seche. Les valeurs de l’indice de selectivite alimentaire de Chesson (1983), indiquent que le waterbuck exerce une selection positive sur la plupart des graminees consommees.

The telematic debate on the theme “Global History and Global Policies” has provided historians from different historiographic traditions and distant countries – Piero Bevilacqua, Guillermo Castro, Ranjan Chakrabarti, Gabriella Corona, Kobus du Pisani, John McNeill, Donald Worster – with an opportunity to compare views on the different interpretive paradigms of global environmental history. A wide range of subjects was covered: the definition of the discipline’s field of research, themes, and chronological scope; the relationship between global and local; the role of the West in history and historiography; the perspective of the dominated; the discipline’s role in policy making; and its relationship with the natural sciences. The discussion has yielded a rich harvest of reflections on global environmental history as a paradigm for the interpretation of the past and a cultural and political instrument for action in the present. Important indications have emerged as regards the research paths a global environmental historian can follow to contribute to this field of studies.

The small size of many African protected areas makes adjacent rangelands potentially important in the local survival of wild animals. In order to assess the importance of pastoral areas to wild ungulates, we studied density and habitat choice of wild ungulates and cattle in Lake Mburo National Park, Uganda, the adjacent exclusively pastoral Nshara Dairy Ranch and on private land consisting of a mixture of ranching and subsistence farms. Transects, in the three land-use zones, were walked during the wet season and the data were analysed by DISTANCE sampling technique. We found significantly higher total density of wild ungulates on the dairy ranch compared with the National Park and private land. There was no significant difference in total wild animal density between the National Park and private land. Impala (Aepyceros melampus), zebra (Equus quagga), bushbuck (Tragelaphus scriptus) and waterbuck (Kobus ellipsiprymnus) had significantly higher densities on the dairy ranch compared to the National Park. Only eland (Taurotragus oryx) density was higher in the National Park compared to private land. Wild ungulates and cattle showed a high degree of habitat overlap, generally preferring open grassland. Our study shows that high densities of wild ungulates are not necessarily associated with protected areas. Pastoral areas may be important for populations of wild herbivores during the growing season despite a pronounced presence of livestock.

Resource manipulation, such as the creation and maintenance of grazing lawns, may shape the structure of herbivore communities. We tested the hypothesis that grazing lawns contribute towards the subsistence of the Kobus kob kob in a dystrophic West African savanna, where kob and Hippopotamus amphibius both occur. Comparison of the foliage of grazing lawns and ungrazed swards shows that hippo lawns are more nutritious with regard to both structure and nutrients; kob lawns are higher in nutrients only. Up to the early dry season, hippo lawns meet kob energy and protein demand, thereafter, the shortness of the sward limits intake. Kob lawns always provide sub-maintenance values. Grazing on ungrazed swards is least profitable. We suggest that grazing lawns are essential for the daily subsistence of mesoherbivores, particularly on nutrient-poor soil, and that megaherbivores facilitate their food supply, for at least part of the year.

The kob (Kobus kob kob) population studied in the Comoe National Park, Ivory Coast, suffered unsustainable losses to poaching before and during the investigation. This in turn did not only lead to low densities but also changed the population composition, resulting in a strong female bias in all age classes except juveniles. The relative rarity of adult males probably had negative effects on the population recruitment. Although mating occurred throughout the year and birth intervals were short, at least in some females, annual birth-rate was low compared to earlier studies in the same area. Effective conservation measures are urgently needed to ensure the future existence of the studied population. Resume La population de kobes (Kobus kob kob) etudiee au Parc National de Comoe, en Cote d'Ivoire, a souffert de pertes insoutenables dues au braconnage, avant et pendant les investigations. Ceci a entraine non seulement de faibles densites mais aussi des changements de la composition de la population, avec pour resultat un biais important en faveur des femelles dans toutes les classes d'âge sauf les juveniles. Bien que les accouplements aient lieu toute l'annee et que les intervalles entre les naissances soient courts, en tout cas pour certaines femelles, le taux de natalite annuel etait faible compare aux etudes realisees precedemment dans la region. Il faut absolument prendre des mesures de conservation urgentes et efficaces pour garantir l'avenir de la population etudiee.