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Drusilla profunda nov.sp. (Figs 45 and 183) T y p e m a t e r i a l Holotype, Sabah, Danum Valley, B.R.L., f.i.t., 14-16.II.2007, G. de Rougemont leg. (CROU). D e s c r i p t i o n: Length 3.93 mm. Body shiny, reddish, elytra reddish-brown, abdomen yellowish-red, antennae reddish-brown with the two basal antennomeres, the base of third and the eleventh yellowish-red, legs yellowish-red. Second antennomere shorter than the first, third longer than the second, fourth as long as wide, fifth to tenth transverse. Eyes very large in dorsal view. Fore-body devoid of reticulation, that of the abdomen very transverse and evident. Puncturation of the head fine and very superficial, that of the abdomen absent. Granulation of the pronotum and elytra fine, very superficial and not very salient. There is a salient tubercle between the antennae. Median sulcus of the pronotum shallow and ending in a deep posterior dimple. C o m p a r a t i v e n o t e s: In the form of the spermatheca, the new species is similar to D. semimonticola PACE, 2008 also from Borneo. It differs in the bicoloured abdomen, whereas in D. semimonticola the abdomen is uniformly yellowish-red, in the deep apical umbilicus of the distal bulb of the spermatheca, which is short in D. semimonticola, and in the proximal bulb of the spermatheca which is narrower (0.013 mm), than the maximum width (0.052 mm) of the distal bulb, whereas in D. semimonticola the proximal bulb of the spermatheca is as wide (0.066) as the maximum width of the distal bulb. E t y m o l o g y: The new species is named "deep" in reference to the deep apical umbilicus of the distal bulb of the spermatheca., Published as part of Pace, R., 2014, Aleocharinae from Sabah (Borneo) collected by Guillaume de Rougemont (Coleoptera, Staphylinidae), pp. 727-794 in Linzer biologische Beiträge 46 (1) on page 757, DOI: 10.5281/zenodo.5306753

Pelioptera plenitudinis nov.sp. (Figs 13 and 93) T y p e m a t e r i a l Holotype, Sabah, Danum Valley, B.R.L., f.i.t., 14-16.II.2007, G. de Rougemont leg. (CROU). D e s c r i p t i o n: Length 2.12 mm. Body shiny, yellowish-brown, head and elytra brown, antennae brown with the two basal antennomeres yellowish-brown, legs yellow. Second antennomere shorter than the first, third shorter than the second, fourth to tenth transverse. Eyes longer than the postocular region in dorsal view. Fore-body devoid of reticulation, that of the abdomen irregular, polygonal, evident. Puncturation of the head very superficial and absent on frons. Granulation of pronotum and elytra very superficial and evanescent, that of the abdomen fine and evident. The pronotum bears four punctures disposed in a square, but the puncture of the right anterior angle is missing. No secondary sexual characters on the female fifth free abdominal tergite. Spermatheca: Fig. 93. C o m p a r a t i v e n o t e s: The spermatheca of the new species is similar in shape to that of P. bituberculata PACE, 2003 from West Malaysia but the apical umbilicus of the distal bulb of the spermatheca is enormous in the new species, and absent in of P. bituberculata. Moreover the length of the spermatheca of the new species is 0.2 mm, while in P. bituberculata it is 0.27 mm. E t y m o l o g y: The name of the new species means "of the fullness" in reference to the distal bulb of the spermatheca with its enormous apical umbilicus that fills the entire internal space., Published as part of Pace, R., 2014, Aleocharinae from Sabah (Borneo) collected by Guillaume de Rougemont (Coleoptera, Staphylinidae), pp. 727-794 in Linzer biologische Beiträge 46 (1) on page 738, DOI: 10.5281/zenodo.5306753

Chaetosogonocephus ruficollis nov.sp. (Figs 20 and 108) T y p e m a t e r i a l Holotype, Sabah, Danum Valley, B.R.L., f.i.t., 14-16.II.2007, G. de Rougemont leg. (CROU). D e s c r i p t i o n: Length 3.63 mm. Body shiny, reddish, head and pronotum yellowish-red, antennae brown with the two basal antennomeres and the base of third yellowish-red, legs reddish. Second antennomere shorter than the first, third longer than the second, fourth to tenth transverse, eleventh as long as the three preceding antennomeres taken together. Eyes very large in dorsal view. Body devoid of reticulation. Head devoid of puncturation or granulation, except a large puncture near the posterior margin of each eye. Puncturation of the pronotum evident. Granulation of elytra fine and salient. Free abdominal tergites impunctate and devoid of granulation. Female fifth free tergite with striae along the whole posterior margin, female sixth free tergite strongly punctate. C o m p a r a t i v e n o t e s: The antennae and the spermatheca of the new species are similar to those of C. borneensis PACE, 2008 also from Borneo. The new species differs in the greater length of the spermatheca, (0.24 mm; in C. borneensis 0.15 mm), and in the proximal portion of the spermatheca forming a complete coil, whereas in C. borneensis the corresponding coil is incomplete. E t y m o l o g y: The new species is named "red neck" in reference to its reddish pronotum., Published as part of Pace, R., 2014, Aleocharinae from Sabah (Borneo) collected by Guillaume de Rougemont (Coleoptera, Staphylinidae), pp. 727-794 in Linzer biologische Beiträge 46 (1) on page 743, DOI: 10.5281/zenodo.5306753

Orphnebius acutissimus nov.sp. (Figs 33 and 144-145) T y p e m a t e r i a l Holotype, Sabah, Danum Valley, B.R.L., f.i.t., 14-16.II.2007, G. de Rougemont leg. (CROU). D e s c r i p t i o n: Length 3.6 mm. Body shiny, reddish, antennae reddish with the three basal antennomeres and apical half of the eleventh yellowish-red, legs yellowishred. Second antennomere shorter than the first, third longer than the second, fourth to tenth strongly transverse. Eyes enormous. Body devoid of reticulation. Pronotum impunctate and devoid of granulation. Pronotum with only two strong discal punctures. Granulation of the elytra fine and sparse. Free abdominal tergites concave and bare. Male fifth free tergite with evident longitudinal striae. Aedeagus: Figs 144-145. C o m p a r a t i v e n o t e s: The apex of the aedeagus is acute in ventral view, unlike in any of the other species known from Borneo. E t y m o l o g y: The name of the new species refers to the apex of the aedeagus which is very acute in ventral view., Published as part of Pace, R., 2014, Aleocharinae from Sabah (Borneo) collected by Guillaume de Rougemont (Coleoptera, Staphylinidae), pp. 727-794 in Linzer biologische Beiträge 46 (1) on pages 749-750, DOI: 10.5281/zenodo.5306753

Pelioptera danumensis nov.sp. (Figs 12 and 90-92) T y p e m a t e r i a l Holotype, Sabah, Danum Valley, B.R.L., f.i.t., 14-16.II.2007, G. de Rougemont leg. (CROU). D e s c r i p t i o n: Length 2.57 mm. Body shiny, black, posterior margin of the three basal free abdominal tergites and pygidium reddish-brown, antennae black with the three basal antennomeres brown, legs yellow. Second antennomere shorter than the first, third longer than the second, fourth and fifth as long as wide, eighth to tenth transverse. Eyes longer than the postocular region in dorsal view. Body devoid of reticulation except on the fifth free abdominal tergite where it is very transverse and superficial. Puncturation of the head slightly evident. Granulation of pronotum and elytra very close and superficial, that of the abdomen fine and very superficial. Male fifth abdominal tergite with a postero-median tubercle. Aedeagus: Figs 90-91; male sixth free tergite: Fig. 92. C o m p a r a t i v e n o t e s: The aedeagus of the new species is similar to that of P. bituberculata PACE, 2003 from West Malaysia, but it is clearly more curved to the ventral side and its apical part, in ventral view, is very wide, while it is very narrow in P. bituberculata. E t y m o l o g y: The new species takes its name from the Danum Valley., Published as part of Pace, R., 2014, Aleocharinae from Sabah (Borneo) collected by Guillaume de Rougemont (Coleoptera, Staphylinidae), pp. 727-794 in Linzer biologische Beiträge 46 (1) on pages 737-738, DOI: 10.5281/zenodo.5306753

Drusilla rougemontiana nov.sp. (Figs 47 and 185-188) T y p e m a t e r i a l Holotype, Sabah, Danum Valley, B.R.L., f.i.t., 14-16.II.2007, G. de Rougemont leg. (CROU). Paratypes: 14 exx., same provenance as the holotype (CROU, MSNV). D e s c r i p t i o n: Length 4.3-4.54 mm. Body shiny, blackish-brown, abdomen yellowish-red with the free third to sixth free abdominal tergites brown, antennae blackish-brown with the two basal antennomeres and the base of third reddish-brown, legs yellow with knees yellowish-brown. Second antennomere shorter than the first, third longer than the second, fourth to tenth transverse. Eyes longer than the postocular region in dorsal view. Body devoid of reticulation except on the sixth free tergite on which it is very transverse and superficial. Puncturation of the head sparse and superficial, absent on a narrow longitudinal median band, that of the pronotum strong anteriorly, to the sides thin, that of the elytra very superficial and clear. Median sulcus of the pronotum superficial, ample median concavity of the pronotum limited posteriorly by a transverse edge and a triangular posterior median impression. Male fifth free abdominal tergites with six tubercles on the posterior margin and two anterior median tubercles. Aedeagus: Figs 186-187; spermatheca: Fig. 185; male sixth free tergite: Fig. 188. C o m p a r a t i v e n o t e s: The habitus of this new species is similar to D. sculpticollis PACE, 2008, but the aedeagus and the spermatheca are different. The apex of the aedeagus, in ventral view, is more constricted and longer than that of D. sculpticollis, and the proximal portion of the spermatheca of the new species is not wound in a coil as in D. sculpticollis. E t y m o l o g y: The new species is dedicated to its collector, our colleague, the staphylinid specialist Guillaume de Rougemont., Published as part of Pace, R., 2014, Aleocharinae from Sabah (Borneo) collected by Guillaume de Rougemont (Coleoptera, Staphylinidae), pp. 727-794 in Linzer biologische Beiträge 46 (1) on page 758, DOI: 10.5281/zenodo.5306753

Atheta (Poromicrodota) borneotibialis nov.sp. (Figs 11 and 89) T y p e m a t e r i a l Holotype, Sabah, Danum Valley, B.R.L., f.i.t., 14-16.II.2007, G. de Rougemont leg. (CROU). D e s c r i p t i o n: Length 2.42 mm. Body shiny, reddish-brown with fourth free abdominal tergite brown, antennae brown with the two basal antennomeres reddishbrown, yellow legs. Second antennomere shorter than the first, third as long as the second, fourth to seventh longer that wide, eighth and ninth as long as wide, tenth transverse. Eyes longer than the postocular region in dorsal view. Head and pronotum devoid of reticulation, that of the elytra superficial, that of the abdomen very transverse. Granulation of head and pronotum fine, very superficial, that of the elytra very superficial and evident, that of the abdomen fine and very superficial. Spermatheca: Fig. 89. C o m p a r a t i v e n o t e s: For the form of the spermatheca the new species is similar to A. subaegra CAMERON, 1939 from India of which I have examined the female holotype labelled "Haldwani Dist., Kumaon, D.G. Champion, Atheta suabegra CAM., Type" (MHNL). The new species differs from it by the median and posterior tibiae which each bear two long bristles, which are absent in A. subaegra, in the short apical umbilicus of the distal bulb of the spermatheca, which is deep in A. subaegra, and in its smaller size: length, 0.25 mm; in A. subaegra 0.39 mm. E t y m o l o g y: The name of the new species means "Borneo tibial" alluding to the middle and posterior tibiae with long bristles., Published as part of Pace, R., 2014, Aleocharinae from Sabah (Borneo) collected by Guillaume de Rougemont (Coleoptera, Staphylinidae), pp. 727-794 in Linzer biologische Beiträge 46 (1) on page 736, DOI: 10.5281/zenodo.5306753

Psephothetemusa rougemonti nov.sp. (Figs 6 and 79) T y p e m a t e r i a l Holotype, Sabah, Danum Valley, B.R.L., f.i.t, 14-16.II.2007, G. de Rougemont leg. (CROU). D e s c r i p t i o n: Length 1.66 mm. Body shiny, yellowish-red, antennae brown with the three basal antennomeres pale yellow, legs yellow. Second antennomere as long as the first, third shorter than the second, fourth to tenth transverse. Eyes longer than the postocular region in dorsal view. Body devoid of reticulation. Puncturation of the head indistinct, that of the pronotum scarcely visible, except for four strong punctures, two on either side of the midlongitudinal band. Granulation of the elytra fine, evident and very superficial. Basal half of the first free abdominal tergites with imbricate sculpture, second to fourth free tergites covered with salient longitudinal striae, on the fifth free tergite these striae are superficial. Spermatheca: Fig. 79. C o m p a r a t i v e n o t e s: The form of the spermatheca of the new species presents an affinity with P. introflexa PACE, 2002 from Borneo, but while the apical umbilicus of the distal bulb of the spermatheca of P. introflexa is narrow and long, in the new species the same is wide and short. The distal bulb of the spermatheca of the new species is oval transverse, while in P. introflexa is sub-spherical. E t y m o l o g y: The new species is dedicated to its collector, our colleague, the staphylinid specialist Guillaume de Rougemont., Published as part of Pace, R., 2014, Aleocharinae from Sabah (Borneo) collected by Guillaume de Rougemont (Coleoptera, Staphylinidae), pp. 727-794 in Linzer biologische Beiträge 46 (1) on pages 732-733, DOI: 10.5281/zenodo.5306753

Apimela perarmata nov.sp. (Figs 59 and 229-230) T y p e m a t e r i a l Holotype, Sabah, Danum Valley, B.R.L., f.i.t., 14-16.II.2007, G. de Rougemont leg. (CROU). Paratypes: 1 and 2, same provenance as the holotype (CROU, MSNV). D e s c r i p t i o n: Length 2-2.12 mm. Body shiny, brown, head third and fourth free abdominal tergites blackish-brown, antennae black with the two basal antennomeres reddish-brown, legs dirty yellow. Second antennomere shorter than the first, third shorter than the second, fourth and fifth as long as wide, sixth to tenth transverse. Eyes shorter than the postocular region in dorsal view. Fore-body devoid of reticulation, that of the abdomen transverse and strong. Head very finely and densely punctate. Granulation of pronotum and elytra very dense and superficial, that of the first three free tergites fine and close, that of the two following less close. Longitudinal median impression of the pronotum feeble. Aedeagus: Fig. 229; spermatheca: Fig. 230. C o m p a r a t i v e n o t e s: The aedeagus of the new species is similar to that of A. kinabaluensis PACE, 2008 also from Borneo, but the length of the aedeagus of the new species is 0.17 mm, while the length of the aedeagus of A. kinabaluensis is 0.22 mm. The internal sclerotized pieces of the aedeagus of the new species is includes a curved lamina extending to the apex, that of A. kinabaluensis a thin tubule. E t y m o l o g y: The name of the new species "highly armed" refers to the strongly sclerotised pieces of the aedeagus., Published as part of Pace, R., 2014, Aleocharinae from Sabah (Borneo) collected by Guillaume de Rougemont (Coleoptera, Staphylinidae), pp. 727-794 in Linzer biologische Beiträge 46 (1) on pages 765-766, DOI: 10.5281/zenodo.5306753

Orphnebius acutus nov.sp. (Figs 31 and 140-141) T y p e m a t e r i a l Holotype, Sabah, Danum Valley, B.R.L., f.i.t., 14-16.II.2007, G. de Rougemont leg. (CROU). D e s c r i p t i o n: Length 2-2.2 mm. Body shiny, yellowish-red, including antennae and legs. Second antennomere shorter than the first, third as long as the second, fourth to tenth strongly transverse. Eyes longer than the postocular region in dorsal view. Body devoid of reticulation. Puncturation of the head indistinct, that of the pronotum reduced to four punctures, two discal and two lateral; puncturation of the elytra fine and very superficial. Free abdominal tergites concave and bare. Male fifth free tergite with strong elongate punctures. Aedeagus: Figs 140-141. C o m p a r a t i v e n o t e s: The apex of the aedeagus of the new species bears a ventral preapical tooth. The only species known with a similar tooth on the aedeagus is O. vorax PACE, 2000 from Thailand, but in that species the tooth is on the dorsal preapical region and not the ventral side as in the new species. The length of the aedeagus of the new species is of 0.78 mm, that of O. vorax 0.67 mm. E t y m o l o g y: The name of this new species is derived from the acute apex of the aedeagus, as seen in both lateral and ventral views., Published as part of Pace, R., 2014, Aleocharinae from Sabah (Borneo) collected by Guillaume de Rougemont (Coleoptera, Staphylinidae), pp. 727-794 in Linzer biologische Beiträge 46 (1) on page 749, DOI: 10.5281/zenodo.5306753, {"references":["PACE R. (2000): Aleocharinae della Thailandia (Coleoptera, Staphylinidae) (144 ° Contributo alla conoscenza delle Aleocharinae). - Bollettino del Museo regionale di Scienze naturali di Torino 17: 39 - 86."]}

Drusilla danumensis nov.sp. (Figs 48 and 189-191) T y p e m a t e r i a l Holotype, Sabah, Danum Valley, B.R.L., f.i.t, 14-16.II.2007, G. de Rougemont leg. (CROU). Paratypes: 6 exx., same provenance as the holotype (CROU, MSNV). D e s c r i p t i o n: Length 3.35-3.48 mm. Body shiny, head and pronotum blackishbrown, elytra black with the base dirty yellow, abdomen yellowish-red with posterior portion of the paratergites blackish-brown and first to third free tergites reddish-brown on their mid-longitudinal area, fourth free tergite yellowish-red in basal half and blackish-brown in distal half, fifth free tergite brown, antennae reddish-brown with the three basal antennomeres and the eleventh yellowish-red, legs yellowish-red with femora yellow, except to the distal extremities which are yellowish-red. Second antennomere shorter than the first, third longer than the second, fourth to ninth as long as wide, tenth transverse. Eyes longer than the postocular region in dorsal view. Body devoid of reticulation. Puncturation of the head fine and absent on the longitudinal median band, that of the elytra evident and deep. Granulation of the pronotum fine, very superficial and salient. Two large median punctures on each free tergite that are otherwise bare. There is a deep depression between the antennae. Pronotum with a deep median sulcus, an ample anterior median impression, a deep impression on either side and two strong punctures near the median sulcus. Aedeagus: Figs 190-191; spermatheca: Fig. 189. C o m p a r a t i v e n o t e s: In the form of the spermatheca and partly that of the aedeagus, and in the colour of the abdomen this new species is similar D. fontis PACE, 2008 also from Borneo. It differs in the transverse pronotum with length/width ratio of 0.88, whereas in D. fontis the pronotum is longer than wide with a length/width ratio of 1.10. The ventral profile of the aedeagus of the new species has an acute angle near the "crista apicalis", which is absent in D. fontis. The apex of the aedeagus of the new species, in ventral view, is not angled as the apex of the aedeagus of D. fontis. E t y m o l o g y: The name of the new species is derived from the Danum Valley., Published as part of Pace, R., 2014, Aleocharinae from Sabah (Borneo) collected by Guillaume de Rougemont (Coleoptera, Staphylinidae), pp. 727-794 in Linzer biologische Beiträge 46 (1) on pages 758-759, DOI: 10.5281/zenodo.5306753

Drusilla sabahensis nov.sp. (Figs 44 and 181-182) T y p e m a t e r i a l Holotype, Sabah, Danum Valley, B.R.L., f.i.t., 14-16.II.2007, G. de Rougemont leg. (CROU). D e s c r i p t i o n: Length 3.93 mm. Body shiny, blackish-brown, antennae brown with the two basal antennomeres and the base of third yellowish-red, legs yellowish-red, the femora yellow with distal extremity brown. Second antennomere shorter than the first, third longer than the second, fourth to seventh longer than wide, eighth as long as wide, ninth and tenth transverse. Eyes very large, in dorsal view. Fore-body devoid of reticulation, that of the abdomen very transverse and superficial, but the male sixth free tergite with slightly transverse and evident reticulation. Puncturation of the head superficial and absent on the longitudinal median band. Puncturation of pronotum and elytra strong and very superficial. There is a transverse sulcus between the antennae. Pronotum with a deep median sulcus ending posteriorly in a deep dimple. Aedeagus: Figs 181-182. C o m p a r a t i v e n o t e s: In the form of the aedeagus and in the habitus the new species is similar to D. foeda PACE, 2008 also from Borneo, but the middle and posterior femora are clearly bicoloured, the intermediary antennomeres are longer than wide (transverse in D. foeda), and the apex of the aedeagus in ventral view is abruptly dilated, (not dilated in D. foeda). E t y m o l o g y: The new species takes its name from Sabah., Published as part of Pace, R., 2014, Aleocharinae from Sabah (Borneo) collected by Guillaume de Rougemont (Coleoptera, Staphylinidae), pp. 727-794 in Linzer biologische Beiträge 46 (1) on pages 756-757, DOI: 10.5281/zenodo.5306753

Chaetosogonocephus danumensis nov.sp. (Figs 21 and 109-111) T y p e m a t e r i a l Holotype, Sabah, Danum Valley, B.R.L., f.i.t., 14-16.II.2007, G. de Rougemont leg. (CROU). Paratype: 1, same provenance as the holotype (CROU). D e s c r i p t i o n: Length 2.34-2.36 mm. Body Shiny, yellowish-red, antennae brown with the two basal antennomeres and base of the third pale yellow, legs yellowish-red. Second antennomere as long the first, third shorter than the second, fourth to ninth antennomeres strongly transverse, tenth weakly transverse, eleventh as long as the two and a half preceding antennomeres together. Eyes very large in dorsal view. Body devoid of reticulation or granulation. Puncturation of the head very superficial, denser between the antennae and the eye, largely absent on the longitudinal median band and occipital area. Granulation of pronotum and elytra salient, fine and sparse. Free abdominal tergites impunctate and devoid of granulation. Male fifth free tergite with strong and dense puncturation, male sixth free tergite with strong puncturation and very superficial only on the basal half, sparse posteriorly. Aedeagus: Figs 109-110; spermatheca: Fig. 111. C o m p a r a t i v e n o t e s The new species is similar to C. kinabaluensis PACE, 2008, also from Borneo, in the ventral profile of the aedeagus, but in ventral view the aedeagus of the new species is narrowed to the apex, while it is wide in C. kinabaluensis, and the antennae are brown with yellow base in the new species, while in C. kinabaluensis they are entirely yellow. The proximal portion of the spermatheca of the new species is folded in tight U shape, whereas in C. kinabaluensis the proximal portion of its spermatheca is merely slightly curved. E t y m o l o g y: The new species takes its name from the Danum Valley., Published as part of Pace, R., 2014, Aleocharinae from Sabah (Borneo) collected by Guillaume de Rougemont (Coleoptera, Staphylinidae), pp. 727-794 in Linzer biologische Beiträge 46 (1) on page 744, DOI: 10.5281/zenodo.5306753

Gyrophaena manus nov.sp. (Figs 1 and 65-68) T y p e m a t e r i a l Holotype, Sabah, Danum Valley, B.R.L., f.i.t., 14-16.II.2007, G. de Rougemont leg. (CROU). Paratypes: 8 exx., same provenance as the holotype (CROU, MSNV). D e s c r i p t i o n: Length 2-2.2 mm. Body shiny, brown, antennae yellow with the three basal antennomeres pale yellow, legs yellow. Second antennomere shorter than the first, third shorter than the second, fourth transverse, antennomeres five to ten transverse. Eyes longer than the postocular region in dorsal view. Reticulation of the fore-body very superficial, on pronotum and elytra very transverse, that of the abdomen irregular, polygonal and evanescent. Puncturation of the head irregularly distributed and absent on the longitudinal median band, that of the pronotum evident and distributed as in figure 67. Granulation of the elytra sparse and salient, that of the abdomen evident, the granules longitudinally elongate. Aedeagus: Fig. 65; male sixth free abdominal tergite: Fig. 68; pronotum: Fig. 67. C o m p a r a t i v e n o t e s: Based on the simple linear form of the aedeagus, this new species is included within the group of G. appendiculata MOTSCHOULSKI, 1859 from India, which also includes G. borneensis CAMERON, 1943, of which I have examined 1 male and 2 females of the type series labelled "Sandakan, N. Borneo, G. borneensis CAMERON, fungus" (MSNL). In the shape of the male sixth free abdominal tergite the new species is near to G. microcicatricosa PACE, 2001 from Mt Kinabalu. The new species differs from G. microcicatricosa in the yellowish-red antennae (brown in G. microcicatricosa) and in the bi-sinuate aedeagus (curved aedeagus in G. microcicatricosa). E t y m o l o g y: The distal portion of the aedeagus of the new species in the shape of a hand with forearm suggested the name of the new species., Published as part of Pace, R., 2014, Aleocharinae from Sabah (Borneo) collected by Guillaume de Rougemont (Coleoptera, Staphylinidae), pp. 727-794 in Linzer biologische Beiträge 46 (1) on page 730, DOI: 10.5281/zenodo.5306753, {"references":["PACE R. (2001): Nuove specie del genere Gyrophaena MANNERHEIM del Monte Kinabalu (Borneo) (Coleoptera, Staphylinidae). - Revue suisse de Zoologie 108: 699 - 737."]}

Drusilla borneoapicalis nov.sp. (Figs 49 and 192-194) T y p e m a t e r i a l Holotype, Sabah, Danum Valley, B.R.L., f.i.t, 14-16.II.2007, G. de Rougemont leg. (CROU). Paratypes: 1 and 1, same provenance as the holotype (CROU, MSNV). D e s c r i p t i o n: Length 4.54-4.69 mm. Body shiny, head brown, pronotum reddish, elytra blackish-brown with base reddish, abdomen yellowish-red with posterior margin of the first four free tergites largely brown, the fifth and sixth free tergites brown, antennae reddish-brown with the two basal antennomeres, the base of third and eleventh yellowish-red, legs yellowish-red with basal ¾ of the femora yellow. Second antennomere shorter than the first, third longer than the second, fourth to tenth longer than wide. Eyes very large in dorsal view. Body devoid of reticulation. Granulation of the head salient but absent on the longitudinal median band. Puncturation of the pronotum strong but absent on the anterior portion of the lateral depressions. Granulation of the elytra very superficial and salient. Free tergites bare, only the fifth free tergite with two strong median punctures. The head bears a salient tubercle between the antennae, and the pronotum a deep median sulcus. Aedeagus: Figs 192-193; spermatheca: Fig. 194. C o m p a r a t i v e n o t e s: The habitus and the spermatheca of the new species are similar to those of D. bruneiensis PACE, 2008, but the eyes of the new species are larger, longer than the postocular region, whereas in D. bruneiensis the eyes are as long or almost as the postocular region. The spermatheca of the new species is shorter (0.14 mm) than that of D. bruneiensis (0.19 mm). The aedeagus of the new species is different in several respects, including the apex, which is narrow in ventral view, whereas it is broad in D. bruneiensis, and the internal sclerotised pieces of the aedeagus are devoid of the numerous thorns present in D. bruneiensis. E t y m o l o g y: The name of the new species "Borneo apical" refers to the apex of the aedeagus which is broad in lateral view and narrow in ventral view., Published as part of Pace, R., 2014, Aleocharinae from Sabah (Borneo) collected by Guillaume de Rougemont (Coleoptera, Staphylinidae), pp. 727-794 in Linzer biologische Beiträge 46 (1) on page 759, DOI: 10.5281/zenodo.5306753

Chaetosogonocephus minor nov.sp. (Figs 19 and 106-107) T y p e m a t e r i a l Holotype, Sabah, Danum Valley, B.R.L., f.i.t, 14-16.II.2007, G. de Rougemont leg. (CROU). Paratype: 1, same provenance as the holotype (CROU). D e s c r i p t i o n: Length 1.84-1.96 mm. Body shiny, brown, sides of the pronotum, humeral angles of elytra, posterior margins of the free abdominal tergites and pygidium reddish, antennae and legs yellowish-red. Second antennomere shorter than the first, third as long as the second, fourth to tenth strongly transverse, eleventh as long as the seven preceding antennomeres taken together. Eyes very large in dorsal view. Body devoid of reticulation. Puncturation of the head little evident, very superficial and absent on the longitudinal median band. Granulation of pronotum and elytra fine and very superficial, absent on the longitudinal median band and near the posterior margin of the pronotum. Free abdominal tergites impunctate and devoid of granulation. Male fifth free tergite with posterior median striae, male sixth free tergite densely punctate. Aedeagus: Figs 106-107. C o m p a r a t i v e n o t e s: In the length of the eleventh antennomere, the new species is similar to C. notaticornis PACE, 2008 also from Borneo, but the aedeagus is 0.2 mm long, while that of C. notaticornis measures 0.25 mm. The ventral profile of the aedeagus of the new species is slightly curved, while is clearly curved in C. notaticornis. The internal sclerotised piece of the aedeagus of the new species is short, whereas it is very long in C. notaticornis. E t y m o l o g y: The new species is called "smaller" for its small body size in comparison with the species described above., Published as part of Pace, R., 2014, Aleocharinae from Sabah (Borneo) collected by Guillaume de Rougemont (Coleoptera, Staphylinidae), pp. 727-794 in Linzer biologische Beiträge 46 (1) on page 743, DOI: 10.5281/zenodo.5306753

Drusilla caputserpentis nov.sp. (Figs 42 and 178) T y p e m a t e r i a l Holotype, Sabah, Danum Valley, B.R.L., f.i.t, 14-16.II.2007, G. de Rougemont leg. (CROU). D e s c r i p t i o n: Length 7.12 mm. Pronotum and elytra opaque, rest of the body shiny. Head blackish-brown, pronotum and elytra brown, abdomen yellowish-red, posterior margin of the first to fourth free abdominal tergites brown, fifth free tergite reddishbrown, antennae blackish-brown with the two basal antennomeres reddish-brown and apex of the eleventh yellow, legs yellowish-red. Second antennomere shorter than the first, third longer than the second, fourth to tenth longer than wide. Eyes shorter than the postocular region in dorsal view. Reticulation of the fore-body evident, that of the abdomen very transverse and superficial. Puncturation of the head evident, fairly close and absent on a narrow longitudinal median band and on the frons, that of pronotum and elytra very dense and deep, that of the abdomen fine and fairly close. Pronotum with median sulcus at the bottom of an ample longitudinal concavity and with feeble lateral impressions. Spermatheca: Fig. 178. C o m p a r a t i v e n o t e s: The spermatheca of the new species shares some of the characters of the spermatheca of D. perdensa PACE, 2004 from Thailand. Both have a triangular appendix on the distal bulb of the spermatheca, and the proximal portion of the spermatheca is folded. However the intermediary portion of the spermatheca of the new species is much dilated near the proximal bulb, which is not the case in D. perdensa. The apical umbilicus of the distal bulb of the spermatheca of the new species is narrow: 0.024 mm wide, whereas in D. perdensa it measures 0.036 mm. E t y m o l o g y: The name of the new species, meaning "snake head" refers to the shape of the distal bulb of the spermatheca., Published as part of Pace, R., 2014, Aleocharinae from Sabah (Borneo) collected by Guillaume de Rougemont (Coleoptera, Staphylinidae), pp. 727-794 in Linzer biologische Beiträge 46 (1) on pages 755-756, DOI: 10.5281/zenodo.5306753, {"references":["PACE R. (2004): Falagriini, Deremini, Athetini e Thamiaraeini del Borneo. - Revue suisse de Zoologie 111: 791 - 850."]}

Chaetosogonocephus luteicollis nov.sp. (Figs 22 and 112-113) T y p e m a t e r i a l Holotype, Sabah, Danum Valley, B.R.L., f.i.t., 14-16.II.2007, G. de Rougemont leg. (CROU). Paratype: 1, same provenance as the holotype (CROU). D e s c r i p t i o n: Length 1.93-2 mm. Body shiny, yellowish-red, head reddish, posterior half of the elytra brown, antennae reddish-brown with the two basal antennomeres yellow, legs reddish. Second antennomere shorter than the first, third longer than the second, fourth as long as wide, fifth to tenth transverse, eleventh as long as the two preceding antennomeres together. Eyes very large in dorsal view. Body devoid of reticulation or granulation. Puncturation of the head indistinct. Granulation of the pronotum superficial and sparse, that of the elytra salient and sparse. Free abdominal tergites impunctate and devoid of granulation. Male and female fourth free tergite with postero-lateral striae, fifth free tergite with longitudinal striae on the entire surface, sixth free tergite with strong and very superficial Puncturation, and with four striae posteriorly. Aedeagus: Figs 112-113; spermatheca: Fig. 111. C o m p a r a t i v e n o t e s: The aedeagus of this new species is broadly curved to the ventral side as in C. notaticornis PACE, 2008 also from Borneo, but the eleventh antennomere is as long as the three preceding antennomeres together, and not as long as the six preceding antennomeres as in C. notaticornis. The internal sclerotised piece of the aedeagus of the new species is short (0.1 mm), whereas in C. notaticornis it measures 0.15 mm. The colour of the body of C. notaticornis is uniformly yellowish-red, while in the new species it is reddish-brown with yellow pronotum. E t y m o l o g y: The new species is named "yellow neck" in reference to its reddishyellow pronotum., Published as part of Pace, R., 2014, Aleocharinae from Sabah (Borneo) collected by Guillaume de Rougemont (Coleoptera, Staphylinidae), pp. 727-794 in Linzer biologische Beiträge 46 (1) on page 744, DOI: 10.5281/zenodo.5306753

Lambanus borneensis nov.sp. (Figs 36 and 150-156) T y p e m a t e r i a l Holotype, Sabah, Danum Valley, B.R.L., f.i.t., 14-16.II.2007, G. de Rougemont leg. (CROU). Paratypes: 3, same provenance as the holotype (CROU, MSNV). D e s c r i p t i o n: Length 5.8-6.1 mm. Body shiny, black, antennae reddish-brown with the eleventh antennomere yellowish-red, legs yellowish-red. Second antennomere shorter than the first, third longer than the second, fourth to seventh longer than wide, eighth and ninth as long as wide, tenth transverse. Eyes longer than the postocular region in dorsal view. Reticulation of head and elytra superficial, that of the pronotum very evanescent, absent on abdomen. Puncturation of the head very superficial and close, absent on the longitudinal median band, that of the pronotum strong and close, sparse in the bottom of the deep median concavity, that of the elytra evident, that of the abdomen sparse and clearly visible. On either side of the deep median concavity of the pronotum there is a salient conical protuberance and lateral depressions. Aedeagus: Figs 151-152; male sixth free abdominal tergite: Fig. 153. E t y m o l o g y: The new species takes its name from Borneo., Published as part of Pace, R., 2014, Aleocharinae from Sabah (Borneo) collected by Guillaume de Rougemont (Coleoptera, Staphylinidae), pp. 727-794 in Linzer biologische Beiträge 46 (1) on page 752, DOI: 10.5281/zenodo.5306753

Drusilla borneoruficollis nov.sp. (Figs 53 and 209-213) T y p e m a t e r i a l Holotype, Sabah, Danum Valley, B.R.L., f.i.t, 14-16.II.2007, G. de Rougemont leg. (CROU). Paratypes: 9 exx., same provenance as the holotype (CROU, MSNV). D e s c r i p t i o n: Length 6.3-6.81 mm. Body shiny, pronotum and elytra weakly shiny. Body blackish-brown, pronotum reddish, elytra black, antennae blackish-brown with the two basal antennomeres and the base of third yellowish-red, tibiae and distal half of the femora brown, basal half of the femora yellowish-red, tarsi reddish. Second antennomere shorter than the first, third longer than the second, fourth to tenth longer than wide. Eyes very large in dorsal view. Head devoid of reticulation, that of the pronotum evident, that of the elytra strong, that of the abdomen very transverse and evident, but on the fifth free tergite the reticulation is irregular, polygonal and evident. Granulation of the pronotum and elytra dense and salient, that of the abdomen superficial. There is a longitudinally sulcate tubercle between the antennae (Fig. 213), disk of the head with a Y-shaped furrow. Pronotum with median sulcus and a depression on either side of it. Male fifth free tergite with four salient granules. Aedeagus: Figs 209- 210; spermatheca: Fig. 211; male sixth free tergite: Fig. 212. C o m p a r a t i v e n o t e s: The habitus of the new species is similar to that of D. bruneiorum PACE, 2008 also from Borneo, except for the colour of the elytra and abdomen. The large ventral gibbosity of the aedeagus of the new species, and the very long spermatheca, the proximal portion of which is wound in numerous tight coils, also distinguish the new species from D. bruneiorum. E t y m o l o g y: The name of the new species, meaning "Borneo red neck" refers to its reddish pronotum., Published as part of Pace, R., 2014, Aleocharinae from Sabah (Borneo) collected by Guillaume de Rougemont (Coleoptera, Staphylinidae), pp. 727-794 in Linzer biologische Beiträge 46 (1) on pages 761-762, DOI: 10.5281/zenodo.5306753

Pseudatheta rougemonti nov.sp. (Figs 7 and 80-81) T y p e m a t e r i a l Holotype, Sabah, Danum Valley, B.R.L., f.i.t., 14-16.II.2007, G. de Rougemont leg. (CROU). D e s c r i p t i o n: Length 1.66 mm. body shiny, brown, elytra blackish-brown with base yellow, abdomen blackish-brown with the two basal free tergites, base of the third and pygidium yellow; antennae brown with the two basal antennomeres yellow, apex of the eleventh dirty yellow, legs yellow. Second antennomere as long as the first, third shorter than the second, fourth as long as wide, fifth to tenth antennomeres transverse. Eyes a little longer than the postocular region in dorsal view. Reticulation of the head superficial, that of the pronotum absent, that of the elytra slightly evident, that of the abdomen evident and irregularly polygonal. Puncturation of the head very superficial and clear. Granulation of the pronotum dense and fine, that of the elytra very superficial, also fine, that of the abdomen superficial, except near the posterior margin of all free tergites on which the granules are salient. Male fifth free abdominal tergite elongate with dense granules. Aedeagus: Figs 80-81. C o m p a r a t i v e n o t e s: In the presence of an acute ventral lamina of the aedeagus, the new species is similar to both P. borneensis PACE, 2007 and to P. kinabaluensis PACE, 2007, both from Borneo. The new species is distinct from P. borneensis in the strongly transverse intermediary antennomeres, which are only slightly transverse in P. borneensis, in the aedeagus, which is smaller and devoid of an apical flagellum (apical flagellum present in P. borneensis) and in the rectilinear ventral appendix of the aedeagus, which is sinuate in P. borneensis. The new species is distinguished from P. kinabaluensis by its brown abdomen with yellowish-red pygidium, (abdomen uniformly dirty yellow in P. kinabaluensis), and by the rectilinear ventral appendix of the aedeagus, which is curved in P. kinabaluensis. E t y m o l o g y: The new species is dedicated to its collector, our colleague, the staphylinid specialist Guillaume de Rougemont., Published as part of Pace, R., 2014, Aleocharinae from Sabah (Borneo) collected by Guillaume de Rougemont (Coleoptera, Staphylinidae), pp. 727-794 in Linzer biologische Beiträge 46 (1) on page 733, DOI: 10.5281/zenodo.5306753

Drusilla sabahorum nov.sp. (Figs 51 and 199-202) T y p e m a t e r i a l Holotype, Sabah, Danum Valley, B.R.L., f. it., 14-16.II.2007, G. de Rougemont leg. (CROU). Paratypes: 1 and 1, same provenance as the holotype (CROU, MSNV). D e s c r i p t i o n: Length 5-5.3 mm. Body shiny, reddish, pronotum and abdomen yellowish-red, antennae blackish-brown with the two basal antennomeres and the base of third yellowish-red, legs yellow. Second antennomere shorter than the first, third longer than the second, fourth longer than wide, fifth to seventh as long as wide, eighth to tenth transverse. Eyes longer than the postocular region in dorsal view. Reticulation of the head strong, absent on frons and on the pronotum and elytra, that of the abdomen very transverse and very superficial. Puncturation of the head fine and scarcely visible, absent on a broad longitudinal median band. Puncturation of the pronotum irregularly distributed and evident, that of the elytra strong and very superficial. There is a salient tubercle between the antennae and two on the disk between which the surface is deeply concave. Pronotum with deep median concavity confluent posteriorly with a fovea, on either side deeply engraved. Male second free tergite with postero-median appendix strongly carinate and truncate (Fig. 202). Posterior margin of the male fourth free tergite with large tubercles, fifth with strong tubercles on the whole surface. Aedeagus: Figs 199-200; spermatheca: Fig. 201; abdomen of the male: Fig. 202. C o m p a r a t i v e n o t e s: In its habitus and the form of the spermatheca, the new species is similar to D. aerea (CAMERON, 1933) also from Borneo, of which I have examined 1 male and 1 female labelled as follows: "Borneo, Dr. Cameron Myrmedonia aerea Cam. " (MNHL). The new species differs from D. aerea in that the spermatheca is scarcely sinuous from the distal end to the proximal bulb, whereas it is very sinuous in D. aerea, in the presence of a truncate thorn on the posterior margin of the male first free abdominal tergite, and in the long narrow aedeagus without protruding flagellum, whereas in D. aerea the aedeagus is broad with long flagellum. E t y m o l o g y: The new species is dedicated to the inhabitants of Sabah., Published as part of Pace, R., 2014, Aleocharinae from Sabah (Borneo) collected by Guillaume de Rougemont (Coleoptera, Staphylinidae), pp. 727-794 in Linzer biologische Beiträge 46 (1) on pages 760-761, DOI: 10.5281/zenodo.5306753, {"references":["CAMERON M. (1933): Staphylinidae (Col.) from Mount Kinabalu. - Journal of the Federated Malay States Museums 17: 338 - 360."]}

Mesomegaskela rougemonti nov.sp. (Figs 17 and 101-103) T y p e m a t e r i a l Holotype, Sabah, Danum Valley, B.R.L., f.i.t., 14-16.II.2007, G. de Rougemont leg. (CROU). Paratypes: 5 exx., same provenance as the holotype (CROU, MSNV). D e s c r i p t i o n: Length 5-5.2 mm. Body shiny, reddish, head and elytra brown, antennae reddish-brown, legs reddish. The whole body covered in very superficial short and adherent, silky pubescence. Posterior margin of the elytra with eleven long bristles. Abdomen without reticulation and with fine and very superficial Puncturation. Posterior margin of each of the four basal free tergites with ten long bristles. Aedeagus: Figs 101- 102; spermatheca: Fig. 103. C o m p a r a t i v e n o t e s: In its habitus and the form of the spermatheca, the new species is similar to M. adesi PACE, 1999 from Hong Kong. It differs in the pronotum which has a length/width ratio of 0.85, in M. adesi 0.87, and above all in the form of the spermatheca. Whereas in M. adesi the proximal portion of the spermatheca is wound in four coils, in the new species the coils are more numerous (Fig. 103), and more convoluted. E t y m o l o g y: The new species is dedicated to its collector, our colleague, the staphylinid specialist Guillaume de Rougemont., Published as part of Pace, R., 2014, Aleocharinae from Sabah (Borneo) collected by Guillaume de Rougemont (Coleoptera, Staphylinidae), pp. 727-794 in Linzer biologische Beiträge 46 (1) on page 740, DOI: 10.5281/zenodo.5306753

Outachyusa borneensis nov.sp. (Figs 9 and 85) T y p e m a t e r i a l Holotype, Sabah, Danum Valley, B.R.L., f.i.t., 14-16.II.2007, G. de Rougemont leg. (CROU). Paratypes: 2, same provenance as the holotype (CROU, MSNV). D e s c r i p t i o n: Length 2.4-2.9 mm. Body opaque, brown, pronotum yellowishbrown, posterior margin of the elytra dirty yellow, antennae brown, legs pale yellow. Second antennomere as long as the first, third shorter than the second, fourth and fifth longer than wide, sixth and seventh as long as wide, eighth to tenth transverse. Eyes longer than the postocular region in dorsal view. The whole body covered with dense and fine granules and silky pubescence. Spermatheca: Fig. 85. C o m p a r a t i v e n o t e s: The habitus of the new species is similar to that of O. velox (CAMERON, 1939) from India, of which I have examined the type series of 17 specimens labelled "Dehra Dun, Dr. Cameron, 14.X.1922, Brachyusa velox Cam. " (MNHL). The fourth to tenth antennomeres of the new species are shorter than those of O. velox, and the proximal portion of the spermatheca is almost rectilinear in the new species, strongly curved in O. velox. E t y m o l o g y: The new species takes its name from Borneo., Published as part of Pace, R., 2014, Aleocharinae from Sabah (Borneo) collected by Guillaume de Rougemont (Coleoptera, Staphylinidae), pp. 727-794 in Linzer biologische Beiträge 46 (1) on page 735, DOI: 10.5281/zenodo.5306753

Drusilla bilobata nov.sp. (Figs 43 and 179-180) T y p e m a t e r i a l Holotype, Sabah, Danum Valley, B.R.L., f.i.t., 14-16.II.2007, G. de Rougemont leg. (CROU). D e s c r i p t i o n: Length 5 mm. Body shiny, brown, pronotum reddish, abdomen yellowish-red with the fourth and fifth free abdominal tergites reddish, antennae brown with the two basal antennomeres and the base of third yellowish-red, legs yellowish-red. Second antennomere shorter than the first, third longer than the second, fourth as long as wide, fifth to tenth transverse. Eyes longer than the postocular region in dorsal view. Reticulation of head and pronotum evident, that of the elytra superficial, that of the abdomen very transverse and clearly visible. Puncturation of the fore-body fine, very superficial and evident, that of the abdomen fine and superficial. The median sulcus of the pronotum does not reach the anterior margin and posteriorly it ends in a deep median dimple. Spermatheca: Fig. 179; female sixth free tergite: Fig. 180. C o m p a r a t i v e n o t e s: The habitus of the new species is similar to that of D. spissatheca PACE, 2008 also from Borneo, but the maximum width of the distal bulb of the spermatheca of the new species is greater (0.06 mm), than in spissatheca (0.048 mm). E t y m o l o g y: The name of the new species "two lobed" refers to the two large lobes of the sixth free abdominal tergite of the female., Published as part of Pace, R., 2014, Aleocharinae from Sabah (Borneo) collected by Guillaume de Rougemont (Coleoptera, Staphylinidae), pp. 727-794 in Linzer biologische Beiträge 46 (1) on page 756, DOI: 10.5281/zenodo.5306753

Pace, R. (2014): Aleocharinae from Sabah (Borneo) collected by Guillaume de Rougemont (Coleoptera, Staphylinidae). Linzer biologische Beiträge 46 (1): 727-794, DOI: http://doi.org/10.5281/zenodo.5306753, {"references":["BERNHAUER M. (1911): Zur Staphylinidenfauna Ostindiens und der Sundainseln (3. Beitrag. Schluss). - Entomologische Blatter fur Biologie und Systematik der Kafer 7: 86-93.","BERNHAUER M. (1915): Neue Staphyliniden der indo-malaiischen Fauna, insbesondere der Sunda-Insel Borneo. - Verhandlungen der Zoologisch-Botanischen Gesellschaft in Wien 65: 134-158.","CAMERON M. (1920): New species of Staphylinidae from Singapore, Part III. - Transaction of the Entomological Society of London 1920: 212-284.","CAMERON M. (1933): Staphylinidae (Col.) from Mount Kinabalu. - Journal of the Federated Malay States Museums 17: 338-360.","CAMERON M. (1937): Fauna Javanica. The Staphylinidae Collected by Mr. F.C. Drescher. - Tijdschrift voor Entomologie 80: 1-37.","CAMERON M. (1939a): New Staphylinidae (Col.) from New Guinea. - Annals and Magazine of natural History of London 3: 139-152.","CAMERON M. (1939b): The fauna of British India, including Ceylon and Burma. Coleoptera, Staphylinidae, 4 (Part 1). pp. 1-410. London.","CAMERON M. (1943a): New species of Staphylinidae (Col.) from Borneo. - Entomologist's Monthly Magazine 79: 39-42.","CAMERON M. (1943b): New species of Staphylinidae (Col.) from Borneo. - Entomologist's Monthly Magazine 79: 139-143.","FAUVEL A. (1905): Staphylinides de Java recueillies par M. le Dr. K. Kraepelin et M. le Dr. Koningsberger en 1904. - Mitteilungen aus dem Naturhistorischen Museum in Hamburg 22: 75-86.","HAMMOND P.M. (1984): An annotated Check-List of Staphylinidae (Insecta: Coleoptera) recorded from Borneo. - Sarawak Museum Journal 33: 187-218.","KISTNER D.H. (1990): Termitophilous and myrmecophilous Staphylinidae collected in Sierra Leone by W. Rossi (Coleoptera). - Accademia Nazionale dei Lincei 387, Quaderno 265: 155-164.","KISTNER D.H. & R. JACOBSON (1975): The Natural History of the Myrmecophilous Tribe Pygostenini (Coleoptera: Staphylinidae). - Sociobiology 1: 155-384.","KISTNER D.H. & M.J. MCNAIRM (1991): A new Genus and Two New species of Predacious Staphylinidae Associated with Pheidolegeton in Sulawesi (Coleoptera: Aleocharinae) with Notes on theis Behavior. - Sociobiology 18: 305-322.","KISTNER D.H., BERGHOFF S.M. & U. MASCHWITZ (2003): Myrmecophilous Staphylinidae (Coleoptera) associated with Dorylus (Dichthadius) laevigatus (Hymenoptera: Formicidae) in Malaysia with studies of their behavior. - Sociobiology 41: 203-264.","KLIMASZEWSKI J. & A. SMETANA (1990): The species of Aleochara GRAVENHORST from the Mount Kinabalu National Park, Sabah, northern Borneo (Coleoptera: Staphylinidae: Aleocharinae). - Annals of the Transvaal Museum 35: 157-169.","KRAATZ G. (1859): Die Staphyliniden-Fauna von Ostindien, insbesondere der Insel Ceylan. - Archiv fur Naturgeschichte 25: 1-45.","LEACH W.E. (1819): New genera. - In: Samuelle. - The Entomologist's useful Compendium, London 1819: 496 pp.","MOTSCHULSKY V.I. (1857. Enumeration des nouvelles Especes de Coleopteres rapportes de ses voyages. - Bulletin de la Societe Imperiale des Naturalistes de Moscou 31: 204-264.","PACE R. (1984): Aleocharinae delle Mascarene, parte I: tribu Myllaenini, Pronomaeini, Oligotini e Bolitocharini (Coleoptera Staphylinidae) (XLV Contributo alla conoscenza delle Aleocharinae). - Revue suisse de Zoologie 91: 3-36.","PACE R. (1986): Aleocharinae dell'Asia sudorientale raccolte da G. de Rougemont (Coleoptera, Staphylinidae). - Bollettino del Museo civico di Storia naturale di Verona 23: 139-237.","PACE R. (1988): Sphuridaethes loebli, nuovo genere e nuova specie di Aleocharinae termitofila del Borneo (Coleoptera Staphylinidae) (XCIX Contributo alla conoscenza delle Aleocharinae). - Revue suisse de Zoologie 95: 979-985.","PACE R. (1989a): Aleocharinae attere del Monte Kinabalu (Borneo) (Coleoptera Staphylinidae) (XCVIII Contributo alla conoscenza delle Aleocharinae). - Revue suisse de Zoologie 96: 3-8.","PACE R. (1989b): Monografia del genere Leptusa KRAATZ (Coleoptera Staphylinidae) (LXXV Contributo alla conoscenza delle Aleocharinae). - Memorie del Museo civico di Storia naturale di Verona (II S.) A: 1-307.","PACE R. (1993): Nuove Aleocharinae Orientali (Coleoptera, Staphylinidae). - Bollettino del Museo civico di Storia naturale di Verona 16: 127-180.","PACE R. (1999a): Aleocharinae di Hong Kong (Coleoptera, Staphylinidae). - Revue suisse de Zoologie 106: 663-689.","PACE R. (1999b): Aleocharinae della Cina: Parte V (conclusione) (Coleoptera,Staphylinidae). - Revue suisse de Zoologie 106: 107-164.","PACE R. (2000): Aleocharinae della Thailandia (Coleoptera, Staphylinidae) (144° Contributo alla conoscenza delle Aleocharinae). - Bollettino del Museo regionale di Scienze naturali di Torino 17: 39-86.","PACE R. (2001): Nuove specie del genere Gyrophaena MANNERHEIM del Monte Kinabalu (Borneo) (Coleoptera, Staphylinidae). - Revue suisse de Zoologie 108: 699-737.","PACE R. (2002a): Nuovi generi di Aleocharinae del Borneo (Coleoptera, Staphylinidae). - Revue suisse de Zoologie 109: 189-240.","PACE R. (2002b): Specie dei generi Eusteniamorpha CAMERON e Autalia LEACH del Borneo (Coleoptera, Staphylinidae). - Revue suisse de Zoologie 109: 295-323.","PACE R. (2002c): Le specie del genere Brachida MULSANT & REY del Monte Kinabalu (Borneo) (Coleoptera, Staphylinidae). - Revue suisse de Zoologie 109: 491-509.","PACE R. (2003a): Homalotini del Borneo (Coleoptera, Staphylinidae). - Revue suisse de Zoologie 110: 255-313.","PACE R. (2003b): Gyrophaenini dei generi Sternotropa, Pseudoligota e Adelarthra del Monte Kinabalu (Borneo, Sabah) (Coleoptera, Staphylinidae). - Revue suisse de Zoologie 110: 761-781.","PACE R. (2004): Falagriini, Deremini, Athetini e Thamiaraeini del Borneo. - Revue suisse de Zoologie 111: 791-850.","PACE R. (2005): Pronomaeini del Borneo (Coleoptera, Staphylinidae). - Revue suisse de Zoologie 112: 21-45.","PACE R. (2007a): Le specie del genere Orphnebius MOTSCHULSCKY, 1858, nel Borneo (Coleoptera, Staphylinidae). - Revue suisse de Zoologie 114: 743-769.","PACE R. (2007b): Le specie di Oligotini, Leucocraspedini, Hygrinomini, Placusini, Bolitocharini e Diestotini nel Borneo (Coleoptera, Staphylinidae). - Revue suisse de Zoologie 114: 771-815.","PACE R. (2008a): Generi e specie della tribu Lomechusini del Borneo (Coleoptera, Staphylinidae). - Revue suisse de Zoologie 115 (1): 107-156.","PACE R. (2008b): Le specie di Thamiaraeini, Oxypodini, Hoplandriini e Aleocharini del Borneo (Coleoptera, Staphylinidae). - Revue suisse de Zoologie 115 (1): 157-183.","PACE R. (2008c): Specie del genere Pseudoplandria FENYES,1921 nel Borneo (Coleoptera, Staphylinidae). - Revue suisse de Zoologie 115 (2): 397-431.","PACE R. (2008c): Le specie di Thamiaraeini, Oxypodini, Hoplandriini e Aleocharini del Borneo (Coleoptera, Staphylinidae). - Revue suisse de Zoologie 115 (1): 157-183.","PACE R. (2008d): Specie del genere Pseudoplandria FENYES, 1921 nel Borneo (Coleoptera, Staphylinidae). - Revue suisse de Zoologie 115 (2): 397-431.","PASNIK G. (1999): A new species of the genus Taxicera MULSANT et REY, 1873 from Borneo (Coleoptera, Staphylinidae: Aleocharinae). - Acta zoological cracoviensis 42: 369-371.","SAWADA K. (1982): Atheta and its allies of Southeast Asia. Oriental species described by V. von Motschulsky and G. Kraatz. - Contributions from the Biological Laboratory, Kyoto University 26: 141-187.","SEEVERS C.H. (1957): A monograph on the termitophilous Staaphylinidae (Coleoptera). - Fieldiana: Zoology 40: 344 pp.","WALKER F. (1859): Characters of some apparently undescribed Ceylon insects. - Annals and Magazine of Natural History 3: 52.","WASMANN E. (1898): Eine neue dorylophile Tachyporinen-Gattung aus Sudafrika. - Wiener Entomologische Zeitung 17: 101-103."]}

Rougemontius borneensis nov.sp. (Figs 24 and 117-119) T y p e m a t e r i a l Holotype, Sabah, Danum Valley, B.R.L., f.i.t., 14-16.II.2007, G. de Rougemont leg. (CROU). Paratype: 1, same provenance as the holotype (CROU). D e s c r i p t i o n: Length 3 mm. Fore-body a little shiny, abdomen shiny. Body yellowish-brown, head brown, abdomen yellowish-red, antennae yellowish-brown with antennomeres seven to ten pale yellow, legs dirty yellow. Antennae geniculate, scape long, second antennomere shorter than the first, third longer than the second, fourth as long as wide, fifth and sixth transverse, seventh to tenth much longer than wide. Eyes strongly developed. Reticulation of head and pronotum evident only in the bottom of the sulci, that of the elytra evident only in the bottom of the longitudinal depressions of each elytron, that of the abdomen fine. Body devoid of puncturation or granulation. Head with two swellings on each side of the ample median sulcus, and behind the eyes, a posterior transverse sulcus. Pronotum with two protuberant swellings in front to the anterior angles, a wide and deep longitudinal median depression posteriorly divided in two sulci. Posterior angles of the elytra. bulging and very protuberant, disk of each elytron with an ample, clear and deep longitudinal depression and another smaller one between this depression and the scutellum. Spermatheca sclerites absent, replaced by a semicircular structure (Fig. 117). E t y m o l o g y: The new species takes its name from Borneo., Published as part of Pace, R., 2014, Aleocharinae from Sabah (Borneo) collected by Guillaume de Rougemont (Coleoptera, Staphylinidae), pp. 727-794 in Linzer biologische Beiträge 46 (1) on pages 741-742, DOI: 10.5281/zenodo.5306753

Borneochara rougemonti nov.sp. (Figs 64 and 239-243) T y p e m a t e r i a l Holotype, Sabah, Danum Valley, B.R.L., f.i.t., 14-16.II.2007, G. de Rougemont leg. (CROU). D e s c r i p t i o n: Length 1.81 mm. Body shiny, yellowish-red, head and elytra reddish-brown, antennae reddish with the three basal antennomeres yellow, legs yellow. Second antennomere shorter than the first, third shorter than the second, fourth as long as wide, fifth to tenth transverse. Eyes very large in dorsal view. Body devoid of reticulation. Puncturation of the head invisible, that of the pronotum very sparse. Granulation of the elytra superficial and sparse. Free abdominal tergites bare and convex with long bristles on the posterior margins. The pronotum bears ample lateral impressions. Aedeagus: Figs 239-240. E t y m o l o g y: The new species is dedicated to its collector, our colleague, the staphylinid specialist Guillaume de Rougemont., Published as part of Pace, R., 2014, Aleocharinae from Sabah (Borneo) collected by Guillaume de Rougemont (Coleoptera, Staphylinidae), pp. 727-794 in Linzer biologische Beiträge 46 (1) on page 769, DOI: 10.5281/zenodo.5306753

Drusilla serrulae nov.sp. (Figs 40 and 169-173) T y p e m a t e r i a l Holotype, Sabah, Danum Valley, B.R.L., f.i.t., 14-16.II.2007, G. de Rougemont leg. (CROU). Paratypes: 11, same provenance as the holotype (CROU, MSNV). D e s c r i p t i o n: Length 4.5-4.69 mm. Fore-body slightly shiny, abdomen shiny. Body brown, abdomen reddish with the third to fifth free abdominal tergites brown, antennae blackish-brown with the two basal antennomeres and the base of third yellowish-red, legs yellowish-red. Second antennomere shorter than the first, third longer than the second, fourth to tenth transverse. Eyes longer than the postocular region in dorsal view. Reticulation of the fore-body evident, that of the abdomen transverse and clearly visible. Puncturation of head and abdomen very superficial and close, absent on frons. Strong granulation denser in the median occipital area. Granulation of pronotum and elytra dense and evident, more salient near the suture of the elytra. There is a salient umbonate elevation between the antennae. The pronotum bears a fine median sulcus. Male fifth free abdominal tergites with two large posterior median punctures, arranged in a transverse line. Aedeagus: Figs 169-170; spermatheca: Fig. 173, male sixth free tergite: Fig. 171; female sixth free tergite: Fig. 172. C o m p a r a t i v e n o t e s: The habitus and shape of the spermatheca of this new species point to an affinity with D. operosa PACE, 1986 also from Borneo. However the fourth antennomere of the new species is transverse, whereas that of D. operosa is longer than wide. The intermediary portion of the spermatheca between the distal and proximal bulbs is short (0.07 mm) in the new species, whereas it is 0.12 mm long in D. operosa. E t y m o l o g y: The name of the new species means "serrate" (from the Latin serrula = a saw) in reference to one of the internal sclerotised pieces of the aedeagus which is shaped like the teeth of a saw., Published as part of Pace, R., 2014, Aleocharinae from Sabah (Borneo) collected by Guillaume de Rougemont (Coleoptera, Staphylinidae), pp. 727-794 in Linzer biologische Beiträge 46 (1) on pages 754-755, DOI: 10.5281/zenodo.5306753, {"references":["PACE R. (1986): Aleocharinae dell'Asia sudorientale raccolte da G. de Rougemont (Coleoptera, Staphylinidae). - Bollettino del Museo civico di Storia naturale di Verona 23: 139 - 237."]}

Orphnebius concavus nov.sp. (Figs 34 and 146-147) T y p e m a t e r i a l Holotype, Sabah, Danum Valley, B.R.L., f.i.t, 14-16.II.2007, G. de Rougemont leg. (CROU). D e s c r i p t i o n: Length 3.48 mm. Body shiny, reddish, posterior ¼ of the elytra brown, antennae brown with the three basal antennomeres and apex of the eleventh yellowish-red, legs yellowish-red. Second antennomere shorter than the first, third longer than the second, fourth and fifth as long as wide, seventh to tenth transverse. Eyes very large. Body devoid of reticulation. Head impunctate, devoid of granulation. Puncturation of the pronotum evident and sparse, with two large anterior punctures. Granulation of the elytra clearly visible, fairly dense. Free abdominal tergites convex and bare. Male sixth free tergite with a semicircular median fold. Aedeagus: Figs 146-147. C o m p a r a t i v e n o t e s: The aedeagus of this new species bears two lateral lamina between which the surface is concave. A similar aedeagus has never been observed before. E t y m o l o g y: The name of the new species refers to the ventral concavity of the aedeagus., Published as part of Pace, R., 2014, Aleocharinae from Sabah (Borneo) collected by Guillaume de Rougemont (Coleoptera, Staphylinidae), pp. 727-794 in Linzer biologische Beiträge 46 (1) on page 750, DOI: 10.5281/zenodo.5306753

Drusilla divergens nov.sp. (Figs 38 and 162-163) T y p e m a t e r i a l Holotype, Sabah, Danum Valley, B.R.L., f.i.t., 14-16.II.2007, G. de Rougemont leg. (CROU). Paratypes: 2, same provenance as the holotype (CROU, MSNV). D e s c r i p t i o n: Length 7-7.12 mm. Body shiny, blackish-brown, abdomen reddish, posterior half of the male fourth free abdominal tergite and the whole fourth brown, antennae reddish, legs yellowish-red. Second antennomere shorter than the first, third longer than the second, fourth to tenth longer than wide. Eyes longer than the postocular region in dorsal view. Reticulation of the fore-body evident and fine. Reticulation of the abdomen absent. Puncturation of the head visible, but absent on a narrow longitudinal median band and on frons. Puncturation of the elytra evident and close, that of the abdomen fine. Aedeagus: Figs 162-163. C o m p a r a t i v e n o t e s:Thenewspeciesissimilarto D. foeda PACE, 2008 also from Borneo. It differs in the lenghth/width ratio of the pronotum which is 0.93, whereas in D. foeda it is 1.11, in the antennomeres four to eight which are longer than wide, whereas in D. foeda they are transverse, and in the broad, truncate apex of the aedeagus, which in D. foeda is acute. E t y m o l o g y: The specific epithet of the new species refers from the divergence of the characters of the aedeagus from those of D. foeda PACE., Published as part of Pace, R., 2014, Aleocharinae from Sabah (Borneo) collected by Guillaume de Rougemont (Coleoptera, Staphylinidae), pp. 727-794 in Linzer biologische Beiträge 46 (1) on pages 753-754, DOI: 10.5281/zenodo.5306753

Drusilla trina nov.sp. (Figs 52 and 203-208) T y p e m a t e r i a l Holotype, Sabah, Danum Valley, B.R.L., f.i.t., 14-16.II.2007, G. de Rougemont leg. (CROU). Paratypes: 6 and 3, same provenance as the holotype (CROU, MSNV). D e s c r i p t i o n: Length 8-8.2 mm. Fore-body weakly shiny, abdomen shiny. Body black, antennae black with the two basal antennomeres and the base of third reddish, legs black with basal half of the femora yellow. Second antennomere shorter than the first, third longer than the second, fourth to tenth longer than wide. Eyes longer than the postocular region in dorsal view. Reticulation of the body strong, that of pronotum and elytra evident and that of the abdomen weakly transverse and clearly visible. Granulation of head and pronotum salient and very close, that of the elytra very dense. Puncturation of the abdomen very superficial, evident on the three basal free tergites, sparse on the following tergites. Head with a Y-shaped median sulcus, a carina between the antennae and devoid of granulation. Pronotum with an ample median concavity, deep median sulcus and lateral impressions. First free tergite of the male with three lobes on the posterior margin (Fig. 203), the second with a triangular baso-median area with transverse striae, third with a basal median area, fourth with six posterior marginal tubercles, fifth with fine salient tubercles (Fig. 203). Aedeagus: Figs 204-205; spermatheca: Fig. 206; male sixth free tergite: Fig. 207; female sixth free tergite: Fig. 208; abdomen: Fig. 203. C o m p a r a t i v e n o t e s: The habitus and the form of the aedeagus of this new species are comparable to D. bruneiorum PACE, 2008 also from Borneo, but the fifth to eighth antennomeres are longer than wide, whereas they are transverse in D. bruneiorum, the anterior and middle legs are yellowish-red in bruneiorum, and the aedeagus of the new secies is broadly curved to the ventral side, while it more sharply angled in D. bruneiorum. E t y m o l o g y: The name of the new species, meaning "triple" refers to the posterior margin of the first free tergites which are tri-lobed in the male., Published as part of Pace, R., 2014, Aleocharinae from Sabah (Borneo) collected by Guillaume de Rougemont (Coleoptera, Staphylinidae), pp. 727-794 in Linzer biologische Beiträge 46 (1) on page 761, DOI: 10.5281/zenodo.5306753

Aleochara (Aleochara) sabahensis nov.sp. (Figs 63 and 237-238) T y p e m a t e r i a l Holotype, Sabah, Danum Valley, B.R.L., f.i.t., 14-16.II.2007, G. de Rougemont leg. (CROU). Paratypes: 2, same provenance as the holotype (CROU, MSNV). D e s c r i p t i o n: Length 8.5-9 mm. Body shiny, the abdomen slightly iridescent. Body black, sides of the elytra brown, antennae and legs black, tarsi brown. Second antennomere shorter than the first, third longer than the second, fourth to tenth strongly transverse. Eyes longer than the postocular region in dorsal view. Body devoid of reticulation. Puncturation of head and pronotum umbilicate, evident and close, that of the elytra dense and finer than that of the pronotum, that of the abdomen composed of dense and evident elongate punctures. The disc of pronotum bears four punctures disposed in a square and a lateral puncture. Aedeagus: Figs 237-238. C o m p e r a t i v e n o t e s: The aedeagus of the new species is similar to that of A. paleonigra PACE, 1999 (olim A. nigra KRAATZ, 1859), but the internal sclerotised piece of the aedeagus is different in the two species. In lateral profile a plate in the distal portion of the internal sclerotised piece of the aedeagus is shaped like a bid’s head, whereas in A. paleonigra this plate is sickle-shaped. The internal sclerotised piece of the aedeagus of the new species is less developed in length than that of A. paleonigra. These comparisons are based on my examination of specimens of the type series of A. nigra Kraatz labelled as follows: " Ceylon, J. Nietner, Aleochara nigra Kr " (DEI). E t y m o l o g y: The new species takes its name from Sabah., Published as part of Pace, R., 2014, Aleocharinae from Sabah (Borneo) collected by Guillaume de Rougemont (Coleoptera, Staphylinidae), pp. 727-794 in Linzer biologische Beiträge 46 (1) on page 768, DOI: 10.5281/zenodo.5306753, {"references":["KRAATZ G. (1859): Die Staphyliniden-Fauna von Ostindien, insbesondere der Insel Ceylan. - Archiv fur Naturgeschichte 25: 1 - 45."]}

Tetrabothrus femoralis nov.sp. (Figs 25 and 120-122) T y p e m a t e r i a l Holotype, Sabah, Danum Valley, B.R.L., f.i.t., 14-16.II.2007, G. de Rougemont leg. (CROU). Paratypes: 5 exx., same provenance as the holotype (CROU, MSNV). D e s c r i p t i o n: Length 5.5-5.9 mm. Body shiny, reddish, antennae reddish, legs yellowish-red with reddish-brown knees with basal ¾ of the femora pale yellow. Second antennomere shorter than the first, third longer than the second, fourth to tenth strongly transverse. Eyes shorter than the postocular region in dorsal view. Body devoid of reticulation. Puncturation of head and pronotum fine, absent on the longitudinal median band of both. Granulation of the elytra fine, very superficial and little evident. Four basal sulci of the free abdominal tergites deep. Aedeagus: Figs, 120-121; spermatheca: 122. C o m p a r a t i v e n o t e s: The aedeagus of this new species is more similar to that of T. indicus CAMERON, 1939 from Bengal, than to that T. borneensis CAMERON, 1943 from Borneo, of which I have seen specimens of the type series. The new species differs from T. indicus in the apex of the aedeagus, which is very narrow in ventral view (less narrow in T. indicus), in the distance between the "crista apicalis" and bottom of the curved preapical portion which measures 0.1 mm, (0.05 mm in T. indicus). The internal sclerotised piece of the aedeagus of the new species is less strong than that of T. indicus. The male holotype of T. indicus is labelled "Samsing, 1800, Kalimpong, Bengal, XII.1933, Balwant Singh, in soil under elephant dung, Tetrabothrus indicus Cam., Type."(MHNL). E t y m o l o g y: The pale yellow colour of the basal ¾ of the femora suggested the name of the new species., Published as part of Pace, R., 2014, Aleocharinae from Sabah (Borneo) collected by Guillaume de Rougemont (Coleoptera, Staphylinidae), pp. 727-794 in Linzer biologische Beiträge 46 (1) on page 745, DOI: 10.5281/zenodo.5306753

Chaetosogonocephus nobilis nov.sp. (Figs 18 and 104-105) T y p e m a t e r i a l Holotype, Sabah, Danum Valley, B.R.L., f.i.t., 14-16.II.2007, G. de Rougemont leg. (CROU). D e s c r i p t i o n: Length 3 mm. Body shiny, brown, pygidium reddish, antennae and legs yellowish-red. Second antennomere shorter than the first, third longer than the second, fourth to tenth strongly transverse. Eyes very large in dorsal view. Body devoid of reticulation. Head impunctate. Granulation of the pronotum and elytra fine, that of the abdomen superficial and present only near the posterior margin of each free tergite. Male fifth free tergite with strong elongate punctures, male sixth free tergite covered with strong puncturation. Aedeagus: Figs 104-105. C o m p a r a t i v e n o t e s: The habitus of the new species is very similar to that of C. borneensis PACE, 2008 also from Borneo. It differs in the eleventh antennomere being as long as the seven preceding antennomeres taken together, while in C. borneensis the eleventh antennomere is as long as the three preceding antennomeres. The form of the aedeagus is also different. That of the new species has two ventral lamina in ventral view, which are absent in the aedeagus of C. borneensis, and the internal sclerotised piece of the aedeagus of the new species is not hooked as that of C. borneensis. E t y m o l o g y: The new species is called "noble" for its greater body size, suggesting superiority or nobility in comparison with the other species described in this paper., Published as part of Pace, R., 2014, Aleocharinae from Sabah (Borneo) collected by Guillaume de Rougemont (Coleoptera, Staphylinidae), pp. 727-794 in Linzer biologische Beiträge 46 (1) on page 742, DOI: 10.5281/zenodo.5306753

Malayloeblius borneensis nov.sp. (Figs 23 and 114-116) T y p e m a t e r i a l Holotype, Sabah, Danum Valley, B.R.L., f.i.t., 14-16.II.2007, G. de Rougemont leg. (CROU). Paratypes: 3, same provenance as the holotype (CROU, MSNV). D e s c r i p t i o n: Length 2.3-2.54 mm. Body shiny, brown, base of the free abdominal tergites yellowish-brown, antennae brown, legs yellow with yellowish-brown femora. Antennae clubbed (Fig. 23), eyes reduced. Fore-body devoid of reticulation, that of the abdomen very superficial. Puncturation of the head very superficial, evident and widely spaced between the antennae and with some sparse more evident punctures. Puncturation of the pronotum like that of the head, with strong punctures aligned on each side of the longitudinal median band. Puncturation of the elytra deep with some strong punctures, that of the abdomen very close and very superficial and with strong punctures aligned transversely on all free tergites. Aedeagus: Figs 114-115; spermatheca: Fig. 116. C o m p a r a t i v e n o t e s: In its habitus and the form of the spermatheca, the new species is similar to M. sausai HLAVÁČ & MARUYAMA, 2004 from Malaysia. The aedeagus is clearly different, slender, with parallel sides in ventral view, and with a long flagellum of the internal sclerotised pieces of the aedeagus in the new species, whereas the aedeagus is stumpy and without a flagellum of the internal sclerotised pieces in M. sausai. The distal bulb of the spermatheca is short in the new species, while is very long in M. sausai. E t y m o l o g y: The new species takes its name from Borneo., Published as part of Pace, R., 2014, Aleocharinae from Sabah (Borneo) collected by Guillaume de Rougemont (Coleoptera, Staphylinidae), pp. 727-794 in Linzer biologische Beiträge 46 (1) on pages 740-741, DOI: 10.5281/zenodo.5306753

Dikraspedella borneensis nov.sp. (Figs 15 and 95-97) T y p e m a t e r i a l Holotype, Sabah, Danum Valley, B.R.L., f.i.t., 14-16.II.2007, G. de Rougemont leg. (CROU). Paratypes: 4 and 3, same provenance as the holotype (CROU, MSNV). N o t e: The ligula of the new species is furnished with two bristles at the base of apical lobes. This was not observed when I described the genus Dikraspedella PACE, 2002; the two bristles were perhaps detached and lost in the process of preparation and attaching the label to the specimen. D e s c r i p t i o n: Length 1.96-3.48 mm. Body shiny and blackish-brown, free abdominal tergites one and two yellowish-brown, antennae brown with the two basal antennomeres yellowish-brown, legs yellowish-red. Second antennomere shorter than the first, third as long as the second, fourth to tenth strongly transverse. Eyes as long as the postocular region in dorsal view. Reticulation of head and pronotum very superficial, that of the elytra strong, that of the abdomen irregular, polygonal, evident. Puncturation of the head fairly deep and absent on the mid-longitudinal band, that of the pronotum of the male dense and evident on either side of the ample discal concavity, the fundus of which is impunctate, that of the elytra very close and superficial. Granulation of the abdomen salient only near the posterior margin of each free tergite. Pronotum of the male with ample discal concavity, absent in the female. The disc of the pronotum bears f disposed in a square in both sexes. Each elytron with four strong punctures, two in a longitudinal line near the suture and two in a longitudinal line near the sides. Tibiae externally spinose. Aedeagus: Figs 95-96; spermatheca: Fig. 97. C o m p a r a t i v e n o t e s: In the form of the spermatheca, the new species is similar to D. kinabaluensis PACE, 2002, also from Borneo but the apical umbilicus of the distal bulb of the spermatheca is narrow in the new species, wide in D. kinabaluensis. The spermatheca of the new species is 0.23 mm long, whereas that of D. kinabaluensis measures 0.57 mm. The aedeagus of the new species is 0.32 mm long, that of D. kinabaluensis 0.27 mm. E t y m o l o g y: The new species takes its name from Borneo., Published as part of Pace, R., 2014, Aleocharinae from Sabah (Borneo) collected by Guillaume de Rougemont (Coleoptera, Staphylinidae), pp. 727-794 in Linzer biologische Beiträge 46 (1) on page 739, DOI: 10.5281/zenodo.5306753

Lambanus rougemonti nov.sp. (Figs 37 and 157-158) T y p e m a t e r i a l Holotype, Sabah, Danum Valley, B.R.L., f.i.t., 14-16.II.2007, G. de Rougemont leg. (CROU). D e s c r i p t i o n: Length 7.12 mm. Body shiny, black, antennae brown with the basal antennomeres blackish-brown and eleventh yellowish-red, legs brown with femora yellowish-red. Second antennomere shorter than the first, third longer than the second, fourth to ninth longer than wide, tenth as long as wide. Eyes shorter than the postocular region in dorsal view. Reticulation of head and pronotum superficial, that of the elytra evident, that of the abdomen very transverse and very superficial, except on the fifth free tergite on which the reticulation is weakly transverse and more evident. Puncturation of the head dense and superficial, that of pronotum and elytra also dense, but evident. Spermatheca: Fig. 157; female sixth free tergite: Fig. 158. C o m p a r a t i v e n o t e s: This specimen is clearly not the female of Lambanus borneensis nov.sp. described above, and must be attributed to a new species, because antennomeres eight and nine are longer that wide, the puncturation of the head is also present on the longitudinal median band, there are two large punctures near the anterior margins of the eyes, and the abdominal tergites are transversely reticulate. E t y m o l o g y: The new species is dedicated to its collector, our colleague, the staphylinid specialist Guillaume de Rougemont., Published as part of Pace, R., 2014, Aleocharinae from Sabah (Borneo) collected by Guillaume de Rougemont (Coleoptera, Staphylinidae), pp. 727-794 in Linzer biologische Beiträge 46 (1) on page 752, DOI: 10.5281/zenodo.5306753

Keratodegnathus rougemonti nov.sp. (Figs 27-28 and 128-134) T y p e m a t e r i a l Holotype, Sabah, Danum Valley, B.R.L., f.i.t, 14-16.II.2007, G. de Rougemont leg. (CROU). Paratypes: 2 and 1, same provenance as the holotype (CROU, MSNV). D e s c r i p t i o n: Length 5.2-5.9 mm. Body shiny, reddish, elytra reddish-brown, abdomen yellowish-red, antennae brown with scape reddish-brown and antennomeres ten and eleven white, legs yellowish-red. Antennae inserted behind the level of the front of the eyes, mandibles of the male and female bearing horns near the base. Scape very long, second antennomere shorter than the scape, third longer than the second, fourth to tenth much longer than wide. Eyes a little longer than the postocular region in dorsal view. Body devoid of reticulation. Head and pronotum impunctate and devoid of granulation, except for two large discal punctures on the pronotum. Granulation of the elytra fine, very sparse and evident. Aedeagus: Figs 128-129; spermatheca: Fig. 130. E t y m o l o g y: The new species is dedicated to its collector, our colleague, the staphylinid specialist Guillaume de Rougemont., Published as part of Pace, R., 2014, Aleocharinae from Sabah (Borneo) collected by Guillaume de Rougemont (Coleoptera, Staphylinidae), pp. 727-794 in Linzer biologische Beiträge 46 (1) on page 747, DOI: 10.5281/zenodo.5306753

Pseudoplandria collaris nov.sp. (Figs 62 and 236) T y p e m a t e r i a l Holotype, Sabah, Danum Valley, B.R.L., f.i.t., 14-16.II.2007, G. de Rougemont leg. (CROU). D e s c r i p t i o n: Length 3.33 mm. Body shiny, yellowish-red, antennae yellowishred with the two basal antennomeres and apical half of the eleventh yellow, legs yellowish-red. Second antennomere as long as the first, third shorter than the second, fourth and fifth longer than wide, sixth as long as wide, seventh to tenth transverse. Eyes longer than the postocular region in dorsal view. Body devoid of reticulation. Puncturation of head and pronotum fine and very superficial. Granulation of elytra and abdomen fine, very superficial and evident. The disc of the pronotum bears four large punctures disposed in a square. Base of the third and fourth free abdominal tergites with strong elongate punctures. Spermatheca: Fig. 236. C o m p a r a t i v e n o t e s: The only other known species with a spermatheca similar to that of this new species is P. kinabaluicola PACE, 2008 also from Borneo, but the antennomeres four to seven are transverse in the new species, whereas they are elongate in P. kinabaluicola. The distal bulb of the spermatheca of the new species is more developed than that of P. kinabaluicola and the proximal portion of the spermatheca is shorter in the new species, longer in P. kinabaluicola. E t y m o l o g y: The name of the new species, meaning "relating to the neck" refers to the very transverse pronotum., Published as part of Pace, R., 2014, Aleocharinae from Sabah (Borneo) collected by Guillaume de Rougemont (Coleoptera, Staphylinidae), pp. 727-794 in Linzer biologische Beiträge 46 (1) on pages 767-768, DOI: 10.5281/zenodo.5306753

Porus borneensis nov.sp. (Figs 56 and 223-226) T y p e m a t e r i a l Holotype, Sabah, Danum Valley, B.R.L., f.i.t., 14-16.II.2007, G. de Rougemont leg. (CROU). Paratypes: 3 exx., same provenance as the holotype (CROU, MSNV). D e s c r i p t i o n: Length 5-5.3 mm. Body shiny, brown, abdomen yellowish-red, antennae yellowish-brown with the two basal antennomeres and the base of third yellowish-red, legs yellowish-red. Second antennomere shorter than the first, third as long as the second, fourth to seventh longer than wide, eighth as long as wide, ninth and tenth transverse. Eyes as long as the postocular region in dorsal view. Reticulation of the head evident, that of the pronotum strong, that of the elytra very superficial, that of the abdomen evanescent. Puncturation of the head evident and moderately close, absent on the longitudinal median band, that of the pronotum evanescent, that of the abdomen scarcely visible. Granulation of the elytra close and superficial. Male first free abdominal tergite with two long parallel lateral appendices. Aedeagus: Figs 223-224; male sixth free tergite: Fig. 226. C o m p a r a t i v e n o t e s: This new species differs from P. ochraceus WESTWOOD, 1839 from India, in the shape of the pronotum, which is as wide as long, whereas it is transverse in P. ochraceus, in the antennomeres four to six, which are longer than wide, whereas in P. ochraceus they are transverse. E t y m o l o g y: The new species takes its name from Borneo., Published as part of Pace, R., 2014, Aleocharinae from Sabah (Borneo) collected by Guillaume de Rougemont (Coleoptera, Staphylinidae), pp. 727-794 in Linzer biologische Beiträge 46 (1) on page 764, DOI: 10.5281/zenodo.5306753

Mimacrotona borneensis nov.sp. (Figs 58 and 228) T y p e m a t e r i a l Holotype, Sabah, Danum Valley, B.R.L., f.i.t, 14-16.II.2007, G. de Rougemont leg. (CROU). D e s c r i p t i o n: Length 1.54 mm. Body shiny, yellowish-red, elytra reddish-brown, antennae yellowish-red with the two basal antennomeres yellow, legs yellow. Second antennomere longer than the first, third shorter than the second, fourth to tenth strongly transverse. Eyes longer than the postocular region in dorsal view. Reticulation of the head very superficial, that of pronotum and abdomen abseent, that of the elytra evident. Puncturation of the head indistinct. Granulation of pronotum and elytra fine and very superficial. First to third free abdominal tergites covered with squamous sculpture, the fourth and fifth with sparse Puncturation. Spermatheca: Fig. 228. C o m p a r a t i v e n o t e s: In its habitus and structure of the spermatheca this new species is similar to M. kinabaluensis PACE, 2008 also from Borneo, but the head and elytra of the new species are brown (yellowish-red in M. kinabaluensis), the spermatheca has an oval distal bulb (spherical in M. kinabaluensis), and the proximal portion of the spermatheca is curved and not rectilinear as in M. kinabaluensis. E t y m o l o g y: The new species takes its name from Borneo., Published as part of Pace, R., 2014, Aleocharinae from Sabah (Borneo) collected by Guillaume de Rougemont (Coleoptera, Staphylinidae), pp. 727-794 in Linzer biologische Beiträge 46 (1) on page 765, DOI: 10.5281/zenodo.5306753

Pseudoplandria fortis nov.sp. (Figs 60 and 231-232) T y p e m a t e r i a l Holotype, Sabah, Danum Valley, B.R.L., f.i.t., 14-16.II.2007, G. de Rougemont leg. (CROU). D e s c r i p t i o n: Length 5.9 mm. Body shiny, reddish, head brown, antennae brown with the two basal antennomeres and the base of the third yellowish-red, legs yellowishred. Second antennomere shorter than the first, third longer than the second, fourth to tenth transverse. Eyes longer than the postocular region in dorsal view. Body devoid of reticulation. Puncturation of the fore-body evident and very superficial, on the pronotum denser anteriorly and on either side of the median line. Frons carinate. Disc of pronotum with four large punctures disposed in a square. Elytra of the male with a salient oblique carina extending half way to the posterior margin and an oblique less salient carina near the humeral angle. Male fifth free abdominal tergite with a salient posterior median carina progressively deeply furrowed posteriorly. Aedeagus: Figs 231-232. C o m p a r a t i v e n o t e s: No known species of this genus has secondary sexual characters on the elytra and on the abdomen resembling those of the new species. The strongly sclerotised internal piece of the aedeagus is also unique to this species. E t y m o l o g y: The new species is named "strong" after the strongly sclerotized internal piece of the aedeagus., Published as part of Pace, R., 2014, Aleocharinae from Sabah (Borneo) collected by Guillaume de Rougemont (Coleoptera, Staphylinidae), pp. 727-794 in Linzer biologische Beiträge 46 (1) on pages 766-767, DOI: 10.5281/zenodo.5306753

Myrmecopella borneensis nov.sp. (Figs 30 and 138-139) T y p e m a t e r i a l Holotype, Sabah, Danum Valley, B.R.L., f.i.t., 14-16.II.2007, G. de Rougemont leg. (CROU). Paratype: 1, same provenance as the holotype (CROU). D e s c r i p t i o n: Length 3.45-3.48 mm. Body shiny, yellowish-red, each elytron with a brown macula extending from the humeral angle to the postero-lateral angle, on the posterior margin of the third free abdominal and a median macula on the fourth tergite; antennae reddish-brown with the basal antennomere yellowish-red, legs yellowish-red. Second antennomere shorter than the first, third longer than the second, fourth longer than wide, fifth as long as wide, sixth to tenth strongly transverse. Eyes shorter than the postocular region in dorsal view. Body devoid of reticulation. Puncturation of the head fine and very superficial. Pronotum finely longitudinally furrowed like the grooves of an old phonographic record. Granulation of the elytra very fine and superficial, that of the abdomen sparse and evident. Aedeagus: Figs 138-139. C o m p a r a t i v e n o t e s: The genus Myrmecopella KISTNER & MCNAIRN, 1991 described from Sulawesi, is new to Borneo. The new species differs from hitherto known species in the absence of a median sulcus on the pronotum. The aedeagus of the new species is short and compact, whereas it is long and narrow in the other species. E t y m o l o g y: The name of the new species is derived from Borneo., Published as part of Pace, R., 2014, Aleocharinae from Sabah (Borneo) collected by Guillaume de Rougemont (Coleoptera, Staphylinidae), pp. 727-794 in Linzer biologische Beiträge 46 (1) on page 748, DOI: 10.5281/zenodo.5306753

Zyras (Glossacantha) plenus nov.sp. (Figs 54 and 218) T y p e m a t e r i a l Holotype, Sabah, Danum Valley, B.R.L., f.i.t., 14-16.II.2007, G. de Rougemont leg. (CROU). D e s c r i p t i o n: Length 9 mm. Body shiny, head black, pronotum blackish-brown, elytra and abdomen yellowish-red, fourth to sixth free abdominal tergites brown, antennae brown with the two basal antennomeres and the base of third yellowish-red, legs yellowish-red. Antennae compressed from the fourth antennomere. Second antennomere shorter than the first, third longer than the second, fourth to tenth as long as wide. Eyes a little longer than the postocular region in dorsal view. Body devoid of reticulation, which is present only on the fourth free tergite on which is very superficial, and on the fifth free tergite on which it is distinct and irregularly polygonal. Puncturation of head moderately dense and superficial, absent on the longitudinal median band, that of the pronotum evident, close on either side of the impunctate longitudinal median band; on the rest of the pronotum the Puncturation is sparse. Puncturation of the elytra a moderately dense and strong, that of the abdomen sparse. Spermatheca: Fig. 218. C o m p a r a t i v e n o t e s: The distal bulb of the spermatheca has a lateral prolongation as in the spermatheca of Z. inversus PACE, 2001 from India. However the sixth free abdominal tergite of the female is not split to half as in Z. inversus, and the spermatheca is less developed than that of Z. inversus, with the apical umbilicus of the distal bulb shallow, whereas in Z. inversus it is very deep. E t y m o l o g y: The name of the new species, meaning "full", refers to the distal bulb of the spermatheca, which is normally empty, but in this case appears nearly filled by the thickness of the cuticle., Published as part of Pace, R., 2014, Aleocharinae from Sabah (Borneo) collected by Guillaume de Rougemont (Coleoptera, Staphylinidae), pp. 727-794 in Linzer biologische Beiträge 46 (1) on page 763, DOI: 10.5281/zenodo.5306753, {"references":["PACE R. (2001): Nuove specie del genere Gyrophaena MANNERHEIM del Monte Kinabalu (Borneo) (Coleoptera, Staphylinidae). - Revue suisse de Zoologie 108: 699 - 737."]}

Orphnebius parabigladiosus nov.sp. (Figs 35 and 148-149) T y p e m a t e r i a l Holotype, Sabah, Danum Valley, B.R.L., f.i.t., 14-16.II.2007, G. de Rougemont leg. (CROU). D e s c r i p t i o n: Length 3.9 mm. Body shiny, head black, pronotum brown, elytra yellowish-brown, abdomen yellowish-red, antennae blackish-brown with the three basal antennomeres reddish-brown, legs yellowish-red. Second antennomere shorter than the first, third longer than the second, fourth to tenth longer than wide. Eyes as long as the postocular region in dorsal view. Body devoid of reticulation. Puncturation of the head reduced to some fine and very superficial punctures near the eyes. Pronotum with only six very evanescent punctures. Granulation of the abdomen salient and fairly dense. Free abdominal tergites concave and bare, male first free tergite with a long thorn on either side, male fifth free tergite strongly punctate. Anterior margin of the pronotum very sinuate posteriorly to its half. Aedeagus: Figs 148-149. C o m p a r a t i v e n o t e s: The habitus of the new species is similar to that of O. bigladiosus (BERNHAUER, 1915) from Sarawak, of which I have examined the male holotype labelled "Madang, 1000 ft. 2.11.19, Sarawak, ded. Moulton 41, Deroleptus bigladiosus Brnh., Typus" (FMHNC). The new species differs from it in the short lateral thorns on the abdomen of the male (long in O. bigladiosus), in the shorter intermediary antennomeres and in the aedeagus curved to the dorsal side, while it is rectilinear in O. bigladiosus. The internal sclerotised part of the aedeagus is composed of two strongly sclerotised pieces in lateral view, whereas in O. bigladiosus only one is strongly sclerotised. The posterior margin of the male sixth free abdominal tergite of O. bigladiosus is tri-lobed with two thorns between the lobes. The posterior margin of the male sixth free tergite of the new species is linear. E t y m o l o g y: The name of the new species mean "close to O. bigladiosus " from the ancient Greek παρά = near., Published as part of Pace, R., 2014, Aleocharinae from Sabah (Borneo) collected by Guillaume de Rougemont (Coleoptera, Staphylinidae), pp. 727-794 in Linzer biologische Beiträge 46 (1) on pages 750-751, DOI: 10.5281/zenodo.5306753, {"references":["BERNHAUER M. (1915): Neue Staphyliniden der indo-malaiischen Fauna, insbesondere der Sunda-Insel Borneo. - Verhandlungen der Zoologisch-Botanischen Gesellschaft in Wien 65: 134 - 158."]}

Orphnebius ocularis nov.sp. (Figs 32 and 142-143) T y p e m a t e r i a l Holotype, Sabah, Danum Valley, B.R.L., f.i.t., 14-16.II.2007, G. de Rougemont leg. (CROU). D e s c r i p t i o n: Length 2.7 mm. Body shiny, yellowish-red, head reddish-black, antennae with the three basal antennomeres yellowish-red. Second antennomere shorter than the first, third as long as the second, fourth to tenth weakly transverse. Eyes very large. Body devoid of reticulation. Puncturation of the head fine, sparse and absent on the longitudinal median band. Puncturation of the pronotum fine except for two large discal punctures. Free abdominal tergites concave and bare. Male fifth free tergite with strong puncturation that does not extend to reach the posterior margin, which is striate. Aedeagus: Figs 142-143. C o m p a r a t i v e n o t e s: The aedeagus of the new species is similar to that of O. directus PACE, 2007 also from Borneo, but the eyes of the new species are enormous, so the postocular region is obsolete, whereas in O. directus the eyes are as long as the postocular region. The aedeagus of the new species is broadly curved to the ventral side, while that of O. directus is only siglthly curved. The internal sclerotised piece of the aedeagus of the new species is strong and long 0.13 mm long, that of O. directus is less strong and 0.1 mm long. E t y m o l o g y: The name of the new species refers to its enormous eyes., Published as part of Pace, R., 2014, Aleocharinae from Sabah (Borneo) collected by Guillaume de Rougemont (Coleoptera, Staphylinidae), pp. 727-794 in Linzer biologische Beiträge 46 (1) on page 749, DOI: 10.5281/zenodo.5306753

Drusilla borneostricta nov.sp. (Figs 50 and 195-198) T y p e m a t e r i a l Holotype, Sabah, Danum Valley, B.R.L., f.i.t., 14-16.II.2007, G. de Rougemont leg. (CROU). D e s c r i p t i o n: Length 7-7.57 mm. Body shiny, blackish-brown, abdomen reddish with the fourth to sixth free abdominal tergites brown, antennae brown with the two basal antennomeres yellowish-red, tarsi and anterior tibiae reddish, posterior and middle tibiae pale yellow, anterior and middle femora yellowish-brown, posterior femora brown with basal half pale yellow. Second antennomere shorter than the first, third longer than the second, fourth to tenth longer that wide. Eyes longer than the postocular region in dorsal view. Reticulation of the fore-body strong, that of the abdomen absent except on the fifth free tergite on which an irregular polygonal reticulation is evident. Granulation of the fore-body very close and superficial, absent on the longitudinal median band of the head. The head bears two small deep fovea near the anterior margin of each eye. Median sulcus of the pronotum deep, the surface very salient near the posterior angles of the pronotum, declivous toward the broad posterior median fovea. There is a feeble lateral keel on the elytra that is salient on the disk. The elytra of the male bear large granules along the posterior 2/3rds of the suture. Free abdominal tergites bare with two median tubercles in transverse line on the second to fourth free tergites. Male first free tergite with a median appendix on the posterior margin (Fig. 198), and beneath this, on the base of the following tergite, a triangular area is flattened and strigose. Male fifth free tergite broadly concave on the median line and impressed on the sides and with two convergent posterior median keels (Fig. 198). Aedeagus: Figs 195-186; male sixth free tergite: Fig. 197; male abdomen: Fig. 198. C o m p a r a t i v e n o t e s: In its habitus and the form of the aedeagus the new species is similar to D. kinabaluensis PACE, 1986 also from Borneo, but the colour of the abdomen is different, the "crista apicalis" is little developed, whereas it is strongly developed in D. kinabaluensis, and the preapical region of the aedeagus is broad in ventral view, whereas it is narrow in D. kinabaluensis. E t y m o l o g y: The name of the new species, meaning "Borneo narrow", refers to the narrow pronotum., Published as part of Pace, R., 2014, Aleocharinae from Sabah (Borneo) collected by Guillaume de Rougemont (Coleoptera, Staphylinidae), pp. 727-794 in Linzer biologische Beiträge 46 (1) on pages 759-760, DOI: 10.5281/zenodo.5306753, {"references":["PACE R. (1986): Aleocharinae dell'Asia sudorientale raccolte da G. de Rougemont (Coleoptera, Staphylinidae). - Bollettino del Museo civico di Storia naturale di Verona 23: 139 - 237."]}

Brachida danumensis nov.sp. (Figs 4 and 77) T y p e m a t e r i a l Holotype, Sabah, Danum Valley, B.R.L., f.i.t., 14-16.II.2007, G. de Rougemont leg. (CROU). D e s c r i p t i o n: Length 1.8 mm. Body shiny, yellowish-red, posterior half of the elytra brown, pygidium yellow, antennae brown with the two basal antennomeres and base of the third yellow. Second antennomere as long as the first, third shorter than the second, fourth to tenth transverse. Eyes longer than the postocular region in dorsal view. Body devoid of reticulation. Puncturation of the head very superficial and evident, absent on the mid-longitudinal median band. Puncturation of the pronotum obsolescent. Granulation of the elytra very superficial. Spermatheca: Fig. 77. C o m p a r a t i v e n o t e s: In the shape of the spermatheca the new species is similar to B. crassiuscula (KRAATZ, 1859) from Sri Lanka, of which I have examined 7 specimens of the type series labelled " Ceylon, J. Nietner, Homalota crassiuscula KRAATZ " (DEI). The new species can be distinguished from B. crassiuscula by the intermediary portion of the spermatheca, between the distal bulb and appendix of the proximal bulb, which is about as long as the maximum width of the distal bulb, whereas this intermediary portion of the spermatheca is absent in B. crassiuscula. B. subadunca PACE, 2002 from Borneo has the intermediary portion of the spermatheca shorter than the maximum width of the distal bulb of the spermatheca. E t y m o l o g y: The new species takes its name from the Danum Valley., Published as part of Pace, R., 2014, Aleocharinae from Sabah (Borneo) collected by Guillaume de Rougemont (Coleoptera, Staphylinidae), pp. 727-794 in Linzer biologische Beiträge 46 (1) on pages 731-732, DOI: 10.5281/zenodo.5306753, {"references":["KRAATZ G. (1859): Die Staphyliniden-Fauna von Ostindien, insbesondere der Insel Ceylan. - Archiv fur Naturgeschichte 25: 1 - 45."]}

Strabocephalium borneorum nov.sp. (Figs 26 and 125-127) T y p e m a t e r i a l Holotype, Sabah, Danum Valley, B.R.L., f.i.t., 14-16.II.2007, G. de Rougemont leg. (CROU). Paratypes: 8 exx., same provenance as the holotype (CROU, MSNV). D e s c r i p t i o n: Length 5.4-5.9 mm. Body shiny, reddish; head, antennae and legs brown. Second antennomere shorter than the first, third longer than the second, fourth to tenth transverse. Eyes longer than the postocular region in dorsal view. Head, pronotum and abdomen devoid of reticulation, that of the elytra evident, forming an irregular polygonal mesh. Puncturation of the head very superficial and very close, absent on the longitudinal median band. Puncturation of the pronotum invisible except for a few punctures on each side of the median line. Granulation of the elytra evident and not very close. Free abdominal tergites bare, posterior ¾ of the male fifth free tergite strongly punctate, the posterior margin of the male fifth tergite with six tubercles. Aedeagus: Fig. 126; spermatheca: Fig. 125; female sixth free tergite: Fig. 127. C o m p a r a t i v e n o t e s The new species is very similar to S. mirabile BERNHAUER, 1911 from Borneo. It differs in the eyes being as long as the postocular region, whereas in S. mirabile the eyes are shorter than the postocular region. The posterior margin of the female sixth free abdominal tergite is plurilobate in the new species, whereas is serrate in S. mirabile (Fig. 124). The distal bulb of the spermatheca is transverse in the new species (Fig. 125), and spherical in S. mirabile (Fig. 123). The proximal bulb of the spermatheca of the new species is sub-spherical, whereas it is pearshaped in S. mirabile. E t y m o l o g y: The new species is dedicated to the inhabitants of Borneo., Published as part of Pace, R., 2014, Aleocharinae from Sabah (Borneo) collected by Guillaume de Rougemont (Coleoptera, Staphylinidae), pp. 727-794 in Linzer biologische Beiträge 46 (1) on page 746, DOI: 10.5281/zenodo.5306753, {"references":["BERNHAUER M. (1911): Zur Staphylinidenfauna Ostindiens und der Sundainseln (3. Beitrag. Schluss). - Entomologische Blatter fur Biologie und Systematik der Kafer 7: 86 - 93."]}